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SMITHSONIAN INSTITUTION 
UNITED STATES NATIONAL MUSEUM 


Butuetin 199 


AN ANNOTATED CHECKLIST AND KEY 
TO THE REPTILES OF MEXICO 
EXCLUSIVE OF THE SNAKES 


BY 


HOBART M. SMITH ann EDWARD H. TAYLOR 











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UNITED STATES 
GOVERNMENT PRINTING OFFICE 
WASHINGTON : 1950 


a a a 
For sale by the Superintendent of Documents, U. S. Government Printing Office 
Washington 25, D. C. - Price 75 cents (Paper) 


ADVERTISEMENT 


The scientific publications of the National Museum include two 
series, known, respectively, as Proceedings and Bulletin. 

The Proceedings series, begun in 1878, is intended primarily as a 
medium for the publication of original papers, based on the collections 
of the National Museum, that set forth newly acquired facts in biology, 
anthropology, and geology, with descriptions of new forms and re- 
visions of limited groups. Copies of each paper, in pamphlet form, 
are distributed as published to libraries and scientific organizations 
and to specialists and others interested in the different subjects. The 
dates at which these separate papers are published are recorded in the 
table of contents of each of the volumes. 

The series of Bulletins, the first of which was issued in 1875, con- 
tains separate publications comprising monographs of large zoological 
groups and other general systematic treatises (occasionally in several 
volumes), faunal works, reports of expeditions, catalogs of type 
specimens, special collections, and other material of similar nature. 
The majority of the volumes are octavo in size, but a quarto size has 
been adopted in a few instances in which large plates were regarded 
as indispensable. In the Bulletin series appear volumes under the 
heading Contributions from the United States National Herbarium, in 
octavo form, published by the National Museum since 1902, which 
contain papers relating to the botanical collections of the Museum. 

The present work forms No. 199 of the Bulletin series. 


ALEXANDER WETMORE, 
Secretary, Smithsonian Institution. 


II 


CONTENTS 


Page 
Introduction_..=..--.------------------------------<-------------- 1 
Itinerary and gazetteer of localities for the Walter Rathbone Bacon 

Expedition, 1938-1941 __----------------------------------------- 4 
Sia Reptilia ss nee eee eae 12 
Subclass Anapsida--------------------------------------------- 12 
Order Testudines-_-_---------------------------------------- 12 
Suborder Atheca......--------------------------------- 13 

Family Dermochelyidae----------------------------- 13 
Genus-Dermochelys=-=-"* >= = ------------- 13 

Suborder Carettoides. === 2 2 Sees = 22 === 14 

Family Cheloniidae_------------------------------- 14 

Genus Lepidochelys_____-----=----------------- 14 

Conus: Carcitg=— eee eee 22 2 552525 15 

@onus- Eretmocheltys-——-- 2-2 e = =—- --=- == ---—— 16 

Ganus’ Chelonta= = 00 ern ee 8 SS e --=- 17 

Suborder Trionychoidea-------------------------------- 18 

Family Trionychidae--_----------------------------- 18 

Genus Amydas ee 2 See rst eee 2 === 18 

Suborder Cryptodira_---------------------------------- 18 

Family Dermatemyidae----------------------------- 19 

Genus Dermatemys. ot 2 ee === --=-= 19 

Family Chelydridae_-__----------------------------- 20 

Genus Chelydrac 1 2 2 ee eee oa a-s- S a 20 

Family Kinosternidae- -_-_--------------------------- 21 

Genus Kanosternon=22- 2-22 21 

Genus Claudius ee ee === 26 

Genus Staurotypus:._=--*2== 2). =--------------- 27 

Family Testudinidae-_-_----------------------------- 27 

Genus Gopherus..--------_--------------------- 28 

Family Emyidae------.---------------------------- 28 

Genus Geoemydas.—=— = -- + 2-2-2 - - =~ -- 29 

Genus: Pseudemys- =. 2 22222 9-2 22 =--=-----=-===- 31 

Genus Chrysemys..----------------------------- 33 

Genus Terrapene_----_------_---=---=----------- 34 

Genus Malaclemys-__--------------------------- 36 

Genus: Clemmijsa= = - ss = Se 2 - = -==== 37 

Subclass Lepidosauria------------------------------------------ 37 
Order Squamata--_...-------------------------------------- 37 
Suborder Amphisbaenia_-------------------------------- 37 

Family Bipedidae-_--------------------------------- 37 

Genus-Bipes =-222 2 eae + == -- = - 38 


IV BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Class Reptilia—Continued 
Subclass Lepidosauria—Continued 


Order Squamata—Continued Page 
Suborder Saurias 2225522 Sea eee re eS oes 39 
Barmily Gekkonidaes 22.522 ae Se eee ee ae ee 40 
Genus: Coleony@ 22-2822 sess eee ee eee eS 41 
Genus’ Gonatod esa se Ot es 2 eee 2 45 
GenustPhylodactyluses sre Sie ee or a ee 46 
(GentseeCad cl yl 1Ls a eee 49 
Genus:hemidactyluss 22 Be ee ee ae ee 49 
Genlisiarrstelizgen ce <= ees. ee Oe ea 51 
Genusei2cr0 piss ee ne ee Se eee 51 
Genus Sphaerodactylus= s22. 2.) ee Se 52 
amily Iguanidae® 252 lacs. ad) fe ae es Se 53 
Ge rnnt sc oe se ee ae = 55 
Genus Conythophanes= — = =22- 2 o> a ee 68 
Genusviaemanctusicss ao) ee eee eee 69 
GeniisBasiltseuss eee ee ees eee ac 
Genus quang= = = 2th ane he se se oe SC 72 
Genus Cienosaura.o. 2222" ee Pe 73 
Genus Pnyaltosaurus. eee. 32a ea ee 75 
GentspDinsosaunus: S22 ee ee eee 77 
Genus SGuromalus... ase eee se ee 78 
Genus: Holbrookiag lsat ee ee a ee 81 
Genus alles wis ee 85 
GentissUmG@s = 22002 35 8st ares ae ee: Oe Se 89 
Genus! Petrosatiis=e ae ae e e ee e  e 90 
Genus) Ste plosa tise ee ee ae 91 
Genus Crotaphytusio n= =a Pe 91 
Genus Gambelia: 2523-5 22422 ee a ee See 93 
Genuse@hnynosomassss 2a eee 94 
GenusySceloponus=2- =e = 2 ee a a eee 105 
Genus: Satotss 5265. - 2S ee as es ae 139 
Genus {WU rosaunus 2) a et sh OS 140 
Gents UC es ee ee ee is Ae rh re 147 
Pamily Nantusudac== 2: 235.2 smn oe eS 151 
Genus: Lepidophiyarna-- ea eae ae a ee 151 
Genus Gatgetas cs) 22-2 Sis ee oan ees Se 153 
Gents. Xantustal-— tee ee cee 154 
Family Scmeidae-2 2 5_ 2 Oe es ee se ee 155 
Genus Mabugyas: . seo are Se eee 155 
Genus! Sercella = 2.) + pear oe ee 156 
Genus :Euimeccse= = 25h anaes ee ee 160 
Family Anelytropsidae sso. 293-222 - ==. 4252525 170 
Genus Anelytropsts.< tere eee 2S 170 
Family ‘Tetidae. 2. << jee ee ee oe kk ee 170 
Genus: Ameiva oe ee ee ee 170 
Genus Cnemidophoris. = 8-9 > 2 eee 2 eee 174 
Genus Gymnophthalmus_— 2-262. - 25 --43255------ 192 
Family Helodermidae=— 2. =. 6-8 See =. 2 5 192 


Genus Helodenmase = soe eee ee eee 192 


CONTENTS 


Class Reptilia—Continued 
Subclass Lepidosauria—Continued 
Order Squamata—Continued 
Suborder Sauria—Continued 
Ramilyeaneriaag~ © 20 <1 k/o 02 Lk ee, ee ee 
(CORURECCLER ES ak Sh tee ee ee ee 
GENUS PAUT ONE A. she tee fa ba en 
GoeH ES ESTEE 2 a re ok em Eee 
Gens Gerri nongis oi) eke i an 
Genus ganic Pee ne a apie a ee ae hs ee 
BamilyeXenosauridaens sen = — Se heen ns ee eee eee 
(Genus E7205 01107115 S ee ea 
amily Anmieliaaesa= 2222 -=~ = 222 kee ne eee ee 
GennseAnnsclas 3-20. BPs ee ee 
Suborder Serpentes.! 

PTR OLHONEAT CHOSAUTIA SA ras ee A ois etee ere e e e ee eS LS oes 
Ordertoriespacy 0 Le eS ie eee BAL ee Peo ey 
Hamil yiCrocodylidae= 2) =- * See were) Se ae 
Gents rocodi/ ussite ee oe ee ee 
Bamily Allipatoridae 4-225 2228 oe 2 oe oe oa 
WET UIS CLO BETA ae is a a A tet hea eh 2 
PEST CORTINA Oe See te ee eee Sn Oe Ute Pee ae 
a ens Dae oes Rate DRM tk Take see ae PIB eS 


1See U. 8. Nat. Mus. Bull. 187, 1945, p. 12. 





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AN ANNOTATED CHECKLIST AND KEY TO 
THE REPTILES OF MEXICO, EXCLUSIVE 
OF THE SNAKES 


By HosBart M. SMITH and EDWARD H. TAYLOR 





INTRODUCTION 


Tuts volume constitutes the last of a series of checklists and keys 
to the herpetological fauna of Mexico. The first, “An Annotated 
Checklist and Key to the Snakes of Mexico” (U. S. Nat. Mus. Bull. 
187), was published on October 5, 1945. The second, “An Annotated 
Checklist and Key to the Amphibia of Mexico” (U. S. Nat. Mus. 
Bull. 194), appeared on June 17, 1948. In this, the third volume, the 
turtles, amphisbaenians, lizards, and crocodilians are treated. The 
preparatory work that preceded these checklists was pursued as time 
permitted from 1932 to the present, a period of about 17 years. It 
has involved a number of expeditions to Mexico at our own expense ” 
and the collecting of more than 50,000 herpetological specimens. 

Except in rare instances the materials have all passed under the 
scrutiny of one or the other of us, and the detailed results of these 
studies are embodied in more than a hundred short or longer reports. 
Our own ideas have often changed with the acquisition of materials 
not originally available. 

It was hoped that the long-awaited work on turtles by Stejneger 
and Hartweg would see the light before the appearance of the turtle 
checklist, but since there is still no certainty of the completion of this 
work in the immediate future it does not seem wise to delay on this 
account. 

We have refrained from anticipating certain forms not now recog- 
nized that are present even in our own collection; the specimens repre- 
senting them are referred without comment to the recognized form 
they most closely resemble. A number of such forms belong to Anolis, 
no adequate revision of which has yet been made, although the senior 
author is contemplating such a work. Certain other groups, especially 
Cnemidophorus and Uta, merit reinvestigation. 

There are many other unsolved problems in Mexican herpetology; 
much more exploration remains to be performed. Probably no single 

2 From 1938 to 1941 I was financed by the Smithsonian Institution on several expeditions, through tenure 
of a Walter Rathbone Bacon Traveling Scholarship.—H. M. S. 
1 


2 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


problem of distribution is more demanding of explanation than the 
extreme dearth of Plethodontidae in the Guerrero “‘island”’ and in the 
high western plateau, and the apparent replacement of that family in 
the latter region, largely by the Ambystomidae. 

The treatment of forms in this volume differs little from that of the 
two preceding volumes, except that the name of the collector of each 
type is added. This name often is as important perhaps as the name 
of the author of a species. Unfortunately, the collector is frequently 
unknown or uncertain. This is true of the collections from Mexico 
studied by Wiegmann, in which case we have given Deppe credit 
for obtaining most of the reptiles. The other two men constituting 
the collecting party, Count Von Sack and Schiede, were, we believe, 
engaged primarily in the collection of orchids. However, it is not 
at all improbable that some forms accredited to Deppe were actually 
taken by one of the others. 

We refrain here from considering the implications that may be 
deduced from a perusal of the combined data of the three volumes, 
such as an investigation of faunal origins, lines of migration, relative 
age of taxonomic groups, and faunal areas of ecological relationships. 
However, we are fully aware that such consideration would doubtless 
yield valuable information. 

The number of forms recognized in the herpetofauna of Mexico, 
including those forms described since the first of these checklists was 
published, is relatively large. The following table represents the 
numbers in the various groups: 


TaBLE 1.—Number of forms recognized in the herpetofauna of Mexico 





Species 
Order or suborder Families Genera and sub- 

species 
Gymnophionas se sey erecen ee eeeae 1 2 2 
@audatate see ees ee eee 4 16 65 
Sahieitiaite ees = = Ho cee ti re ee re i. 25 163 
*Restudines#s 251%. ereiiad. 1 jbens weeiaaas 8 18 49 
Amphisbaenian — = = ss — thee. See ee ere 1 1 3 
SS UU ree ee Re re On ns eee Re 10 47 394 
Serpentest sitar: ais 24 deroereree. fyspen 8 80 486 
WOniCa tae so 2 a ok ee oe. Pet eee 2 2 3 
Totalieeaaie. aii ayes sew) ber ype 41 191 1, 165 


That there will in the future be considerable shifting of species to 
the subspecific category, and changing forms we regard as subspecies 
to the species category, is to be anticipated. On occasion we have 
found even ourselves at variance on certain problems of this sort. 
We both feel, however, that the currently and deservedly popular 
trend toward recognition of subspecies where they exist has brought 


CHECKLIST OF REPTILES OF MEXICO 3 


with it, almost inevitably, occasional overzealous reduction of species 
to subspecies. Such changes are made without due demand for facts, 
but instead with speculations that often merely serve as an excuse to 
tamper with an accepted arrangement that may be equally as plausible. 

One of the most exasperating tasks has been the unraveling of certain 
purely nomenclatorial knots. Several names, for example, are of 
controversial orthography. Worthy of special mention are Agkistro- 
don vs. Ancistrodon, Cnemidophorus tessellatus vs. C. tesselatus, Cory- 
tophanes vs. Corythophanes, and Kinosternon vs. Cinosternon. Opinion 
36 of the International Commission on Zoological Nomenclature 
interprets the permission given by Article 19 to correct any ‘‘error 
of transcription” to mean also “error of transliteration,’ whether 
the original author was aware of his error or not. By such procedure, 
the names Ancistrodon, tessellatus, Corythophanes, and Cinosternon 
would be recommended. On the other hand, Moore, Weller, and 
Knight (Journ. Paleont., vol. 16, 1942, pp. 250-261) maintain with 
excellent reason that only in very clear-cut and exceptional cases is 
any modification of the original orthography of a generic name 
justified. Their view is strengthened by Blackwelder, Knight, and 
Sabrosky (Science, vol. 108, 1948, pp. 37-38). Maintenance of original 
form would perpetuate Agkistrodon, tesselatus, Kinosternon, and 
Corytophanes. Since the proper procedure is not universally or even 
generally agreed upon, we see little advantage in deviation from 
currently accepted form until some means of real standardization is 
provided by the International Commission on Zoological Nomen- 
clature. 

For the first time in these checklists we here adopt the practice of 
placing a comma between scientific name and author in citations of all 
references except the original. 

We wish to acknowledge the assistance of W. Leslie Burger in the 
laborious task of checking and rechecking various points in the 
manuscript, and in criticizing certain parts, and of Dr. Doris M. 
Cochran, zoologist, division of reptiles and amphibians, United States 
National Museum, who devoted much time to curatorial duties 
connected with the voluminous collection and in making available for 
study the National Museum herpetological collections. We are grate- 
ful likewise for the corrections and improvements suggested by Dr. 
D. F. Hoffmeister, Karl P. Schmidt, and Dr. Frederick Shannon. The 
Graduate Research Boards of the University of Illinois and the 
University of Kansas have generously provided financial support for 
research and clerical work involved in completion of the present 
project, and the Walter Rathbone Bacon Scholarship of the Smith- 
sonian Institution furnished a substantial sum for travel and research 
in Mexico. 


4 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


ITINERARY AND GAZETTEER OF LOCALITIES FOR THE WALTER 
RATHBONE BACON EXPEDITION, 1938-1941 


As previously stated, the senior author was enabled to collect in 
and study material from Mexico by tenure of a Walter Rathbone 
Bacon Traveling Scholarship from 1938 to 1941. The collection 
obtained during those years has formed an important addition to 
other material from Mexico and merits an elaboration of the routes 
traversed by the expedition and the localities at which specimens 
were secured. Such an itinerary was, as a matter of fact, to have 
accompanied a study of the lizards of the Bacon collection, as stated 
in our introduction to the summary of the amphibians (Proc. U. S. 
Nat. Mus., vol. 95, 1945, p. 521). It now seems unlikely that a com- 
plete survey of the Bacon lizards can appear for a number of years, 
for a satisfactory completion of such a study involves revision of a 
number of very sketchily understood genera, such as Anolis and 
Onemidophorus. In the meantime a need for an account of the Bacon 
itinerary and localities continues to exist. We have, accordingly, 
taken this opportunity to present this information in this the final 
checklist. 

The expedition personnel consisted of the senior author and his wife. 
They received hearty support in field work from a number of indi- . 
viduals, through whose combined efforts about 22,000 specimens were 
secured during a period of two years. As closely as can now be deter- 
mined (the lizards not yet having been fully studied), 500 species and 
subspecies (146 amphibians, 160 lizards, 1 amphisbaenian, 170 snakes, 
20 turtles, 3 crocodilians) were secured. 

Among those who contributed to the success of the expedition are 
Dr. Alexander Wetmore, who saw to it that the expedition was prop- 
erly planned and could function smoothly; Dr. Doris M. Cochran, 
who must have nearly equaled the expedition personnel in time ex- 
pended for the collection, inasmuch as the laborious task of cataloging 
the specimens rested with her; Dr. Linton P. Satterthwaite, who 
provided the facilities for our stay at Piedras Negras in the midst of a 
little-known, remote, and austere area; Mr. and Mrs. Dyfrig McH. 
Forbes, who provided for nearly two years a base of operations in 
Veracruz and who were a constant source of companionship, inspira- 
tion, and material; Eizi Matuda, who very generously provided facil- 
ities for a two-month stay at his finca and who sympathetically aided 
us in every possible way to sample the herpetofauna of the area as 
thoroughly as possible; Thomas MacDougall, who in Tehuantepec 
secured for us numerous specimens and arranged for our travel into 
areas otherwise difficult of access; Ernest Rateike, of Palenque, 
Chiapas, who accepted us as a guest in his home for a month; and a 


CHECKLIST OF REPTILES OF MEXICO 5 


host of local residents, far too numerous to mention, who materially 
augmented our collections and made our visits pleasant and successful. 

Inasmuch as we traveled chiefly by car, our routes are best followed 
on aroad map of Mexico. We entered Mexico on October 5, 1938, at 
Ciudad Judrez, Chihuahua, and followed the Chihuahua—El Paso 
highway southward to Vado (October 9). We then turned back to 
Ahumada and followed a poor trail through Carrizal and Rancho 
Nuevo to Progreso (October 10). We camped beside the Rio Santa 
Maria about a mile southwest of Progreso for 5 days, interrupted only 
by one hurried round trip to Casas Grandes for the benefit of a snake- 
bitten cowboy. At this camp we were aided greatly by 10 or 15 
cowboys who brought in material as fast as it could be preserved. 
Our specimen containers full, we determined to go to the border to 
ship specimens and likewise to obtain higher wheels for the car (a 
half-ton panel truck), with which we had encountered numerous diffi- 
culties because of deep ruts and high centers. We left the Progreso 
camp on October 15, passing through Casas Grandes and Ascencién, 
and reached Las Palomas the same day. After two days in Columbus 
and Deming, N. Mex., we retraced our steps (October 20) from Las 
Palomas to Casas Grandes. As the weather had become cool, we 
continued southward along the road through San Buenaventura and 
Carmen to the main E] Paso—Chihuahua highway, thence southward 
to Ciudad Chihuahua. We left there on October 27 for Torreén, 
arriving October 29. We collected in the vicinity of Torreén October 
30 and 31, and on November 1 started for Ciudad Durango. We 
could get no farther than Pedricefia, however, and, after exploring 
with little success a side road to Nazas, returned to Torreén on Novem- 
ber 4. We continued to San Pedro the next day and collected in the 
vicinity of that town until November 10. After a side trip to Parras 
off the Torreén-Saltillo highway, we made an attempt to find Jaral, 
Coahuila, a locality made famous zoologically by Heller and Barber. 
This turned out to be an abandoned ranch near Hipdlito, and, as no 
likely habitat for the montane species recorded from Jaral appeared 
to be nearer than 20 miles or so, we continued to Saltillo (November 
14). We collected in the vicinity of Saltillo, Arteaga, and Mount 
Zapalinamé until November 17, when we left for Monterrey. 

On November 18 we arrived at Hacienda La Clementina (near 
Forlén), and we remained there until November 28, when we drove 
to Laredo to ship another lot of specimens. We returned to La 
Clementina on December 3 and left December 5 for Mexico City. 
We stayed at Huichihuay4n for several days, leaving December 9. 
Our route, with brief stops at or near various towns, then led through 
Mexico City to Tehuacan, Puebla, thence through Orizaba to Potrero 
Viejo, Veracruz, where we arrived on December 11. 


6 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Potrero Viejo remained our headquarters for over a month. We 
made numerous short trips during that time; especially noteworthy 
were those to Palma Sola (on the Veracruz—Orizaba highway), to 
Tezonapa, Veracruz, and Cosolapa, Oaxaca, and to Cuautlapan, 
Orizaba, Cérdoba, Acultzingo, and other localities along the Cérdoba- 
Tehuacan road. We left Potrero Viejo on January 16, 1939, and 
after a brief stop in Mexico City made headquarters in Cuernavaca, 
Morelos, for nearly two weeks, with side trips to Puente de Ixtla 
(Morelos), Cacahuamilpa (Guerrero), and Zempoala (Morelos and 
México). 

Leaving Cuernavaca February 2, we continued toward Acapulco, 
reaching there February 5. We collected in the vicinity of Acapulco, 
with side trips to Coyuca, until February 11. Our route, with fre- 
quent stops, then led northward again to Mexico City, where we 
arrived on March 1. After two days in search of axolotls and other 
ambystomids in the vicinity of Mexico City (with side trips to Texcoco, 
Zumpango, and Chimalhuac4n), we started (March 4) on the road to 
Guadalajara. Our first deviation from this route was on March 9 
and 10, to Patzcuaro. On March 11 we reached Uruapan on another 
side trip and continued southward to Apatzing4n, returning to 
Uruapan on March 19. An attempt was made to find Crotalus poly- 
stictus in the marshes of the eastern end of Lake Chapala, but without 
success, since most of the marshes have been drained and are now under 
cultivation. We reached Guadalajara on March 24 and returned to 
Potrero Viejo on March 26. On March 30 we left Potrero Viejo 
enroute to Laredo with another shipment of specimens, arriving at 
Laredo on April 3, 1939. 

After nearly a month in the United States we left Laredo on April 29, 
1939, and arrived in Potrero Viejo on May 2. From there we went 
to Veracruz by train and secured passage for Alvaro Obregén, arriving 
on May 13. A river boat was then taken, via Ciudad del Carmen, 
to Tenosique, Tabasco, the head of navigation on the Rio Usumacinta. 
Our final destination, Piedras Negras, Guatemala, was reached on May 
21 after two days by mule. We collected in the immediate vicinity 
of Piedras Negras until June 23, when we left for Tenosique. 

From Tenosique (June 30) we continued downstream to Emiliano 
Zapata and there procured mules for a 2-day trip to San Juanito, a 
ranch half a mile from the village of Palenque. There we remained, 
with a side trip only to the ruins of Palenque several miles distant 
in the hills, until August 6, when we left for Alvaro Obregén and, 
immediately thereafter, Potrero Viejo, where we arrived on August 14. 
There we were joined by the junior author and with him left on 
August 18 for Mexico City, where we arrived, via the usual route 
through Tehuac4n and Puebla, on August 22, after numerous brief 


CHECKLIST OF REPTILES OF MEXICO 7 


stops enroute. After afew days spent in the environs of Mexico City 
searching for ambystomids, on August 29 we took the Acapulco road 
and followed it with only one deviation to Tixtla, Guerrero, arriving 
at Acapulco on September 3. We returned to Mexico City by the 
same route, arriving on September 6. Again we collected in the 
immediate environs of Mexico City, chiefly investigating montane 
faunas. On September 10 we left for Guadalajara but turned back 
near Sahuayo. We then, on September 16, started northward on the 
Pachuca Road to El Chico National Park in Hidalgo. After one day 
there we returned, September 18, to Mexico City, where Dr. Taylor 
left us. 

We then returned, September 19, to Potrero Viejo. After a few 
local trips we again returned to the Guadalajara road, leaving Sep- 
tember 29. We took the side road to the Nevado de Toluca (October 
2) and then retraced our steps to Mexico City, arriving October 3. 
After a few days spent in local trips, we turned northward to Laredo 
with another load of specimens, arriving at Laredo October 17. Our 
only side trip enroute was to Galeana, Nuevo Leén, via the road from 
Linares, Tamaulipas (October 13, 14). 

After nearly two months in the United States, we again crossed the 
border at Laredo on December 10, 1939, and headed directly for 
Potrero Viejo, Veracruz, arriving December 15. After a few local 
trips near Potrero Viejo, we left (December 27) for Tehuantepec by 
rail. We arrived on December 30 and remained until January 28, 
1940. A number of side trips were taken, by truck to Cerro Arenal, 
by rail to Matias Romero, Oaxaca, by rail to Salina Cruz, and by rail 
to Tonal4&. Illness necessitated return to Potrero Viejo and ulti- 
mately to Mexico City. We remained there until March 15, when we 
returned to Potrero Viejo. On March 17 we started on a side trip, 
lasting until March 24, which led by highway through Tehuacan, 
Tecamachalco, thence northeastward on the Jalapa road, to Teziutlan 
(Puebla) from El Limén, Totalco, and eastward to Puente Nacional. 

Shortly thereafter we left for Tehuantepec by rail again, arriving 
April 1. We stayed only one day and traveled by rail to Acapetahua, 
whence we went by bus to Escuintla. From there pack animals were 
used to transport our equipment to La Esperanza, a finca about 5 
miles northeast of Escuintla. We arrived there on April 4 and re- 
mained until June 5, 1940. Various short side trips were taken to 
nearby fincas. 

On June 5 we returned to Tehuantepec and, after one day, to 
Potrero Viejo, arriving June 9. About one month later, on July 6, we 
crossed the border at Laredo with no deviations from the direct route 
from Potrero Viejo to the border. 


8 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


We returned immediately to Mexico City by way of the Pan 
American Highway and remained there until August 7, except for a 
trip by rail to Guanajuato, Guanajuato, on July 19-21. On August 8 
we again drove to Potrero Viejo. Within a few days we made a brief 
and final foray along the highway from Acultzingo to Tehuac4n before 
packing all equipment that had been stored at Potrero Viejo. We 
left our headquarters thers for the final time on August 8 and, with 
but a brief stop in Mexico City, traveled slowly northward along the 
Pan American Highway, with a heavy load, crossing the border at 
Laredo on August 24. 

The following gazetteer includes only those localities represented by 
specimens in the Bacon collection that are not to be found on the 
National Geographic Society’s 1939 map of Mexico, Central America, 
and the West Indies, scale 1:5,702,400. Most of the localities are 
plotted on other, larger-scale maps, but inasmuch as these are not 
always readily available we include all missing from the Geographic 
Society’s map, which is readily available to all investigators. The 
localities are arranged alphabetically within the states, which are 
themselves alphabetically arranged. 


CHIAPAS 


Acacoyagua: 3 miles northwest of Escuintla. 

Aguacate: 7 miles north-northeast by north of Palenque. 

Belén: 20 miles southeast of Escuintla. 

Cerro Obando: 3 miles northeast of La Esperanza (which see). 

Colonia Hidalgo: 7 miles northeast of Acacoyagua (which see). 

Colonia Soconusco: 12 miles northeast of La Esperanza (which see). 

Cruz de la Piedra: 2 miles west of La Esperanza (which see). 

Finca Juarez: 10 miles northeast of La Esperanza (which see). 

Javarinero: 13 miles east of Palenque. 

La Esperanza: 3 miles east of Acacoyagua (which see). 

La Magnolia: 3 miles northwest of La Esperanza (which see). 

Las Nubes: On Cerro Obando, 2 miles northeast of La Esperanza (which see). 
Motozintla: 25 miles east-northeast by east of Escuintla. 

Rancho Las Gradas: 2 miles west of La Esperanza (which see), 

Salto de Agua: On Cerro Obando, 1 mile northeast of La Esperanza (which see). 
San Juanito: 1 mile east-northeast of Palenque. 

Santa Rosa: Near Comitdn. 


CHIHUAHUA 


Carmen: 32 miles east-northeast by east of Galeana. 

Carrizal: 10 miles west-southwest by south of Ahumada. 

Ciudad Delicias: 5 miles south of Meoqui. 

Ojo de Federico: A small ranch 8 miles southeast of Ascencién. 

Progreso: 35 miles northeast of Galeana, near Rio Santa Marfa. 

Rancho Nuevo: A small ranch 10 miles west-northwest by west of Carrizal. 
Samalayuca: 25 miles south of Ciudad Judrez. 


CHECKLIST OF REPTILES OF MEXICO 


COAHUILA 


Hip6lito: 25 miles north-northeast of General Cepeda. 
Zapalinamé: A mountain 3 miles south of Saltillo. 


DISTRITO FEDERAL 


Atzacoalco: 10 miles northeast of Mexico City. 
Cafiada de Contreras: 5 miles southeast of Mexico City. 


DURANGO 


La Goma: 12 miles west of Lerdo, on the south side of the Rfo Nazas. 
La Loma: 12 miles west of Lerdo, on the north side of the Rio Nazas. 
Pedricefia: 25 miles east-southeast of Nazas. 


GUATEMALA 


Desempefia: 4 miles southeast of Piedras Negras. 
Pozo de la Jicotea: 2 miles southeast of Piedras Negras. 


GUERRERO 


Agua Bendita: 13 miles south of Taxco. 

Agua del Obispo: 8 miles north of Dos Caminos. 
Cacahuamilpa: 10 miles southeast of Tetecala, Morelos. 
Coyuca: 7 miles northeast of Acapulco. 

El Treinta: 20 miles north of Acapulco. 

Garrapatas: 5 miles southwest of Dos Caminos. 

Julian Blanco: 4 miles north of Dos Caminos. 

Ocotito: 3 miles north of Dos Caminos. 

Omilteme: 3 miles northwest of Chilpancingo. 

Paso del Limonero: 9 miles northeast of Acapulco. 
Tierra Colorada: 3 miles southwest of Garrapatas (which see). 
Xaltianguis: 31 miles northeast of Acapulco. 


HIDALGO 


Barranca de los Horcones: 6 miles south of Durango (which see). 
Chapulhuacan: 19 miles southwest of Tamazunchale, San Luis Potosf. 
Colonia: 6 miles southeast of Pachuca. 

Durango: 15 miles west-southwest by south of Jacala. 

El Chico Parque Nacional: 15 miles northeast of Pachuca. 

Maguey Verde: 7 miles south of Durango. 

Tianguistengo: 10 miles north-northeast of Zacualtipdn. 


MEXICO 


Chalco: 13 miles northeast of Xochimilco. 

Chimalhuacan: 12 miles east of Mexico City. 

Lerma: 10 miles east of Toluca. 

Nevado de Toluca: 32 miles south-southwest by south of Toluca. 
Rio Frio: 18 miles west-northwest of Texmelucdn, Puebla. 
Santa Magdalena: 15 miles east of Mexico City. 

Zempoala: 6 miles west-southwest of Tres Cumbres, Morelos. 


10 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 
MICHOACAN 


Carapa(n): 23 miles north of Paracho. 

El Temazcal: 20 miles east of Morelia. 

La Palma: 10 miles north of Sahuayo. 
Puerto Hondo: 23 miles east of Morelia. 
Rancho San José: 24 miles east of Morelia. 
Tacicuaro: 5 miles east of Quiroga. 


NUEVO LEON 


Mamulique Pass: 45 miles north of Monterrey. 
Santa Caterina: 7 miles west of Monterrey. 


OAXACA 


Cajén de Piedra: 12 miles south-southwest of Salina Cruz. 

Cerro Arenal: 16 miles west of Tehuantepec. 

Cerro de Chipehua: 16 miles southeast of Salina Cruz. 

Cerro de Huamelula: 34 miles west-southwest by south of Tehuantepec. 
Cerro Guengola: 10 miles west-northwest by west of Tehuantepec. 
Cosolapa: 8 miles northwest of Acatlin. 

Coyol: between San Antonio and Las Cruces. 

El] Limén: 25 miles southeast of Tehuantepec. 

Escurano: 15 miles west-northwest of Tehuantepec. 

La Concepcié6n: 32 miles west of Tehuantepec. 

Lachiguiri: 29 miles northeast of Tehuantepec. 

La Gloria: 8 miles southeast of Chimalapa. 

Las Pilas: 4 miles northwest of Tehuantepec. 

Las Vacas: 40 miles west of Tehuantepec. 

Llano Ocotal: 20 miles west-southwest by west of Tehuantepec. 
Matias Romero: 34 miles north of San Gerénimo. 

Mixtequilla: 4 miles northwest of Tehuantepec. 

Palmar: 37 miles west-northwest by west of Tehuantepec. 

Pixixi: 6 miles south of Tehuantepec. 

Portillo Guayabo: 16 miles west of Tehuantepec. 

Portillo Las Vacas: 40 miles west of Tehuantepec. 

Portillo Los Nanches: 32 miles northwest of Tehuantepec. 

Rincén San Pedro: 16 miles northeast of Tehuantepec. 

Rio Grande: 20 miles west-southwest by south of Tehuantepec. 
Rio Hondo: 41 miles west-northwest by west of Tehuantepec. 

San Francisco Guichina: 61 miles west-northwest by west of Tehuantepec. 
San José Manteca: 61 miles west-northwest by west of Tehuantepec. 
San Mateo del Mar: 14 miles east-northeast by east of Salina Cruz. 
San Pedro Quiechapa: 10 miles west of Yautepec. 

Tenango: 24 miles west-southwest by west of Tehuantepec. 

Tres Cruces: 43 miles west-northwest by west of Tehuantepec. 
Yerba Santa: 10 miles west-northwest by west of Tehuantepec. 


PUEBLA 


Alchichica: 10 miles southwest of Perote, Veracruz. 
Cacaloapam: 8 miles southeast of Tlacotepec. 

El Seco: 27 miles northeast of Tepeaca. 

La Virgin: 2 miles north of Cacaloapam. 


CHECKLIST OF REPTILES OF MEXICO 11 


Ozumbilla: 4 miles east-southeast of Morelos. 

Pajaro Verde: 300 feet west of Puebla—Veracruz state line, about 7 miles south- 
east of Morelos. 

Tecamachalco: 15 miles northwest of Tlacotepec. 


SAN LUIS POTOSi 
Pujal: 20 miles north of Tancanhuitz. 
TABASCO 


El Retiro: 13 miles south of Tenosique. 
Los Rieles: 10 miles south of Tenosique. 
Santo Tomas: 12 miles southeast of Tenosique. 


TAMAULIPAS 


Ciudad Mante (=Villa Judrez): 18 miles east-northeast by north of Antiguo 
Morelos. 

Forlén: 18 miles east-southeast of Llera. 

Hacienda La Clementina: 13 miles east-southeast of Llera. 


TLAXCALA 
Apizaco: 8 miles north-northwest by north of Tlaxcala. 
VERACRUZ 


Acultzingo: 9 miles east-southeast by east of Morelos, Puebla. 
Atoyac: 4 miles east-northeast of Potrero Viejo (which see). 
Cerro Gordo: 20 miles east-southeast of Jalapa. 
Cuautlapan: 4 miles southwest of Fortin (which see). 
Cruz Blanca: 8 miles northeast of Perote. 
El Lim6n Totalco: 10 miles southwest of Perote. 
El Maguey: 8 miles east-southeast by east of Potrero Viejo (which see). 
Encero: 8 miles east-southeast of Jalapa. 
Fortin: 5 miles east of Cérdoba. 
Matacabestro: 7 miles north-northwest by north of Joaquin. 
Mata de Cafia: 18 miles east-southeast of Jalapa. 
Medellin: 10 miles south of Veracruz. 
Metlac: 1 mile west of Fortin (which see). 
Ojo de Agua: 2 miles east of Paraje Nuevo (which see), near Potrero Viejo. 
Pan de Olla: 8 miles south of Teziutlin, Puebla. 
Paraje Nuevo: 2 miles east of Pefiuela (which see). 
Paso del Macho: 5 miles northeast of Atoyac. 
Pefiuela: 4 miles southeast of Cérdoba. 
Potrero Viejo: 2 miles east of Paraje Nuevo (which see). 
Presidio: 22 miles southwest of Cérdoba. 
San Juan de La Punta: 15 miles east-southeast by south of Cérdoba. 
San José de Gracia: 13 miles southeast of Cérdoba. 
Palma Sola: 9 miles east-southeast by south of Cérdoba. 
Tequeyutepec: 10 miles northeast of Jalapa. 
Tezonapa: 8 miles northwest of Acatldn. 
Toxtlacuaya: 17 miles northwest of Jalapa. 
Xuchil: 1 mile north of Potrero Viejo (which see). 
861316—50——2 


12 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Class REPTILIA Laurenti 


Reptilia LAurRENTI, Specimen medicum exhibens synopsin reptilium, 1768, p. 19. 
Subclasses —Three subclasses are represented by living members. 
All occur in Mexico. 
KEY TO MEXICAN! SUBCLASSES, ORDERS, AND SUBORDERS OF REPTILIA 


1. Provided with a shell encasing body above and below, consisting of plastron and 


carapace; turtles__subclass Anapsida_____-_-_- order Testudines (p.12)._ 5 
Not provided witha shell” 22 {oS 2 eo wees cen nme marae eee ts Saree 2 
2. Anal slit longitudinal; skull diapsid___-_subclass Archosauria. 


order Loricata (p. 209) 
Anal slit transverse; skull modified diapsid; lower (quadratojugal-quadrate) 
arch interrupted_-_subclass Lepidosauria- ----__-- order Squamata (p. 37) 

3. Two limbs only, the forelegs; body ringed with grooves. 
suborder Amphisbaenia (p. 37) 
Four limbs or none; body not ringed with grooves_______..___.---------- 4 

4. Limbs present or, if not, movable eyelids present; lizards. 

suborder Sauria (p. 39) 


Limbs absent; movable eyelids absent; snakes___.____-_- suborder Serpentes 
6: ¢Novepidermaltscutes*onishell: ans at eee ss eee et es eee 6 
Epidermial-scutes*present=..2 2s awe eee ne eee seam are meee a 


6. Limbs without external evidence of digits, oar-shaped, clawless. 
suborder Atheca (p. 13) 
Limbs with distinct digits, 3 with claws-___-_- suborder Trionychoidea (p. 18) 
7. Limbs oar-shaped, with 1 to 3 claws__.--_----- suborder Carettoidea (p. 14) 
Limbs not oar-shaped, at least foreleg with 4 or 5 claws. 
suborder Cryptodira (p. 18) 


Subclass ANAPSIDA Williston 


Anapsida WILLIsToN, Journ. Geol., vol. 25, 1917, p. 419. 
Orders.—A single living order exists, the Testudines. 


Order TESTUDINES Batsch 


Testudines Batscu, Versuch Anleit. Kenntn. Gesch. Thier. Mineral., vol. 1, 
1788, p. 437. 

Chelonia MacartTNEY, in Ross, Transl. Cuvier’s Legons d’anatomie comparée..., 
vol. 1, 1802, tab. 3. 

Testudinata OppEL, Die Ordnungen, Familien und Gattungen der Reptilien ... , 
1811, p. 3. 


Suborders.—Five living suborders of Testudines are recognized; 
four occur in Mexico, and the fifth (Pleurodira) is restricted to south- 
ern portions of both hemispheres. 


3 Characters in all keys apply to Mexican forms but not necessarily to those in other parts of the world. 


CHECKLIST OF REPTILES OF MEXICO 13 


Suborder ATHECA Cope 


Athecae Corr, Proc. Amer. Assoc. Adv. Sci., vol. 19, 1870, p. 235. 
Families.—One. 


Family DERMOCHELYIDAE ‘ Fitzinger 


Dermatochelydae Firzincer, Systema reptilium, 1843, p. 30. 


Genera.—One. 
Range.—Tropical oceans of the world. 


Genus DERMOCHELYS Blainville 


Dermochelys BLAINVILLE, Bull. Soc. Philom., 1816, p. 119. 


Genotype.— Testudo coriacea Linnaeus. 

Range.—World-wide, in tropical oceans and occasionally into tem- 
perate waters. 

Species.—As many as three species (or subspecies) may be valid. 
The species has been recorded in Mexico only from the western coast, 
although Atlantic records are to be expected. 


DERMOCHELYS CORIACEA 5 (Linnaeus) 


Testudo coriacea LINNAEUS, Systema naturae, ed. 12, vol. 1, 1766, p. 350. 

Dermochelys coriacea, BLAINVILLE, Bull. Soc. Philom., 1816, p. 119.—BouLENGER, 
Catalogue of the chelonians . . ., 1889, p. 10.—DerranriyaGaua, Tetrapod 
reptiles of Ceylon, vol. 1, 1939, pp. 38-102, figs. 12-34.—ScumiptT, Marine 
Life Occ. Pap., vol. 1, No. 3, 1945, pp. 7-10. 

Sphargis coriacea schlegelit GARMAN, U.S. Nat. Mus. Bull. 25, 1884, p. 303 (type 
unknown; type locality, tropical Pacific and Indian Ocean, here restricted to 
Guaymas, Sonora). 

Dermochelys schlegelii, StEJNEGER, U.S. Nat. Mus. Bull. 58, 1907, p. 485. 


Type.—Unknown. 

Type locality.—Palermo, Sicily, by present restriction. 

Range.—Pacific coast and probably the tropical Atlantic coast. 
Recorded definitely from Sonora: Guaymas; Baja California: Los 
Coronados Islands. 


4 Actually the first family name proposed for this group was Sphargidae Gray (Ann. Philos., vol. 10, 1825, 
p. 212), based upon the genus Sphargis Merrem, 1820, a junior synonym of Dermochelys Blainville, 1816; 
both genera are based upon the same type, Testudo coriacea Linnaeus. Many taxonomists retain the oldest 
family name regardless of the status of its type. We believe there is a certain degree of error courted by 
such a procedure, however, inasmuch as a family name based upon a generic name later found to be a junior 
homonym of an earlier name in another family would, very unfortunately, be preserved; and what if the 
senior homonym had also served as a type forits family? It seems to us that only currently valid generic 
names should be available as types for family names, and that all synonyms and homonyms should be 
regarded unavailable for that purpose. 

5 A common arrangement restricts coriacea to the Atlantic, schlegelii to the Pacific, either as species or sub- 
species. Weare unable to find that anything more than geographic probability has led to such arrangemente 


14 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Suborder CARETTOIDEA® Fitzinger 


Carettoidea Fitzincer, Neue Classification der Reptilien . . ., 1826, p. 5. 
Families.—One. 


Family CHELONIIDAE Gray 


Cheloniadae Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 212. 


Genera.—Four. 
Range.—Tropical oceans, world-wide. 


KEY TO GENERA OF THE FAMILY CHELONIIDAE? 


1. One pair of prefrontal scutes; costal scutes 4; horny cutting edge of lower jaw 
coarsely dentate, that of upper jaw strongly ribbed vertically; bony alveolar 
ridge of upper jaw with a low but regularly raised auxiliary ridge behind 
anterior ridge, which is very strong and terminates anteriorly in a tooth at 
posterolateral corner of premaxillary pit-___._.__._____---- Chelonia (p. 17) 

Two pairs of prefrontals; costal scutes 4-9; horny cutting edge of lower jaw 
smooth or feebly denticulate, that of upper jaw without markedly elevated 
vertical ribbing on its inner surface; bony alveolar surface of upper jaw 
smooth or with a single ridge; this ridge not terminating anteriorly in a 
GS FRSA TPO U0 ap fa ca rea se ce one Le ge es ee re 2 

2. Costal scutes 4 pairs; dorsal scutes usually conspicuously imbricate; bony 
alveolar surface of upper jaw with a sharp-crested ridge. 

Eretmochelys (p. 16) 

Costal scutes 5 pairs or more; dorsal scutes not conspicuously imbricate; bony 
alveolar surface of upper jaw smooth or with a rounded ridge________-_- 3 

3. Four enlarged inframarginal scutes on bridge; dorsal color gray to olive-green; 
maxillaries not in contact, separated by prevomer--__- Lepidochelys (p. 14) 

Three enlarged inframarginal scutes on bridge; dorsal color brown or reddish 
brown; maxillaries in contact between prevomer and premaxillaries. 

Caretta (p. 15) 


Genus LEPIDOCHELYS Fitzinger 


Lepidochelys Firzineer, Systema reptilium, fasc. 1, 1843, p. 30. 
Caouana Gray, Catalogue of the tortoises . . . in the British Museum, 1844, p. 
52 (type, Chelonia olivacea Eschscholtz). 


Genotype.—Chelonia olivacea Eschscholtz. 
Species.—Two are generally recognized. 
Range.—Tropical oceans, world-wide. 


KEY TO SPECIES OF LEPIDOCHELYS 


1. Inframarginal scutes without pores; usually 5 pairs of costal scutes; color usually 
gray; bony alveolar surface of upper jaw with a conspicuous ridge. 

kempii (p. 15) 

Each inframarginal scute usually with a pore at its posterior border; usually 

more than 5 pairs of costal scutes; color olive; bony alveolar surface of upper 

jaw with a low (not conspicuous) ridge___---.._.--------- olivacea (p. 15) 


6 Deraniyagala (Spolia Zeylanica, vol. 24, 1945, p. 98) suggests that this group may be of pleurodirous 
origin. 
1 Adapted from Carr, Proc. New England Zool. Club, vol. 21, 1942, pp. 3-5. 


CHECKLIST OF REPTILES OF MEXICO 15 


LEPIDOCHELYS KEMPII (Garman) 


Thalassochelys (Colpochelys) kempit GARMAN, Bull. Mus. Comp. Zool., vol. 6, 
1880, p. 123. 

Lepidochelys kempii, Baur, Amer. Nat., vol. 24, 1890, p. 487.—Carr, Proc. 
New England Zool. Club, vol. 21, 1942, pp. 4, 8-13, pls. 2, 4. 


Type.—Several cotypes, Mus. Comp. Zool. 

Type locality.—Gulf of Mexico, here restricted to Key West, Fla. 

Range.—Atlantic Ocean and Gulf of Mexico. In Mexico, the entire 
Atlantic coast. The only exact record known is from Quintana 
Roo (Isla de Mujeres). 


LEPIDOCHELYS OLIVACEA (Eschscholtz) 


Chelonia olivacea Escuscuoutz, Zoologischer Atlas, pt. 1, 1829, p. 2, pl. 3. 

Lepidochelys olivacea, GirarD, United States Exploring Expedition... , vol. 
20, Herpetology, 1858, p. 4835.—Drrantyacata, Tetrapod reptiles of Ceylon, 
vol. 1, 1939, pp. 123-163, figs. 43-65.—Carr, Proc. New England Zool. 
Club, vol. 21, 1942, pp. 4-5, pl. 5 (skull). 

Chelonia dussumiert Dumfrin and Brsron, Erpétologie générale . . . , 1835, p. 
557 (type in Mus. Hist. Nat. Paris; type locality, Malabar). 

Caretta remivaga Hay, Proc. U. S. Nat. Mus., vol. 34, 1908, pp. 194-197 (type, 
U.S.N.M. No. 9973; type locality, Ventosa Bay, Oaxaca). 


Type.—Unknown, 

Type locality —Manila Bay, Philippine Islands. 

Range.—Indian and Pacific Oceans. In Mexico, the entire Pacific 
coast. Reported only from the states of Chiapas, Oaxaca, Guerrero, 
Colima, and Sonora (Tiburén Island). 


Genus CARETTA Rafinesque 


Caretta RAFINESQUE, Specchio Sci. Palermo, vol. 2, 1814, p. 66. 

Thalassochelys Firzincer, Ann. Wien Mus., vol. 1, 1835, p. 121 (type, Testudo 
caouana Daudin). 

Caouana CoctEat, in Ramén de la Sagra, Historia ffsica, politica y natural de 
la Isla de Cuba, vol. 4, Rept., 1838, p. 31 (type, Testudo cephalo Schneider). 

Eremonia Gray, Hand list of shield reptiles of the British Museum, 1873, p. 91 
(type, Caouana elongata Gray). 


Genotyye.—Caretta nasuta Rafinesque=Caretta caretta caretta (Lin- 
naeus). 

Species.—One, with two races, is generally recognized. 

Range.—Tropical ocean, world-wide. 


KEY TO SPECIES OF CARETTA 


1. Neural bones usually 9 or more; if fewer, the costal bones usually interrupting 
the neural series by contact with each other at one or more places. 

caretta gigas (p. 16) 

Neural bones 7 or 8, the series rarely interrupted by median contact of costal 

DONGS ss eee ee ee eee PARES ee ae caretta caretta (p. 16) 


16 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


CARETTA CARETTA CARETTA (Linnaeus) 


Testudo careitta LINNAEUS, Systema naturae, ed. 10, vol. 1, 1756, p. 197. 

Caretta caretta, StEJNEGER, Ann. Rep. U. S. Nat. Mus., 1902 (1904), p. 715.— 
Carr, Proc. New England Zool. Club, vol. 21, 1942, p. 5, pls. 1, 3. 

Caretta caretta caretta, DERANIYAGALA, Tetrapod reptiles of Ceylon, vol. 1, 1939, 
p- 164; Spolia Zeylanica, vol. 24, 1945, p. 95. 

Testudo cephalo SCHNEIDER, Algemeine Naturgeschichte der Schildkréten . . ., 
1783, pp. 303-308 (type unknown; type locality, Charleston, S. C., by 
present restriction). 

Testudo caouana Daunpin, Histoire naturelle des reptiles, vol. 2, 1802, p. 55, pl. 
16, fig. 2 (type in Mus. Hist. Nat. Paris; type locality, Jamaica, by present 
restriction). 


Ty pe.—Unknown. 

Type locality —‘“‘About the American Islands,” here restricted to the 
Bermuda Islands. 

Range.—Western Atlantic Ocean; the entire Atlantic coast of 
Mexico. Reported in Mexico only from Yucatan. 


CARETTA CARETTA GIGAS Deraniyagala 


Caretta gigas DERANIYAGALA, Ceylon Journ. Sci., sect. B, vol. 28, 1933, pp. 61-62. 
Caretta caretta gigas, DERANIYAGALA, Tetrapod reptiles of Ceylon, vol. 1, 1939, 
pp. 164-185, figs. 66-73; Spolia Zeylanica, vol. 24, 1945, p. 95. 


Type.—Presumably in Colombo Museum, Ceylon. 

Type locality.—Ceylon. 

Range.—Indian and Pacific Oceans and possibly eastern Atlantic 
Ocean. In Mexico, the entire Pacific coast. Reported only from 
Sinaloa and Baja California. 


Genus ERETMOCHELYS Fitzinger 
Eretmochelys Firzinerr, Systema reptilium, fase. 1, 18438, p. 30. 
Genotype.—Testudo imbricata Linnaeus. 
Species —One is generally recognized, but two species (or sub- 
species) are distinguished by some authorities. 
ERETMOCHELYS IMBRICATA ® (Linnaeus) 


Testudo imbricata LINNAEUS, Systema naturae, ed. 12, 1766, p. 350. 

Eretmochelys imbricata, Firzincer, Systema reptilium, fase. 1, 1848, p. 30.— 
Acassiz, Contributions to the natural history of the United States, vol. 1, 
1857, p. 381.—Deraniyacaua, Tetrapod reptiles of Ceylon, vol. 1, 1939, 
pp. 187-217, figs. 74-85. 

Chelonia lachrymata Cuvimr, Le régne animal, ed. 2, vol. 2, 1829, p. 13 (no type 
or type locality designated; Bermuda Islands by present restriction). 

Chelonia pseudo-mydas Lesson, in Belanger, Voyage aux Indes Oriental. . ., 
Zool., 1834, p. 299 (type unknown; type locality, Atlantic Ocean, here re- 
stricted to the Bermuda Islands). 

Chelonia pseudo-caretta LEssoN, op. cit., p. 302 (type and type locality as in the 
preceding). 


8 The Atlantic Ocean specimens are frequently regarded as Eretmochelys imbricata (or E. i. imbricata), 
the Pacific Ocean ones as Eretmochelys imbricata bissa (or E. i. squamata). We are unable to determine 
characters for such separation. 


CHECKLIST OF REPTILES OF MEXICO £7 


Caretta bissa Rtrreti, Neue Wirbelthiere Abyssiniens, Amphibiens, 1835, p. 4, 
pl. 2 (type unknown; type locality, Red Sea). 

Eretmochelys squamata AGassiz, Contributions to the natural history of the 
United States, vol. 1, 1857, p. 382 (cotypes, Mus. Comp. Zool. Nos. 1415, 
1416; lectoholotype, No. 1416; type locality restricted to Singapore, Straits 
Settlements). 


Type.—Unknown. 

Type locality—American seas, here restricted to the Bermuda 
Islands. 

Range.—Tropical oceans, world-wide; both coasts of Mexico. 
Reported from Baja California, Sonora, Oaxaca, Campeche, and 
Quintana Roo (Isla de Mujeres). 


Genus CHELONIA Brongniart 


Chelonia BRoneniaAkT, Bull. Sci. Soc. Philom., vol. 2, 1800, p. 89. 

Chelone BRonGN1ART, Mem. Sav. Etrang, vol. 1, 1806, p. 610. 

Chelona BurMEIsTER, Handbuch der Naturgeschichte, vol. 2, 1807, p. 731. 

Chelonias RAFINESQUE, Specchio Sci. Palermo, vol. 2, 1814, p. 66. 

Mydas Cocteau, in Ramén de la Sagra, Historia fisica, politica y natural de la 
Isla de Cuba, vol. 4, 1838, p. 22. 

Mydasea Gervats, Dict. Hist. Nat., vol. 3, 1843, p. 457. 

Euchelonia Tscuup1, Untersuchungen tiber die Fauna Peruana . . ., 1846, p. 22. 

Megemys GistEL, Naturgeschichte des Thierreichs, 1848, p. 8 (type of all above, 
Testudo mydas Linnaeus). 


Genotype-—Testudo mydas Linnaeus. 

Species.—Possibly four forms (species or subspecies) are recogniz- 
able. None of these are well enough known to us to diagnose properly. 

Range.—Tropical oceans, world-wide. 


CHELONIA MYDAS ® (Linnaeus) 


Testudo mydas LINNAEUS, Systema naturae, ed. 10, 1758, p. 197. 

Chelonia mydas, BRoNGNIART, Bull. Sci. Soc. Philom., vol. 2, 1800, p. 89.—ScHweEIc- 
cer, Kénigsberg. Arch. Naturw. Math., vol. 1, 1812, p. 412.—DrRantyra- 
GALA, Tetrapod reptiles of Ceylon, vol. 1, 1939, pp. 218-242, figs. 86-95. 

Testudo viridis ScuNEIDER, Algemeine Naturgeschichte der Schildkréten . . ., 
1782, pp. 299-303 (type unknown; type locality, Charleston, S. C., by present 
restriction). 

Chelonia virgata ScHwEIGGER, Prodromi monographiae cheloniorum . . ., 1814, 
p. 21 (type unknown; type locality, ‘‘Seas of Torrid Zone,” here restricted to 
the Bermuda Islands). 

Chelonia maculosa Cuvier, Le régne animal . . ., ed. 2, vol. 2, 1829, p. 13 (type 
in Mus. Hist. Nat. Paris?; type locality not designated, here restricted to 
Ascension Island). 

Chelonia agassizii Bocourt, Ann. Sci. Nat., ser. 5, vol. 10, 1868, p. 122 (type in 
Mus. Hist. Nat. Paris; type locality, mouth of Rio Nagualate, Guatemala) ; 
Mission scientifique au Mexique . . ., Etudes sur les reptiles, livr. 1, 1870, 
pp. 26-28, pl. 6 (color). 


® A common arrangement restricts mydas to the Atlantic Ocean, agassizii to the Pacific, either as species 
or subspecies. We are unable to determine characters Justifying such an arrangement. 


18 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type.—Unknown. 

Type locality.—Ascension Island. 

Range.—Tropical oceans, world-wide, both coasts of Mexico. 
Reported from Veracruz, Yucatan, Quintana Roo (Isla de Mujeres), 
Oaxaca, Guerrero, Sonora (Tiburén Island), and Clarion Island. 


Suborder TRIONYCHOIDEA Fitzinger 
Trionychoidea FirzincrrR, Neue Classification der Reptilien . . ., 1826, p. 7. 


Families—Two, one of which (Carettochelyidae) is restricted to 
New Guinea. 
Family TRIONYCHIDAE Gray 
Trionicidae Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 212. 


Genera.—Seven, only one of which occurs in the Americas. 
Range.—Asia, Africa, North America. 
Genus AMYDA Schweigger 
Amyda ScuHwEIccER, in Geoffroy, Ann. Mus. Hist. Nat. Paris, vol. 14, 1809, p. 1.— 
Conant and Gorn, Occ. Pap. Mus. Zool. Univ. Michigan, No. 510, 1948, pp. 
11-19 (taxonomy). 

Genotype—Amyda javanica Geoffroy |[=Amyda cartilaginea (Bod- 
daert)]. 

Species.—About 21, of which 4 are American; two or three sub- 
species are recognized of one species (A. spinifera). One occurs in 
Mexico.” 

Range.—Asia, North America. 

AMYDA EMORYI (Agassiz) 


Aspidonectes emoryi Acassiz, Contributions to the natural history of the United 
States, vol. 1, 1857, p. 407; vol. 2, 1857, pl. 6, figs. 4, 5. 

Amyda emoryi, STEJNEGER and Barsour, Check list of North American amphib- 
ians and reptiles, 1917, p. 124. 


Type.—Mus. Comp. Zool. Nos. 1901-1910, cotypes; W. H. Emory 
collector. 

Type locality —Rio Grande River, near Brownsville, Tex. 

Range.—Southern Oklahoma and Arizona southward into northern 
Mexico. Recorded from Coahuila: Hacienda Los Borregos (near 
Judrez), San Juan, Cuatro Ciénegas, Hacienda La Gacha; Nuevo 
Leén: Rodriguez; Tamaulipas: Matamoros, Rio Purificacién north of 
Ciudad Victoria; Baja California?. 


Suborder CRYPTODIRA Cope 


Cryptodira Corn, Proc. Amer, Assoc. Adv. Sci., vol. 19, 1870, p. 235. 


Families.—Six, all but one of which (Platysternidae, southern Asia) 
occur in Mexico. 


10 Miiller (Verh. Naturf. Ges. Basel, vol. 6, 1878, p. 641) records A. mutica also from ‘‘Mexico.” The 
record requires confirmation. 


CHECKLIST OF REPTILES OF MEXICO 19 


KEY TO MEXICAN FAMILIES OF CRYPTODIRA 


1. Abdominal scutes in contact with marginals or separated from them only by 
membrane; inguinal scute short, less than half length of bridge; 12 plastral 

SSC UU SS a ea a cee ee te es a 4 

All plastral scutes separated from marginals by a series of inframarginals, or 
inguinal scute very long, half length of bridge or longer; 12 or fewer plastral 


BCULCS Rare See oe See et Seo 2 ates rye Sintogtel ese 2 

2. Sour orbyaniramarpinals — 2s yoo 22} eb bee ee bere Dermatemyidae (p. 19) 
ao LOn a (rarely: 1) inframarginals =< 2. a 2 ee eS 3 

38. Bridge very narrow, covered chiefly by the displaced abdominal scutes, which 
are widely separated from each other medially_____-_- Chelydridae (p. 20) 
Bridge broad, abdominal scutes in normal position__.._Kinosternidae (p. 21) 

4. Digits with no webs whatever; rear foot stump-shaped, plantar surface as 
Eb AC ss Seri oe 17s eis eS Da Be Deg eg dg Testudinidae (p. 27) 
Digits with at least some vestige of webs; rear foot more elongate, plantar 
muniee longer than broads! 22022 3225 2a ee Sees ee Emyidae (p. 28) 


Family DERMATEMYIDAE Gray 


Dermatemyidae Gray, Supplement to the catalogue of the shield reptiles .. ., 
1870, p. 49. 


Genera.—One. 
Range.—Central Mexico to Honduras. 


Genus DERMATEMYS Gray 


Dermatemys Gray, Proc. Zool. Soc. London, 1847, p. 55. 

Chloremys Gray, Proc. Zool. Soc. London, 1870, p. 715 (type, Dermatemys abnormis 
Cope= D. mawii Gray). 

Limnochelone WERNER, Zool. Anz., vol. 24, 1901, p. 297 (type, Limnochelone 
micrura Werner). 


Genotype.—Dermatemys mawii Gray. 
Range.—Atlantic coast from central Veracruz to Honduras. 
Species.—One recognized. 

DERMATEMYS MAWII Gray 


Dermatemys mawit Gray, Proc. Zool. Soc. London, 1847, p. 55.—BouLENGER, 
Catalogue of the chelonians . . ., 1889, pp. 28-29. 

Emys Berardit Dumféri~ and Brisron, in Duméril and Duméril, Catalogue 
méthodique de la collection des reptiles, livr. 1, 1851, p. 11 (type locality, 
environs of Veracruz, Mexico; type in Mus. Hist. Nat. Paris). 

Dermatemys abnormis Corps, Proc. Acad. Nat. Sci. Philadelphia, vol. 20, 1868, 
p. 120 (type, U.S.N.M. No. 6545; Belize River, British Honduras). 

Limnochelone micrura WERNER, Zool. Anz., vol. 24, 1901, p. 298 (type locality, 
Mexico, here restricted to Alvarado, Veracruz; type unknown). 


Type.— Brit. Mus. 

Type locality —Unknown, here restricted to Alvarado, Veracruz. 

Range.—Rivers of the Atlantic coast from central Veracruz to 
Guatemala, probably excluding the northern part of the Yucatan 
Peninsula. Recorded from the states of Veracruz, Oaxaca, Tabasco, 
Yucatan, and Campeche. 


20 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Family CHELYDRIDAE Swainson 


Chelidridae Swarnson, Natural history and classification of fishes, amphibians, 
and reptiles, vol. 2, 1839, p. 116. 

Genera.—Two, one of which (Macrochelys) is restricted to the 
United States. A supposed New Guinea genus, Devisia, is an errone- 
ously labeled Chelydra serpentina. 

Range.—North America south to northern South America. 


Genus CHELYDRA Schweigger 


Chelydra ScuweEicaEr, Kénigsberg. Arch. Naturw. Math., vol. 1, 1812, p. 292. 

Chelonura FLEMING, Philosophy of zoology . . ., vol. 2, 1870, p. 64 (type, Testudo 
serpentina Linnaeus). 

Rapara Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 211 (type as above). 

Saurochelys LATREILLE, Familles naturelles du régne animal . . ., 1825, p. 92 
(type as above). 

Cheliurus Rarinesqus, Atlantic Journ., 1832, p. 64 (type as above). 

Emysaurus Dumé&rit and Brsron, Erpétologie générale, vol. 2, 1835, p. 548 (type 
as above). 

Chelonura Houtsroox, North American herpetology, vol. 1, 1836, p. 139 (type 
as above). 


Genotype.—Testudo serpentina Linnaeus. 

Range.—Southeastern Canada, United States east of the Rocky 
Mountains, south to Ecuador. 

Species.—Three species or subspecies; one is definitely recorded from 
Mexico, another probably occurs there, and the third (acutirostris) is 
restricted to Panama and South America. 


KEY TO MEXICAN SPECIES OF CHELYDRA 


1. Bridge about one-seventh length of plastron; 4 chin barbels; second vertebral 
shield 32 to 34 percent length of carapace; height of skull at quadrate 44 to 

48 percent length of skull to condyle; width of palatine bone 28 to 30 percent 

heme Ch Ofes ull = So severe ae me eens a ey eg ae es ed rossignonii (p. 20) 
Bridge about one-ninth length of plastron; 2 chin barbels; second vertebral 
shield less than 30 percent length of carapace; height of skull at quadrate 

38 to 43 percent length of skull to condyle; width of palatine bone 19 to 

Ze percent lengthiofiskulliya ta. | epg) ae ee ae serpentina (p. 21) 


CHELYDRA ROSSIGNONII (Bocourt) 


Emysaurus rossignonit Bocourt, Ann. Sci. Nat., ser. 5, vol. 10, 1868, pp. 121-122; 
Mission scientifique au Mexique . . ., Etudes sur les reptiles, livr. 1, 1870, 
pp. 18-19, pl. 5, fig. 2. 
Chelydra rossignonii, BouLENGER, Ann. Mag. Nat. Hist., ser. 7, vol. 9, 1902, pp. 
49-51.—ScumiptT, Smithsonian Mise. Coll., vol. 106, No. 8, 1946, pp. 1-9. 
Type.—Mus. Hist. Nat. Paris. 
Type locality —Panzés, Rio Polochic, Guatemala. 
Range.—Atlantic slopes of Guatemala and presumably adjacent 
Mexico southward to Costa Rica. Not yet reported definitely from 
Mexico. 


CHECKLIST OF REPTILES OF MEXICO 21 
CHELYDRA SERPENTINA (Linnaeus) 


Testudo serpentina LINNAEUS, Systema naturae, ed. 10, vol. 1, 1758, p. 199. 
Chelydra serpentina, SchwEIccrER, Kénigsberg. Arch. Naturw. Math., vol. 1, 
1812, p. 293.—Caun, Illinois Biol. Monogr., vol. 16, 1937, pp. 34-45, pls. 3-4. 

Type.—Unknown. 

Type locality —‘‘Warmer region,” here restricted to New Orleans, 
La. 

Range.—North America east of the Rocky Mountains; in Mexico, 
Atlantic slopes south to the Yucatan Peninsula; recorded only from 
the states of Veracruz and Campeche. 


Family KINOSTERNIDAE Agassiz 


Cinosternidae Acassiz, Contributions to the natural history of the United States, 
vol. 1, 1857, p. 347. 


Genera.—Three. 
Range.—North America, south to northern South America. 


KEY TO GENERA OF KINOSTERNIDAE 


1. Plastron very small, cruciform, with 7-9 scutes; bridge narrow, its length 
(parallel to body axis) exceeded by its width; inframarginals little longer 

GH ATNED TOR See cet ree ee ee ee ee nee: he ee nee © See ETE nee seen 2 
Plastron larger, not cruciform, with 10 or 11 scutes; bridge broader, its length 
much exceeding its width; inframarginals about three times as long as broad, 


OTR OIG Cre ee ee eee ie Peta ae Lee ae nt cad e A Soe a Kinosternon (p. 21) 
2. A ligament between inframarginals and plastron__.__------- Claudius (p. 26) 
A suture between inframarginals and plastron_.___--_-- Staurotypus (p. 27) 


Genus KINOSTERNON Spix 


Kinosternon Sp1x, Ranae et testudinis brasiliensis species novae, 1825, p. 17. 
Swanka Gray, Catalogue of the tortoises . . ., 1844, p. 32 (type, Kinosternon 
scorptoides Gray). 
Platythyra Aaassiz, Contributions to the natural history of the United States, 
vol. 1, 1857, p. 430, pl. 5, figs. 12-15 (type, Platythyra flavescens Agassiz). 
Thyrosternum AGASSIZ, op. cit., p. 429 (type, Kinosternum integrum LeConte). 
Genotype.—Kinosternon longicaudatum Spix=K. scorpioides Lin- 
naeus. 
Range.—United States, except the northwestern quarter, south to 
northern South America. 
Species.—About 22 species and subspecies, 12 of which are known to 
occur in Mexico. 


KEY TO MEXICAN SPECIES OF KINOSTERNON 
1. Plastron rounded behind, not incised or indented; stridulation organs 


(on concealed surfaces of shank and thigh) in males poorly developed 
or absent; plastron large, completely closing shell in adults__________ 2 


22 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Plastron incised or indented behind; stridulation organs in males well 
developed or not; plastron at least somewhat smaller, not completely 


closineyshell Gyen in sac uilibsige sper en ee eee Oe le at a 
2. Carapace tricarinate; axillary and inguinal scutes (if present) widely 
Fo() O75 2h 20 a Meee ers eS, Se AU UR SS ee en ae ie 3 
Carapace flat or unicarinate; axillary and inguinal scutes narrowly in 
contactor marrowly) separated: - sza5o82 eee Oe ae 5 


3. Axillary scute usually absent; posterior edge of abdominals convex. 
abaxillare (p. 22) 
Axillary scute present; posterior edge of abdominals straight_________- 4 
4. Anterior lobe of plastron longer than immovable portion of plastron. 
cruentatum cruentatum (p. 23) 
Anterior lobe of plastron as long as or shorter than immovable portion of 


pDIAstron sR eNa f. _ ON EE IL ee eA cruentatum consors (p. 24) 
5. Anterior lobe of plastron longer than fixed portion___.__..._.________-_-- 6 
Anterior lobe of plastron not longer than fixed portion._.__acutum (p. 23) 


6. Gular more than half length of anterior lobe of plastron__creaseri (p. 23) 
Gular less than half length of anterior lobe of plastron__leucostomum (p. 25) 
7. Carapace tricarinate; stridulation organs absent; plastron slightly smaller 


THAN MO PEDIN GAOL CALA DACE see ae ae a ee ee ae eee integrum (p. 25) 
Carapace flat or unicarinate; stridulation organs well developed in males; 

plastron distinctly smaller than opening of carapace__________-_---- 8 

8. Ninth marginal about as high as tenth, much higher than eighth____-- 9 

Ninth marginal much lower than tenth, about height of eighth______-- 10 


9. Gular more than half length of anterior lobe of plastron (60—64 percent); 
length of anterior lobe less than combined length of gular plus length 
of interfemoral suture plus width of nuchal (83-93 percent). 

flavescens stejnegeri (p. 24) 

Gular half or less than half length of anterior lobe of plastron (80-50 percent) ; 

length of anterior lobe greater than combined length of gular plus length 
of interfemoral suture plus width of nuchal (128-196 percent). 

fiavescens flavescens (p. 24) 

10. Anterior vertebral very narrow, widely separated from second marginal 
on each side; gular less than half length of anterior lobe; posterior marginal 
(eleventh) little lower than penultimate (tenth)____-_- herrerai (p. 24) 

Anterior vertebral broad, in contact with second marginal on each side; 
gular variable in length; posterior marginal distinctly lower than penulti- 
TN LO cote WY eee sh dete NON ape AON AE Na ive 5 pig SEER cc ESE Es Re ee 11 

11. Carapace depressed; entire shell twice as broad as deep; head light, dark- 
spotted; sutures between scutes of carapace not distinctly black-bordered, 
those between plastral scutes not or narrowly marked with brown. 

sonoriense (p. 26) 
Carapace peaked, arched as viewed directly from front; shell less than twice 
as broad as deep; head dark, light-spotted, sutures on carapace narrowly 

but distinctly black-edged, those on plastron broadly marked with brown. 
hirtipes (p. 25) 

KINOSTERNON ABAXILLARE Baur 
Kinosternon abazillare Baur, in Stejneger, Journ. Washington Acad. Sci., vol. 15, 
1925, pp. 462-463. 


Type.—U.S.N.M. No. 7518; C. H. Berendt collector. 
Type locality —Tuxtla Gutiérrez, Chiapas, Mexico. 
Range.—Plateau of Chiapas. Recorded only from the type locality. 


CHECKLIST OF REPTILES OF MEXICO 93 
KINOSTERNON ACUTUM Gray 


Kinosternon scorpioides acuta Gray, Synopsis reptilium, 1831, p. 34, pl. 7, fig. 1. 

Kinosternon acutum, STEJNEGER, Proc. U.S. Nat. Mus., vol. 90, 1941, pp. 457-458. 

Cinosternum berendtianum Corr, Proc. Acad. Nat. Sci. Philadelphia, 1865, p. 189 
(Tabasco; U.S.N.M. No. 6517; C. H. Berendt collector). 

Swanka maculata Gray, Proc. Zool. Soc. London, 1869, p. 162 (type locality, 
Cosamaloapam, Veracruz, Mexico; cotype in Brit. Mus.). 

Cinosternon Effeldtit Peters, Monatsb. Berlin Akad. Wiss., 1873, p. 603, pl. 5, 
figs. 1-3 (type locality, “presumably” Veracruz, Mexico, here restricted to 
Cosamaloapam; type Berl. Mus.). 


Type.—Brit. Mus. 

Type locality—Unknown, here restricted to Cosamaloapam, Vera- 
cruz. 

Range.—Atlantic coast from central Veracruz southward to British 
Honduras, excluding the northern part of the Yucatan Peninsula. 
Recorded from the states of Veracruz, Tabasco, and Campeche." 


KINOSTERNON CREASERI Hartweg 


Kinosternon creasert HARTWEG, Occ. Pap. Mus. Zool. Univ. Michigan, No. 277, 
1934, pp. 1-2. 


Type.—Univ. Mich. Mus. Zool. No. 73090, male; Edwin P. Creaser 
collector. 

Type locality—One mile south of the Hacienda, Chichen Itzé, 
Yucatdn. 

Range.—Known only from the general region of the type locality, 
and Quintana Roo (Vivienda de Platanal). 


KINOSTERNON CRUENTATUM CRUENTATUM Dunméril and Bibron 


Clinosternon] cruentatum Dumérit and Bripron, in Duméril and Duméril, Cata- 
logue méthodique de la collection des reptiles, livr. 1, 1851, pp. 16-17.— 
GtnrTHER, Biologia Centrali-Americana, Reptilia and Batrachia, 1885, pp. 
13-14, pls. 9-11.—S1eBrenrock, Zool. Anz., vol. 30, 1906, pp. 98-99. 

Kinosternon cruentatum cruentatum, WETTSTEIN, Sitzb. Akad. Wiss. Wien, math.- 
nat. Kl., Abth. 1, vol. 143, 1934, pp. 14-15.—Scumipt, Publ. Field Mus. Nat. 
Hist., zool. ser., vol. 22, 1941, p. 488. 

K{inosternum] mexicanum LEConre, Proc. Acad. Nat. Sci. Philadelphia, 1854, 
pp. 182-183 (type locality, Mexico, here restricted to San Mateo del Mar, 
Oaxaca; type presumably in Acad. Nat. Sci. Philadelphia). 

Cinosternum triliratum LeCont#, Proc. Acad. Nat. Sci. Philadelphia, 1859, p. 6 
(type locality, ‘‘Mexico,” here restricted to San Mateo del Mar, Oaxaca; 
Pease coll.; Acad. Nat. Sci. Philadelphia?). 

Cinosternon Shawianum Bocourt (nec Bell), Journ. Zool., vol. 5, 1876, pp. 387, 
397-398. 


Type.—Mus. Hist. Nat. Paris. 
Type locality —‘‘Amér. septentr.,’”’ here restricted to San Mateo del 
Mar, Oaxaca. 


1 Gadow’s record (Proc. Zool. Soc. London, 1905, p. 194) for San Mateo, Oaxaca, is considered erroneous. 


24 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Range.—Atlantic and Pacific drainages in Oaxaca, Tabasco, Campe- 
che, and Chiapas.” In Central America, Guatemala. 


KINOSTERNON CRUENTATUM CONSORS Stejneger 


Kinosternon cruentatum consors STEJNEGER, Proc. U. S. Nat. Mus., vol. 90, 1941, 
pp. 458-459. 
Type.—U.S.N.M. No. 13912. 
Type locality —Cozumel Island, Quintana Roo. 
Range.—The type locality and northern Yucatin (Progreso and 
Telchac Puerto). 


KINOSTERNON FLAVESCENS FLAVESCENS (Agassiz) 


Platythyra flavescens Agassiz, Contributions to the natural history of the United 
States, vol. 1, 1857, p. 430; vol. 2, pl. 5, figs. 12-15.—Cann, Illinois Biol. 
Monogr., vol. 16, Nos. 1-2, 1937, pp. 56-62, pls. 6, 7. 

Kinosternon flavescens flavescens, HARTWEG, Occ. Pap. Mus. Zool. Univ. Michigan, 
No. 371, 1938, pp. 2, 3. 


Type.—Unknown. 

Type locality —Texas and Arizona, here restricted to Waco, Tex. 

Range.—Northern Mexico from Coahuila to Tamaulipas; recorded 
in Coahuila (Jaral). In the United States from Arizona and Texas 
north to Colorado and Kansas. 


KINOSTERNON FLAVESCENS STEJNEGERI Hartweg 


Kinosternon flavescens stejnegert HARTWEG, Occ. Pap. Mus. Zool. Univ. Michigan, 
No. 371, 1938, pp. 1-5. 


Type.—Univ. Michigan Mus. Zool. No. 72235; Morrow J. Allen 
collector. 

Type locality—Llano, Sonora (midway between Nogales and 
Hermosillo). 

Range.—Known only from Sonora (type locality) and Durango 
(Pedricefia) ; perhaps also from southern Coahuila (said to intergrade 
with flavescens flavescens at Jaral, Coahuila). 


KINOSTERNON HERRERAI Stejneger 


Kinosternon herrerai STEJNEGER, Journ. Washington Acad. Sci., vol. 15, 1925, 
p. 462.—SHannon and Smiru, Trans. Kansas Acad. Sci., vol. 52, No. 4, 
1949, pp. 497-498. 

Type.—U.S.N.M. No. 61249; Alfonso Herrera, donor. 
Type locality—Xochimilco, Valley of Mexico, Distrito Federal, 

Mexico, in errore; here restricted to La Laja, Veracruz. 

Range.—Known only from the type locality. 


12. A record from La Majada, near Apatzingén, Michoac4n (Schmidt and Shannon, Fieldiana, vol. 31, 
1947, p. 69), is referable to K. integrum. 


CHECKLIST OF REPTILES OF MEXICO 25 
KINOSTERNON HIRTIPES Wagler 


Cinosternon hirtipes WaGuER, Natiirliches System der Amphibien, . . ., 1830, 
pl. 5, figs. 29, 30. 

Cinosternum hirtipes, GinTHER, Biologia Centrali-Americana, 1885, p. 15, pl. 
12-15.—SIEBENROCK, Zool. Anz., vol. 30, 1906, pp. 94-97, figs. 

Cinosternum henrici LEContE, Proc. Acad. Nat. Sci. Philadelphia, 1859, p. 4 
(type presumably in Acad. Nat. Sci. Philadelphia; type locality, New Mexico, 
T. C. Henry collector).—Ditmars, Reptile book, 1907, p. 26, pl. 11, fig. 


Type.——Munich Museum. 

Type locality —Mexico, here restricted to Mazatlan, Sinaloa. 

Range.—Western Texas and southern Arizona southward through 
the main Mexican Plateau, from Chihuahua to México; recorded 
from the states of Chihuahua, Sinaloa, Michoacan, Colima, Guana- 
juato, San Luis Potosi, Hidalgo, México, and from Distrito Federal 
and the Tres Marias Islands. 


KINOSTERNON INTEGRUM LeConte 


Kinosternum integrum LeConrte, Proc. Acad. Nat. Sci. Philadelphia, 1854, p. 133. 

Cinosternum integrum, BOULENGER, Catalogue of the chelonians, rhynchocepha- 
lians and crocodiles in the British Museum, 1889, p. 42. 

Cinosternum scorpioides integrum, SIEBENROCK, Zool. Anz., vol. 30, 1906, pp. 96-97. 

Cinosternum rostellum Bocourt, Journ. Zool., vol. 5, 1876, pp. 391-392 (type 
locality, Guanajuato, Mexico; type in Mus. Hist. Nat. Paris). 


Type.—Acad. Nat. Sci. Philadelphia; Mr. Pease collector. 

Type locality.—Mexico, here restricted to Acapulco, Guerrero. 

Range.—The plateau of Mexico from Sonora to Oaxaca east to 
Veracruz. Known from Tres Marias Islands and the states of Sonora, 
Sinaloa, Nayarit, Colima, Michoac4n, Guerrero, Oaxaca, Morelos, 
Guanajuato, Aguascalientes, Jalisco, San Luis Potosi, Puebla, and 


Veracruz. 
KINOSTERNON LEUCOSTOMUM Duméril and Bibron 


Clinosternon] leucostomum Dumfrit and Brsron, in Duméril and Duméril, 
Catalogue méthodique de la collection des reptiles, livr. 1, 1851, p. 17, figs. 
1-3.—GtnTHER, Biologia Centrali-Americana, Reptilia and Batrachia, 
1885, p. 17, pls. 16, 17.—SimBENnrRock, Zool. Anz., vol. 30, 1906, pp. 97-98. 

Swanka leucostoma, Gray, Catalogue of the shield reptiles in the British Museum, 
pt. 1, Testudinata, 1855, p. 69. 

Swanka maculata Gray, ibid., p. 68 (type locality, ‘‘ Vera Paz,’’ Guatemala, and 
Cosamaloapam, Veracruz, here restricted to the latter; type in Brit. Mus. 
Nat. Hist.). 

Cinosternum brevigulare GUNTHER, Biologia Centrali-Americana, Reptilia and 
Batrachia, 1885, pp. 17-18 (type locality, Playa Vicente, Mexico, Sallé 
collector; type in Brit. Mus. Nat. Hist.). 

Cinosternum cobanum GUNTHER, op. cit., p. 18, pl. 18, fig. B (type in Brit. Mus. 
Nat. Hist.; type locality, Cob4én and Cahabon, Guatemala, here restricted 
to Cob4n). 


Type.—Mus. Hist. Nat. Paris. 
Type locality—‘N. Orléans; Mexique; Rio-Sumasinta (Amér. 


26 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


centr.): M. Morelet. Amér. septentr. o'9 ?Vallée de la Madeleine 
(N. Grenade), ad. et j. 4ge: M. J. Goudot. Santa-Fé de Bogota 
(N. Grenade): M. Lewy, V. V. Vélins No. 28.” Here restricted to 
Cosamaloapam, Veracruz. 

Range.—Atlantic slopes of southern Mexico south of central 
Veracruz; recorded from the states of Puebla, Veracruz, Tabasco, 
Campeche, and Chiapas; records from Yucat&én and Cozumel Island 
presumably are referable to K. creaseri and K. cruentatum consors. 
Ranges through Central America to northern South America. 


KINOSTERNON SONORIENSE LeConte 


Kinosternum sonoriense LEConTE, Proc. Acad. Nat. Sci. Philadelphia, 1854, p. 184. 

Thyrosternum sonoriense, Acassiz, Contributions to the natural history of the 
United States, vol. 1, 1857, p. 428; vol. 2, pl. 5, figs. 8-11 (Cinosternum 
sonoriense LeConte). 

Kinosternon sonoriense, STEJNEGER, Proc. U.S. Nat. Mus., vol. 25, 1902, p. 149.— 
Van DEnBuRGH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 967-970, 
pls. 120-121. 

?Cinosternon punctatum Gray, Catalogue of the shield reptiles in the British 
Museum, 1855, p. 45, pl. 20ce, figs. 5, 6 (type in Brit. Mus.; ‘‘North America,” 
here restricted to Tucson, Ariz.). 


Type.—Acad. Nat. Sci. Philadelphia. 

Type locality Tucson, Ariz. 

Range.—Sonora, Chihuahua, and Durango (Durango); in the 
United States: southeastern California, southern Arizona, New 
Mexico to western Texas. 


Genus CLAUDIUS Cope 


Claudius Corz, Proc. Acad. Nat. Sci. Philadelphia, 1865, p. 187. 


Genotype.—Claudius angustatus Cope. 
Range.—Restricted to the Atlantic coast from Veracruz to British 
Honduras. 
Species.—One. 
CLAUDIUS ANGUSTATUS Cope 


Claudius angustatus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1865, pp. 187, 
189.—Bocourt, Mission scientifique au Mexique..., Etudes sur les 
reptiles, livr. 1, 1870, pp. 20-21, pl. 4 (color). 

Claudius megalocephalus Bocourt, Ann. Sci. Nat., ser. 5, vol. 10, 1868, p. 122 
(type locality, Mexico, here restricted to Tabasco; type in Mus. Hist. Nat. 
Paris). 


Type.—U.S.N.M. Nos. 6518, 6525. 

Type locality.—Tabasco. 

Range.—Veracruz southward to British Honduras. Recorded in 
Mexico from the states of Veracruz, Tabasco, and Campeche. 


CHECKLIST OF REPTILES OF MEXICO DF 


Genus STAUROTYPUS Wagler 


Staurotypus WaGuER, Natiirliches System der Amphibien . . ., 1830, p. 137. 
Stauremys Gray, Proce. Zool. Soc. London, 1864, p. 127 [genotype, Staurotypus 
(Stauremys) salvinii Gray]. 
Genotype.—Terrapene triporcata Wiegmann. 
Range.—Atlantic slopes of southern Mexico south to Guatemala. 
Species.—Two. 
KEY TO SPECIES OF STAUROTYPUS 


1. Suture between humerals longest; abdominals much broader than long; length 


of bridge 5% or more times in length of plastron.__________ salvinii (p. 27) 
Suture between pectorals longest; abdominals as broad as long; length of 
bridge less than 5% times in length of plastron_________ triporcatus (p. 27) 


STAUROTYPUS TRIPORCATUS (Wiegmann) 


Terrapene triporcata WIEGMANN, Isis von Oken, 1828, p. 364. 

Staurotypus triporcatus, WAGLER, Natiirliches System der Amphibien . . ., 1830, 
pl. 5, figs. 44-45. 

Staurotypus salvinit, Bocourt (nec Gray), Mission scientifique au Mexique . . ., 
Etudes sur les reptiles, livr. 1, 1870, pp. 22-23, pl. 5, fig. 3. 

Claudius pictus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1872, pp. 26-27 (a 
name proposed for the preceding specimen of Bocourt; type locality, Vera Paz, 
Guatemala, here restricted to Alta Verapaz; type presumably in Acad. Nat. 
Sci. Philadelphia). 

Type.—Presumably in the Berlin Mus.; Deppe collector. 
Type locality —Rio Alvarado, Veracruz. 
Range.—Atlantic slopes from Veracruz to Guatemala and British 

Honduras (recorded in Mexico only from the states of Veracruz and 


Tabasco). 

STAUROTYPUS SALVINII Gray 

Staurotypus salvinit Gray, Proc. Zool. Soc. London, 1864, p. 127. 

Staurotypus marmoratus FiscuEr, Archiv fir Naturg., 1872, p. 265, pl. 10 (type 
locality, “‘Mexico,” here restricted to Santa Efigenia, Oaxaca; St. Petersburg 
Mus.). 

Claudius severus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1872, pp. 24-26 (type 
locality, Santa Efigenia, Oaxaca; type presumably in Acad. Nat. Sci. Phila- 
delphia). 

Staurotypus biporcatus SiEBENROCK, Zool. Jahrb., Suppl. 10, No. 3, 1909, p. 438 
(type locality restricted here to that of S. salvinii Gray; type here restricted, 
as lectotype, to that of S. salviniz Gray). 


Type—Brit. Mus. 

Type locality —Huamuchil, Guatemala. 

Range.—Pacific slopes from the Isthmus of Tehuantepec to Guate- 
mala (in Mexico recorded only from Oaxaca). 


Family TESTUDINIDAE Gray 


Testudinidae Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 210. 
861316—50——3 


28 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Genera.—About four, of which one occurs in Mexico. 
Range.—World-wide in tropical and subtropical areas, except the 
East Indies and Australia. 
Genus GOPHERUS Rafinesque 


Gopherus RAFINESQUE, Atlantic Journ., vol. 1, 1832, p. 64. 
Genotype.—Testudo polyphemus Daudin. 
Range.—Southern United States and northern Mexico. 
Species.—Three recognized, two of which occur in Mexico. 
KEY TO MEXICAN SPECIES OF GOPHERUS 


1. Carapace dome-shaped, rounded on top; carapace length about twice shell 


hele a (4y— 5 Gyper ce wit) es ee a berlandieri (p. 28) 
Carapace with flat-topped contour; carapace length more than twice shell 
height (86-48: percent) = Bat esses Ee eS agassizii (p. 28) 


GOPHERUS BERLANDIERI (Agassiz) 


Xerobates berlandiert Acassiz, Contributions to the natural history of the United 
States, vol. 1, 1857, p. 447; vol. 2, 1857, pl. 3, figs. 17-19. 
Testudo berlandiert, BouLENGER, Catalogue of the chelonians, rhynchocephalians 
and crocodiles in the British Museum, 1889, p. 156. 
Gopherus berlandieri, StrsNeGER, North Amer, Fauna, No. 7, 1893, p. 161. 
Type.—U.S.N.M. No. 60, two specimens. 
Type locality —Lower Rio Grande, Tex., here restricted to Browns- 
ville. 
Range.—Nuevo Leén, Tamaulipas, and Coahuila; in United 
States: Southern Texas. 


GOPHERUS AGASSIZII (Cooper) 


Xerobates agassizii CooprEr, Proc. California Acad. Sci., vol. 2, 18638, p. 125. 

Testudo agassizii, BOULENGER, Catalogue of the chelonians, rhynchocephalians 
and crocodiles in the British Museum, 1889, p. 156. 

Gopherus agassizit, SrEINEGER, North Amer. Fauna, No. 7, 1898, p. 161.—Woop- 
BuRY and Harpy, Ecol. Monogr., vol. 18, 1948, pp. 145-200, figs. 1-25. 


Type.—Unknown. 

Type locality— Mountains near Fort Mojave, Calif. 

Range.—Baja California, Sonora including Tiburén Island; in the 
United States, southeastern California and southwestern Arizona 
north into Nevada and Utah. 


Family EMYIDAE ” Gray 
Emyidae Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 210. 


13 From Woodbury and Hardy, Ecol. Monogr., vol. 18, 1948, p. 152. 

14 Dugés’s record (La Naturaleza, ser. 2, vol. 1, 1888, pp. 146-147; vol. 2, p. 479) from Bolsén de Mapimf, 
Chihuahua, cannot be authoritatively allocated. It may be referable to this species. 

18 See Dermochelyidae (p. 13) for a discussion of orthography of family names. The present family has 
as its type the genus Hmys, not the genus Hmyda Gray, 1831, which belongs to the family Trionychidae 
(a homonym, Hmyda Rafinesque, 1815, long overlooked but of earlier date, belongs to the Emyidae, but 
being a substitute for the generic name Hmys, it is not available). 


CHECKLIST OF REPTILES OF MEXICO 29 


Genera.—About 25, of which 6 occur in Mexico. 
Range.—World-wide except Australia and East Indies. 


KEY TO MEXICAN GENERA OF EMYIDAE 


1. Plastron hinged, movable; no inframarginals______________ Terrapene (p. 34) 
Plastron fixed, immovable; inframarginals present______________________ 2 

2. Alveolar surface of upper jaw with a ridge either smooth or toothed, parallel to 
Pabialsentbing ed ger . 2.6. 8s apons see oe UE eet ree 8 dee A es 5 
Alveolar surface of upper jaw ridgeless, smooth__-_-_-___-_____..-_____- 3 


3. A strong interrupted median keel oncarapace, with a marked tubercle at the 
end of each vertebral; a low dorsal crest of enlarged compressed scales on 
tail; bridge much shorter than posterior lobe of plastron. 

Malaclemys (p. 36) 
Dorsal keel on carapace absent or continuous, not tuberculate; no crest 


Baecalessonmvall Shu. Seber et ee heel ey Pt Es A ee oe hy 4 
4. Bridge much shorter than posterior lobe of plastron________ Clemmys (p. 37) 
Bridge longer than posterior lobe of plastron_____________ Geoemyda (p. 29) 


5. Rear of carapace perfectly smoothly rounded, edges of all plates even, no 
notches except a median rear one; a terminal notch flanked on either side by 

a projection on upper jaw, and cutting edge of lower jaw smooth. 
Chrysemys (p. 33) 
Rear of carapace at least somewhat serrate; no terminal notch flanked on 
either side by a projection in upper jaw, or if so cutting edge of lower jaw 
SCUTAlChe eee 8 oh eee oe co eee Soe eee Pseudemys (p. 31) 


Genus GEOEMYDA Gray 


Geoemyda Gray, Proc. Zool. Soc. London, 1834, p. 100.—WetTrTsreEIN, Sitzb. Akad. 
Wiss. Wien, math.-naturw. K1., Abth. 1, vol. 148, 1934, pp. 17-19 (key to 
American species). 

Rhinoclemys Gray, Ann. Mag. Nat. Hist., ser. 3, vol. 12, 1863, p. 182 (genotype, 
Geoclemmys annulata Gray). 


Genotype.—Testudo spenglerit Gmelin. 

Range.—India to Japan and the Malay Archipelago; the coasts 
of central Mexico south into South America. 

Species.—Nineteen species and subspecies are known, nine in the 
Americas, four in Mexico. 


KEY TO MEXICAN SPECIES OF GEOEMYDA 


Perigo WOlOW~ as = a Seen DSTO Sa 2 ee Sot Jos aees areolata (p. 30) 
Bridge black, uniform or with a few yellow markings__________________-_ 2 

2. Upper jaw hooked or smooth; plastron dark with a yellow margin_rubida (p. 30) 
Upper jaw notched; plastron yellow, with a broad dark median stripe____- 3 

3. Carapace almost uniform brown, without distinct markings, its entire border 
strongly bowed, nowhere straight; sides of head in front of eye with usually 

2, sometimes 3, bright horizontal lines___.-___- pulcherrima incisa (p. 30) 
Carapace olive-brown, with bright, yellow-red concentric or hieroglyphic 
pattern of lines on each scale; young with dark, light-centered spots on 
costals; only lateral border of carapace strongly bowed; sides of head in 
front of eye always with 3 lines________ pulcherrima pulcherrima (p. 30) 


30 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 
GEOEMYDA AREOLATA (Duméril and Bibron) 


Emys areolata DuméRiL and BrBron, in Duméril and Duméril, Catalogue métho- 
dique de la collection des reptiles . . ., 1851, p. 10. 
Geoemyda areolata, WETTSTEIN, Sitzb. Akad. Wiss. Wien, math.-nat. K1., Abth. 1, 
vol. 148, 1934, p. 18. 
Type —Mius. Hist. Nat. Paris; M. Morelet collector. 
Type locality. El Petén, Guatemala, here restricted to La Libertad. 
Range.—Coast of central Veracruz south to northern Guatemala 
and British Honduras. Recorded in Mexico from Veracruz, Tabasco, 
Campeche, Yucat4n, and Quintana Roo (Cob&, Cozumel Island). 


GEOEMYDA PULCHERRIMA PULCHERRIMA (Gray) 


Emys pulcherrima Gray, Catalogue of the shield reptiles in the British Museum, 
vol. 1, 1855, p. 25, pl. 25, fig. 1—Bocourt, Mission scientifique au Mexique, 
Etudes sur les reptiles, livr. 1, 1870, pp. 15-16, pl. 7, figs. 1, 1a, 1b (color). 

Geoemyda pulcherrima pulcherrima, WETTSTEIN, Sitzb. Akad. Wiss. Wien, math.- 
nat. K1., Abth. 1, vol. 143, 1934, p. 18. 


Type.—Brit. Mus. Nat. Hist. 

Type locality—Mexico, here restricted to Presidio de Mazatlan, 
Sinaloa. 

Range.—Pacific slopes from southern Sonora to Guerrero and 
probably western Oaxaca. Recorded from the states of Sonora, 
Sinaloa, Nayarit, Colima, and Guerrero. 


GEOEMYDA PULCHERRIMA INCISA (Bocourt) 


Emys incisa Bocourt, Ann. Sci. Nat., vol. 10, 1868, p. 121; Mission scientifique 
au Mexique . . ., Etudes sur les reptiles, livr. 1, 1870, pp. 11-13, pls. 1, 2. 

Geoemyda pulcherrima incisa, WeTTSTHIN, Sitzb. Akad. Wiss. Wien, math.-nat. 
Kl., Abth. 1, vol. 148, 1934, p. 18. 


Type.—Mus. Hist. Nat. Paris. 

Type locality —lLa Unién, El Salvador. 

Range.—Pacific slopes of the Isthmus of Tehuantepec southward 
to at least El Salvador. Recorded from Oaxaca and Chiapas. 


GECEMYDA RUBIDA (Cope) 


Chelopus rubida Cops, Proc. Amer. Philos. Soc., vol. 11, 1870, p. 148. 

Geoemyda rubida, StepeENRock, Zool. Jahrb., Suppl. 10, Heft 3, 1909, p. 500. 

Rhinoclemmys mexicana Gray, Supplement to the catalogue of shield reptiles 
. . -, 1870, p. 30 (type locality, “San Juan del Rio”; type in Brit. Mus. Nat. 
Hist.; Rebouch collector). 


Type.—Unknown. 

Type locality —Juchitan, Oaxaca. 

Range.—The Pacific coast from Colima to the Isthmus of Tehuan- 
tepec (recorded from Colima, Michoacan, Guerrero, and Oaxaca"). 


16 The exact location of San Juan del Rio (type locality of Rhinoclemmys mexicana) is uncertain; it prob- 
ably is one of the two cities of that name in the state of Oaxaca, 


CHECKLIST OF REPTILES OF MEXICO 31 


Genus PSEUDEMYS Gray 


Pseudemys Gray, Proc. Zool. Soc. London, 1855 (1856), p. 197. 


Type.—Testudo concinna LeConte. 

Species.—About 8, and a total of about 22 species and subspecies, 
of which 8 forms occur in Mexico. 

Range.—The United States east of the Rocky Mountains, Baja 
California, both coasts of Mexico from the Rio Grande on the east 
and southern Sonora on the west southward to Argentina. 


KEY TO MEXICAN SPECIES OF PSEUDEMYS 


1. Ridge on alveolar surface of upper jaw distinctly toothed; temporal stripe 


Absent.oOr HObiexPpandede san 2a steam a ae eee ee eee ee ee 2 
Ridge on alveolar surface of upper jaw not toothed; temporal stripe or spot 
muchyexpandeds.f42 Alessia) Sree? SS Seek 2 ene SR AEL 3 


2. Astrong notch at apex of upper jaw, flanked on each side by a toothlike projec- 
tion; teeth on alveolar ridge long, numerous; nostrils terminal; snout not 
MOIMtEG, +O btUSe==saee se Ao a Oe ee eee ane floridana texana (p. 33) 

No notch at apex of upper jaw, and no subterminal toothlike projections; teeth 
on alveolar ridge smaller, less numerous; nostrils pierced below and posterior 


to tip of snout; snout pointed, acuminate____-_.__.-._-__-- umbra (p. 33) 
3. A light, round temporal spot, completely isolated from orbital markings__ 4 
Supratemporal light spot elongate, reaching orbit__-__------------------ 5 


4. Temporal light spot completely isolated, not continuous with lines either 
anteriorly or posteriorly; bridge with longitudinal black stripes. 
scripta gaigeae (p. 33) 
Temporal light spot continuous toward rear with longitudinal light stripe; 
bridge. biack spottedes-- 2-2) es eee oe Seo scripta nebulosa (p. 32) 
5. Markings solid, intensely black, replacing or obscuring a linear juvenile pat- 
tern, and including a broad area along middle of plastron, lengthwise along 
middle of bridge, and on posterolateral corners of the marginal scutes above 
and below; these markings evident in bony plates as well as in epidermal 
SCULCS SRE Betas Se ee See eee ee Se ee scripta hiltoni (p. 32) 
Markings notras described soit > MEIER PeRNa seis at ih ohn il Te eee ee SY = 6 
6. Plastral markings always present, either smudged, or, if in lines, the lines 
forming ocelli; supratemporal stripe, if visible, reaching eye. 
scripta elegans (p. 32) 
Plastral markings, if visible, not smudged, but in lines that tend to be arranged 
longitudinally, not ocellate; supratemporal stripe reaching eye or not___ 7 
7. Dark ocellus of each costal scute nearest latter’s posteroventral corner. 
scripta cataspila (p. 32) 
Dark ocelli nearest posterodorsal corners of costals____scripta ornata (p. 31) 


PSEUDEMYS SCRIPTA ORNATA (Gray) 


Emys ornata Gray, Synopsis reptilium, 1831, p. 30.—Bocourt, Mission scientifi- 
que au Mexique .. . , Etudes sur les reptiles, livr. 1, 1870, pl. 3, fig. 1, la. 

Pseudemys scripta ornata, Carr, Herpetologica, vol. 1, 19388, p. 135, figs. 3, 4, 5. 

Emys venusta Gray, Catalogue of the shield reptiles . . . , vol. 1, 1855, p. 24, 
pl. 24a (type locality, Mexico, Honduras and America, here restricted to 
Honduras; types in Brit. Mus.). 


Type.—Brit. Mus., two cotypes. 


32 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type locality —Mazatlan, Sinaloa. 

Range.—Pacific slopes of Mexico southward from southern Sinaloa, 
and both Atlantic and Pacific slopes (presumably) of Central America 
to Panama and possibly northern South America; Atlantic slopes of 
Mexico east of the Isthmus of Tehuantepec (recorded in Mexico from 
the states of Sinaloa, Jalisco, Oaxaca, Chiapas, Tabasco, Campeche, 
and Yucatan, and from Cozumel Island). 


PSEUDEMYS SCRIPTA NEBULOSA (Van Denburgh) 


Chrysemys nebulosa VAN DENBURGH, Proc. California Acad. Sci., ser. 2, vol. 5, 
1895, p. 84, pls. 4-6. 
Pseudemys scripta nebulosa, Carr, Amer. Mus. Nov., No. 1181, 1942, p. i. 


Type.—Destroyed. 
Type locality —tLos Dolores, Baja California, Mexico. 
Range.—The southern half of Baja California. 


PSEUDEMYS SCRIPTA HILTONI Carr 
Pseudemys scripta hiltont Carr, Amer. Mus. Nov., No. 1181, 1942, pp. 1-4, fig. 1-3. 


Type.— Amer. Mus. Nat. Hist. No. 63747; John W. Hilton collector. 

Type locality—Guirocoba, 28 miles southeast of Alamos, Sonora, 
Mexico. 

Range.—Known only from the type locality; presumably occurs in 
northern Sinaloa as well as in Sonora. 


PSEUDEMYS SCRIPTA CATASPILA (Giinther) 


Emys cataspila GUNTHER, Biologia Centrali-Americana, Reptilia and Batrachia, 
1885, p. 4, pl. 6, fig. B. 

Pseudemys scripta cataspila, Carr, Herpetologica, vol. 1, 1938, p. 135. —Hartwse, 
Copeia, 1939, No. 1, p. 55. 

? Emys callirostris Gray, Catalogue of the shield reptiles in the British Museum, 
vol. 1, 1855, p. 25, pl. 12B (type locality uncertain, here restricted to Alva- 
rado, Veracruz; type in Brit. Mus.). 


Type.—Brit. Mus., seven cotypes. 

Type locality —Mexico, here restricted to Alvarado, Veracruz. 

Range.—Atlantic slopes of Mexico from the northern border to the 
Isthmus of Tehuantepec (recorded from the states of Tamaulipas 
and Veracruz). 


PSEUDEMYS SCRIPTA ELEGANS (Wied) 


Emys elegans WiEpD, Reise durch Nord-America, vol. 1, pt. 4, 1838, p. 213. 

Pseudemys elegans, Copr, U. 8. Nat. Mus. Bull. 1, 1875, p. 53. 

Chrysemys scripta elegans, SIEBENROCK, Zool. Jahrb., Suppl., vol. 10, 1909, pp. 
464-465. 

Pseudemys troostii elegans, STEJNEGER and Barpour, Check list North American 
amphibians and reptiles, ed. 4, 1939, p. 167.—Scumipt and Owens, Publ. 
Field Mus. Nat. Hist., zool. ser., vol. 29, 1944, p. 101. 

Pseudemys scripta elegans, CAGLE, Copeia, 1944, No. 2, p. 105. 


CHECKLIST OF REPTILES OF MEXICO 33 


Type.—Unknown. 

Type locality —Fox River at New Harmony, Ind. 

Range.—Northeastern Mexico: Coahuila, Nuevo Leén, and Tama- 
ulipas. Widely distributed in central and southern United States. 


PSEUDEMYS SCRIPTA GAIGEAE " Hartweg 


Pseudemys scripta gaigeae Hartweea, Oce. Pap. Mus. Zool. Univ. Michigan, 
No. 397, 1939, pp. 1-4. 


Type—Univ. Michigan Mus. Zool. No. 66472; Helen T. Gaige 
collector. 

Type locality—Rio Grande River, Boquillas, Brewster County, 
Tex. 

Range.—Northern central Mexico (recorded from Coahuila and 
Durango). 

PSEUDEMYS UMBRA (Bocourt) 

Emys grayi Bocourt, Ann. Sci. Nat., ser. 5, vol. 10, 1868, p. 121; Mission scien- 
tifique au Mexique . . ., Etudes sur les reptiles, livr. 1, 1870, pp. 13-15, 
pl. 3, fig. 2, 2a. 

Emys umbra Bocourt, in O’Shaughnessy, Zool. Rec., vol. 13, 1876 (1878), Rept., 
p. 6 (substitute name for Hmys grayi Bocourt, preoccupied by Emys grayt 
Giinther =—Clemmys caspica caspica [Gmelin]).—GuntTuer, Biologia Cen- 
trali-Americana, Reptilia and Batrachia, 1885, pp. x, 5, pls. 4, 5, 6 (fig. A). 

Type.—Mus. Hist. Nat. Paris. 

Type locality —Rio Nagualate, Guatemala. 

Range.—Pacific slopes of the Isthmus of Tehuantepec, southeast- 
ward to Guatemala. Recorded definitely only from the state of 
Oaxaca; other records from Chiapas may refer either to this or to 
P. s. ornata. 





PSEUDEMYS FLORIDANA TEXANA Baur 
Pseudemys texana Baur, Proc. Amer. Philos. Soc., vol. 31, 1898, p. 223. 
Pseudemys floridana texana, Carr, Copeia, 1938, p. 108. 
Type.—Acad. Nat. Sci. Philadelphia No. 246. 
Type locality —San Antonio, Tex. 
Range.—Central Texas and northeastern Mexico. Recorded only 
from Nuevo Leon. 
Genus CHRYSEMYS Gray 


Chrysemys Gray, Catalogue of the tortoises . .. , 1844, p. 27. 
Genotype.— Testudo picta Schneider. 
Range.—Eastern three-fourths of the United States, extreme north- 
ern Mexico. 
Species.—One, with four subspecies. One race occurs in Mexico. 
17 Schmidt and Owens (Publ. Field Mus. Nat. Hist., zool. ser., vol. 29, 1944, p. 101) regard this as a species 
distinct from that to which elegans belongs, but the distinguishing criterion mentioned appears to be one 


that in reality separates males from females of either form. We have not examined the specimens but re- 
gard all Pseudemys reported by Schmidt and Owens as P. s. elegans. 


34 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


CHRYSEMYS PICTA BELLII (Gray) 


Emys bellit Gray, Synopsis reptilium . . . , 1831, p. 31. 
Chrysemys picta bellii, BisHop and Scumipt, Publ. Field Mus. Nat. Hist., zool. 
ser., vol. 18, 1931, p. 136. 
Type.—Lost; originally in Brit. Mus. 
Type locality —Unknown, here restricted to Manhattan, Kans. 
Range.—Western Illinois to Washington and British Columbia, 
south on the east of the Cascade Mountains to northern Chihuahua 
and southern Texas. Recorded only from the state of Chihuahua: 
Rio Santa Maria, near Progreso. 


Genus TERRAPENE Merrem 


Terrapene MERREM, Tentamen systematis amphibiorum, 1820, p. 27.—DitTmars, 
Zoologica, vol. 17, 1934, pp. 1-44, figs. 1-41. 

Type.— Testudo clausa Gmelin= Terrapene carolina (Linnaeus). 

Range.—North America east of the Rocky Mountains and Arizona, 
south to Yucatén and Nayarit. 

Species.—Eleven species and subspecies are recognized at present, 
six in Mexico. 

KEY TO MEXICAN SPECIES OF TERRAPENE 


1. Plastron with a conspicuous and well-defined pattern of yellow lines on a dark 


background (or dark lines on a yellow background).--__-_-- ornata (p. 36) 
Plastron without a lined pattern, either uniform or suffused here and there 
With, dark ‘colori=_ 222226 ta eds OS oy tel et Oe pe pe ie Le See 2 


2. Pattern of carapace consisting of small, round or ovoid yellow dots 1.5—2 mm. 
in diameter, separated from each other by an average distance of 4 mm. 

klauberi (p. 35) 

Pa tterm MOL eSOS 2 ce ee ee a mete a eta ee a ear 3 

3. Width of carapace 65-68 percent length of carapace, average 66 percent; height 

of shell 40-45 percent length of carapace; 4 rear claws; carapace horn-colored, 

Withoutmarks borders atisubuness=) =e aae es eee ee coahuila (p. 36) 

Width of carapace 68-79 percent length of carapace, average about 73 percent; 

height of shell 45-58 percent length of carapace, average 51 percent; rear 

claws 3 or 4; carapace either marked with yellow or horn-colored, but sutures 


always broadly margined with darkicolorz222) 202 te) 222 eee + 

4. No notch at apex of upper jaws; 4 rear claws_____--_-_------ nelsoni (p. 35) 
Apex of upper jaw. notched: rear claws 3.01 42 22. )222 0-22 ee 5 

5. Rear claws always (so far as known) 4___-___-_-_- mexicana yucatana (p. 35) 


Rear claws generally 3 (81 percent of 22 specimens). 
mexicana mexicana (p. 34) 


TERRAPENE MEXICANA MEXICANA (Gray) 


Cistudo (Onychotria) mexicana Gray, Proc. Zool. Soc. London, 1848 (Feb. 1849), 
vol. 16, pp. 16-17, pl. 2. 

Cistudo mexicana, Gray, Catalogue of shield reptiles in the British Museum, pt. 
1, Testudinata, 1855, p. 40. 

Onychotria mexicana, Ducks, La Naturaleza, ser. 2, vol. 1, 1888, pp. 107-108. 

Cistudo carolina var. mexicana, BOULENGER, Catalogue of the chelonians, rhyn- 
chocephalians and crocodiles in the British Museum, 1889, p. 118. 


CHECKLIST OF REPTILES OF MEXICO 35 


Terrapene mexicana, Baur, Amer. Nat., vol. 27, 1893, p. 677.—DuiTMaRs, 
Zoologica, vol. 17, 1934, pp. 18-19, fig. 28—Mtxtuer, Zool. Anz., vol. 
113, 1936, pp. 97-114, figs. 1-4. 

Terrapene mexicana mexicana, SmirH, Publ. Field Mus. Nat. Hist., zool. ser., 
vol, 24, 1939, pp. 17-18. 

Terrapene goldmant STEJNEGER, 18 Proc. Biol. Soc. Washington, vol. 46, 1933, 
pp. 119-120 (type, U.S.N.M. No. 46251; Chijol or Chijoles, southeastern 
corner of the state of San Luis Potosf, Mexico; E. W. Nelson and E. A. 
Goldman collectors).—Ditmars, Zoologica, vol. 17, 1934, pp. 20-22, figs. 
34-36. 

Terrapene yucatana, Dirmars (nec Boulenger), Zoologica, vol. 17, 1934, figs. 29, 
30. 


Type.—Brit. Mus. Nat. Hist.; two female cotypes; collector un- 
known. 

Type locality—Mexico, restricted by Miller to Tampico, Tamauli- 
pas. 

Range.—Atlantic slopes from central Tamaulipas and eastern 
Nuevo Leén to northern Veracruz. Known only from San Luis 
Potosi, Veracruz, and Tamaulipas. 


TERRAPENE MEXICANA YUCATANA (Boulenger) 


Cistudo yucatana BouLEeNGER, Ann. Mag. Nat. Hist., ser. 6, vol. 15, 1895, p. 330. 
Terrapene yucatana, SrrBENROocK, Zool. Jahrb., Suppl., vol. 10, 1909, p. 492. 
Terrapene mexicana yucatana, Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 
24, 1939, pp. 17-18. 

Type.—Brit. Mus. Nat. Hist. 

Type locality—Northern Yucatan, here restricted to Chichen Itza. 

Range.—Northern part of the peninsula of Yucatan. Recorded 
only from Yucatén and Quintana Roo. 


TERRAPENE NELSONI Stejneger 


Terrapene nelsont STEJNEGER, Journ. Washington Acad. Sci., vol. 15, 1925, p. 
463.—Ditmars, Zoologica, vol. 17, 1934, pp. 19-20, figs. 31-33. 


Type —U.S.N.M. No. 46252; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality Pedro Pablo, Tepic [= Nayarit], Mexico, 2,500 feet 
altitude. 

Range.—Known only from the type locality. 


TERRAPENE KLAUBERI Bogert 


Terrapene klaubert Bocert, Amer. Mus. Nov., No. 1226, 19438, pp. 1-4, figs. 1-13. 


18 Schmidt and Owens (Publ. Field Mus. Nat. Hist., zool. ser., vol. 29, 1944, p. 103) regard T. goldmani 
as a distinct species ‘‘of lowland San Luis Potosf,’’ implying that its closest relative, T’. mexicana, is restricted 
to other areas, presumably highland Mexico inasmuch as lowland San Luis Potosi is scarcely distinguish- 
able faunistically from northern Veracruz, southern Tamaulipas, and eastern Nuevo Leén. The entire 
area, as a matter of fact, is small. However, in spite of a few records of occurrence of 7. mexicana on the 
plateau, notably by Dugés (loc. cit.), we do not regard any of them as reliable, and believe the species is 
restricted to the approximate range here stated. There can, therefore, be no geographical isolation of two 
populations to consider. 


36 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type-—Amer. Mus. Nat. Hist. No. 63751; John W. Hilton col- 
lector. 

Type locality—Rancho Guirocoba, about 18 miles southeast of 
Alamos, Sonora. 

Range.—Known only from the type locality and Sierra de Batuc, 
8 kilometers northeast of Matapé, Sonora. 


TERRAPENE COAHUILA Schmidt and Owens 


Terrapene coahuila ScamMipt and Owens, Publ. Field Mus. Nat. Hist., zool. ser., 
vol. 29, No. 6, 1944, pp. 101-103. 


Type.—Chicago Nat. Hist. Mus. No. 41234; Ernest G. Marsh 
collector. 

Type locality —Quatro Ciénegas, Coahuila, Mexico. 

Range.—Known only from the type locality. 


TERRAPENE ORNATA (Agassiz) 


Cistudo ornata Acassiz, Contributions to the natural history of the United States, 
vol. 1, 1857, p. 445. 

Terrapene ornata, Baur, Science, vol. 17, 1891, p. 191—Durmars, Zoologica, vol. 
17, 1934, pp. 16-18, figs. 24-27. 

Cotypes—U.S.N.M. Nos. 7541, 7542, 7547(?); Mus. Comp. Zool. 
No. 1536, Burlington, Iowa; J. Rauch collector. 

Type locality—The upper Missouri and Jowa, here restricted to 
Council Bluffs, Iowa. 

Range.—Widespread in the United States between the Rocky 
Mountains and the Mississippi River reaching east to Indiana and 
west to Arizona; extreme northern Mexico. Reported only from 
Sonora and Chihuahua. 


Genus MALACLEMYS Gray 


Malaclemys Gray, Catalogue of the tortoises . . ., 1844, p. 28. 


Genotype.—Testudo concentrica Shaw. 

Range.—Atlantic and Gulf coasts of the United States and Mexico. 

Species.—One, of five distinct races, only one of which has been 
found in Mexico. 


MALACLEMYS TERRAPIN LITTORALIS Hay 


Malaclemmys littoralis Hay, Bull. U. S. Bur. Fish., vol. 24, 1904, p. 18, pls. 8, 9, 
12, figs. 2, 3. 
M[alaclemmys] terrapin littoralis, CARR, Copeia, 1946, p. 172. 
Type.—U.S.N.M. No. 33913; W. P. Hay collector. 
Type locality —Rockport, Tex. 
Range.—Coast of Texas southward possibly to the Yucat&n Penin- 
sula. Recorded only from “Mexico” (Strauch). 


CHECKLIST OF REPTILES OF MEXICO oF 


Genus CLEMMYS Ritgen 


Clemmys Riteren, Nova Acta Acad. Leopold.-Carol., vol. 14, pt. 1, 1828, p. 272. 


Genotype.—Testudo punctata Schoepff [= Clemmys guttata (Schnei- 
der)]. 

Range.—Southern Europe, northwestern Africa, southern China 
and Japan, extreme western and the eastern third of the United States. 

Species.—Fifteen species and subspecies are recognized, five in 
North America, one of which enters Mexico. 


CLEMMYS MARMORATA PALLIDA Seeliger 
Clemmys marmorata pallida SEELIGER, Copeia, 1945, pp. 158-159. 


Type.—Mus. Vert. Zool. No. 6716; J. E. Law collector. 

Type locality.—Lower Coyote Creek, near Alamitos, Orange County, 
Calif. 

Range.—Central California near the mouth of the Sacramento River 
south to northwestern Baja California. Recorded only from northern 
Baja California. 


Subclass LEPIDOSAURIA Romer 


Lepidosauria Romer, Vertebrate paleontology, ed. 2, 1945, p. 595. 


Orders.—Two orders are represented by living members; one (Rhyn- 
chocephalia) is restricted to certain islands off North Island, New Zea- 
land, and the other is world-wide in distribution. 


Order SQUAMATA Oppel 


Squamata OppEL, Die Ordnungen, Familien und Gattungen der Reptilien ..., 
1811, p. 14. 


Suborders—Three living suborders are recognized: the Sauria, Ser- 
pentes, and Amphisbaenia. 


Suborder AMPHISBAENIA Gray 


Amphisbaenia Gray, Catalogue of the tortoises, crocodilians and amphisbaenians 
in British Museum, 1844, p. 68 (as an order). 

Families —Two, only one of which occurs in Mexico. When the 
anatomy of the forms of amphisbaenians is better known, it is probable 
that several other families will be recognized. 

Range.—West Indies, tropical parts of South America north to 
Arizona, Florida, and Baja California; Spain, Turkey through Africa. 


Family BIPEDIDAE Stejneger 
Bipedidae STEJNEGER, Science, vol. 21, 1905, pp. 157-158. 


Genera.—One. For the present we prefer to refer all Mexican forms 
to the single genus Bipes despite the differences in digital structure. 


38 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Studies on the cranial anatomy may reveal differences not now 
apparent. 

Range.—The southern tip of Baja California, Arizona, the Rio 
Balsas Valley, and southern Guerrero. 


Genus BIPES Latreille 


Bipes LATREILLE, Histoire naturelle des reptiles, vol. 2, 1802, p. 90. 

Bimanus Oprret, Die Ordnungen, Familien und Gattungen der Reptilien . . ., 
1811, p. 18. 

Hemichirotes Ducks, in Cope, Amer. Nat., vol. 28, 1894, p. 436 (type, Hemichirotes 
tridactylus Dugés). 

Euchirotes Cop, ibid. (type, Euchirotes biporus Cope). 


Genotype.—Bipes canaliculatus Bonnaterre. 


Range.—Baja California, Guerrero, southeastern Arizona. 


Species.—Three. 
KEY TO SPECIES OF BIPES 


MMLC S el oui se a i A SS es a gps cee ee a che ge se 2 
DDT OF ESE S| uC] Aw Od aa as ee mi es tridactylus (p. 38) 

2. All digits equally developed, all generally with claws; 6 preanal pores. 
canaliculatus (p. 38) 
One digit smaller and lacking claw; 2 preanal pores__.____-_-_- biporus (p. 38) 


BIPES TRIDACTYLUS (Dugés) 


Hemichirotes tridactylus Ducis, in Cope, Amer. Nat., vol. 28, 1894, pp. 486-4387, 
fig. 6; La Naturaleza, ser. 2, vol. 2, 1894, pp. 411-412, pl. 20. 
Bipes tridactylus, SmirH and Necker, Anal. Ese. Nac. Cien. Biol., vol. 3, 19438, 
pp. 196-197. 
Type.—Alfredo Dugés Mus., Guanajuato, Guanajuato, Mexico. 
Type locality —Tecpan de Galeana, Guerrero. 
Range.—Known only from type locality (only the type specimen 
known). 


BIPES BIPORUS (Cepe) 


Euchirotes biporus Corr, Amer. Nat., 1894, p. 487, figs. 5-5e; Ann. Rep. U.S. Nat. 
Mus., 1898 (1900), pp. 680-682, fig. 140. 

Bipes biporus, STEJNEGER and Barsoour, Check list of North American amphibians 
and reptiles, 1917, p. 72. 


Type.—U.S.N.M. No. 8568, cotypes (?), T. H. Street collector, and 
No. 12599, 12 specimens, L. Belding collector. 

Type locality —Cape San Lucas, Baja California. 

Range.—The cape region of southern Baja California. Reported 
from La Paz, Cape San Lucas. 


BIPES CANALICULATUS Bonnaterre 


Bitpes] canaliculatus BONNATERRE, Encyclopédie méthodique, Erpétologie, 1789, 
p. 68, pl. 12. 

Bipes canaliculatus, SonNINI and LATREILLE, Histoire naturelle des reptiles, vol. 2, 
1801, p. 90.—Smitx, Journ. Washington Acad. Sci., vol. 39, 1949, pp. 42-43. 


CHECKLIST OF REPTILES OF MEXICO 39 


Chirotes canaliculatus, MrRrREM, Tentamen systematis amphibiorum, 1820, p. 
161.—Bocourt, Mission scientifique au Mexique . . ., Etudes sur les reptiles, 
livr. 7, 1881, pp. 487-488, pl. 21C, figs. 8, 8a-c.—BouLENGER, Catalogue of 
the lizards in the British Museum, vol. 2, 1885, p. 432. 

Lacerta lumbricoides SHaw, Naturalists’ Miscellany, vol. 6, 1795, pl. and text No. 
212 (type locality, Mexico, here restricted to Mexcala, Guerrero; type origi- 
nally in British Museum). 

Lacerta mexicana DonnovorrFr, Zool. Beitrige zur 13. Ausg. Linneschen Natur- 
system, vol. 3, 1798, p. 135 (here restricted to Mexcala, Guerrero). 

Chalcides sulcata Suckow, Anfangsgriinde der theoret. und angewandten Natur- 
geschichte der Thiere, vol. 3, 1798, p. 147 (here restricted to Mexcala, 
Guerrero). 

Chamaesaura propus SCHNEIDER, Historia amphibiorum naturalis et literaria, 
fasc. 2, 1801, p. 211 (here restricted to Mexcala, Guerrero). 


Type.—Not known. 

Type locality—Mexico, here restricted to Mexcala, Guerrero. 

Range.—Known in the basin of Rio Balsas, Guerrero. Reported 
from Mexcala, Tecuaiziapan, and Balsas.” 


Suborder Saurra Macartney 


Sauria MacaRTNEY, in Ross’s translation Cuvier’s Lecons d’anatomie comparée, 
vol. 1, 1802, table 3. 


Families —Nineteen, of which 10 occur in Mexico. 
KEY TO MEXICAN FAMILIES OF SAURIA 


1. Neither forelegs nor hind legs evident externally____..__.----__------- 2 
Both forelegs and hind legs visible externally___._____._------.--_----- 3 

2. No eye opening; eye scarcely or not visible below lateral head scales; one 
unpaired median scale between frontal and rostral_Anelytropsidae (p. 170) 

Eye opening present, although small; a pair of scales on dorsal surface of head 


between frontal and rostral_...................--.- Anniellidae (p. 208) 

3. No movable eyelids; eyes permanently open_____-._-__-_------------- 4 
Pyelids {present «MO VADIOS 23 Ss eee oO ee CU in eo ee A ott 6 

4. All scales around middle of body uniform, flat, cycloid, smooth, in about 13 
TO WV ee Ee NN a ee Els SO Ye ee Teiidae » (p. 170) 

INGE SOs RGN 6. LiePOE RV EA FEES? sya aR hy Sa See Ae ee he 5 


5. Head covered with large, flat plates; ventral scales large, quadrangular, 

abruptly differentiated from granular lateral scales__Kantusiidae (p. 151) 

Dorsal surface of head covered with granular scales; ventral scales smaller, 
rounded, not abruptly differentiated from granular lateral scales. 

Gekkonidae (p. 40) 

6. All scales around middle of body perfectly smooth (sometimes finely striated), 

cycloid, more or less equal in size (those in the middorsal row or pair or 


7. Only one scale in contact with frontal anteriorly; five supraoculars. 
Anguidae #! (p. 194) 


19 A record for ‘*Morelia?,’’ Michoac4n (Dugés, La Naturaleza, ser. 2, vol. 2, 1896, p. 480), is unacceptable 
until verified. 

2 Gymnophthaimus only. 

21 Diploglossus only. 


40 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Two or three scales in contact with frontal anteriorly; three or four supra- 


ocularsy j25528) -.. aa ateett see Se eee eee ae Scincidae (p. 155) 

8. A granular fold along sides of body, abruptly differentiated from much 
enlarged dorsal and ventral scales__-_.__.------------- Anguidae (p. 194) 

No tmoneuts= See hee Peel eet Ee Mee eae Sa Se ees Seay eee ee 9 


9. Ventral scales large, quadrangular, in 8 longitudinal series; dorsals granular. 

Teiidae (p. 170) 

Ventral scales smaller, in more numerous rows_---------------------- 10 

10. Ventral scales quadrangular; scales on top of head small, more or less uniform, 

tubercular; dorsal scales granular with numerous enlarged tubercles__ 11 

Ventral scales pointed or rounded, not quadrangular; body and head scales 

as described Or nots 222.2 22026 cee eases as seers eae eee 12 

11. Digits on hind leg of nearly equal length; enlarged tubercles covering most of 

dorsal surface of body; a series of about four large, paired postmentals on 

midline of chin, immediately back of mental__-_-__- Helodermidae (p. 192) 

Digits on hind leg varying greatly in length, fourth toe three times as long 

as fifth; enlarged tubercles on body much less numerous, not occupying as 

great an area as granules; no enlarged postmentals__Xenosauridae (p. 207) 

12. Head and most of body, except belly, covered with very minute granules; no 
parietal “eye’’; no keels or tubercles along ventral surfaces of digits. 

Gekkonidae (p. 40) 

Head covered with larger scales; a parietal ‘‘eye”’ usually visible; at least 

one tubercle, or several keels, on most or all of lamellae on ventral surfaces 

CO) C0 Oa Smee age gh ee at a Olea en Re gets Iguanidae (p. 538) 


Family GEKKONIDAE Stejneger 


Geckonidae BouLENGsER, Catalogue of the lizards in the British Museum, vol. 
1, 1885, p. 3. 
Gekkonidae StrsJnEGgER, U. 8. Nat. Mus. Bull. 58, 1907, p. 164. 


Subfamilies.—Studies by Noble (Amer. Mus. Nov., No. 4, 1921, 
pp. 1-16) and by M. A. Smith (Rec. Indian Mus., 1933, p. 16) indi- 
cate that neither the Eublepharidae nor the Uroplatidae are worthy 
of either family or subfamily distinction. 

Genera.—About 75 genera are commonly recognized. 

Range.—World-wide, in tropical and semitropical areas. 

Remarks.—Eight genera of this family occur in Mexican territory. 
Three of these have probably been introduced accidentally from other 
parts of the world. Peropus, with a single representative, arrived on 
the west coast of Mexico probably from the Philippines. Hemidac- 
tylus, with three species, has a somewhat more complicated history. 
One species reached the west coast, probably from the Philippines 
(frenatus), another came across the Atlantic to the east coast from 
Europe or Africa (turcicus), and a third (mabowia) probably came 
from somewhere in the West Indies or, not impossibly, from Mada- 
gascar or South Africa where it also occurs. Aristelliger undoubtedly 
was Imported from the West Indies. A fourth genus, Gonatodes, has 


CHECKLIST OF REPTILES OF MEXICO 41 


only recently been reported in Mexico, and may possibly have been 
transported by man from Central America. 

It would appear from collections that all these genera have very 
limited or discontinuous distribution in Mexico. 

The genera Phyllodactylus, Sphaerodactylus, and Thecadactylus very 
probably reached Mexico without the aid of man. 


KEY TO MEXICAN GENERA OF GEKKONIDAE 


1. Eyelid fully developed, body covered with small cycloid somewhat imbricating 
scales, with or without somewhat larger tubercles on body or tail; a few 
preanal pores present; digits cylindrical, never distinctly widened. 

Coleonyx (p. 41) 
Eyelid rudimentary, not capable of covering eye____..____________-__-- 2 

2. Under surface of digits unwidened, covered with flat smooth scales, lacking 

leaflike lamellae; no preanal or femoral pores; diminutive size. 

Gonatodes (p. 45) 
A part of under surface of digits with widened lamellae present; femoral pores 
LESCOL AOSEDUy SIZO VALTIADLO tae sore eet Pak ee ee eee ee ee 3S 
38. A single widened terminal lamella on digits with the claw lateral to it; no 
femoral pores; diminutive in size___.____.______-_-_ Sphaerodactylus (p. 52) 
Digits with one pair or several pairs of widened lamellae_______________-_ 4 
4, A single terminal pair of lamellae, the claw between them; size larger; preanal 
and femoral pores absent in Mexican forms_____-_-- Phyllodactylus (p. 46) 
Part of under surface of digits widened with a single or double series of lamellae, 
usually not present under terminal joint; pores present or absent______-_ 5 

5. Subdigital lamellae undivided, in a single row____---____-_ Aristelliger (p. 51) 

Subdigital lamellae in two rows, divided medially__.____....__..____--___ 6 

6. Tail with enlarged, sometimes spiny tubercles, forming longitudinal rows or 
transverse whorls; body with at least some enlarged tubercles, which may be 
fiat or trihedral; femoral or preanal pores present____Hemidactylus (p. 49) 

Tail and body lacking spines or enlarged tubercles; pores present or absent__ 7 

7. Tail with flat ventral surface, serrated lateral edge, and a median row of en- 
larged scales on ventral surface; a long, doubly curved series of femoral 
pores; a fold of skin on posterior side of thigh and leg_____- Peropus (p. 51) 

Tail rounded, tapering, covered with imbricate scales, a little larger on under 
surface; no femoral or preanal pores; no skin fold on hind leg. 
Thecadactylus (p. 49) 


Genus COLEONYX Gray 


Coleonyx Gray, Ann. Mag. Nat. Hist., vol. 16, 1845, p. 162—KtauBEr, Trans. 
San Diego Soc. Nat. Hist., vol. 10, No. 11, 1945, pp. 133-216. 

Brachydactylus Peters, Monatsb. Akad. Wiss. Berlin, 1863, p. 41 (type, B. 
mitratus Peters). 


Genoiype.—Coleonyz elegans Gray. 

Range.—The genus extends from Southwestern United States to 
Panama. 

Species.—Five, with a total of 12 forms. Four species and 10 
forms are known or are to be expected in Mexico. 


42 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 
KEY TO MEXICAN FORMS OF COLEONYX # 


1. Dorsal surface covered with small granular scales, without larger tubercular 


scales. Sia Petits. bebe 2 Ta eee De ee eee eee eet ARO 2 
Dorsal surface with larger tubercles intermingled with the small granular 
peales=.% =~ = AN F205. chy ge eR Lene Sa ae 5 ee Eases dshes 9 


2. Two light and 3 dark bands between level of arm and leg insertions. 
fasciatus (p. 43) 
Usually 4 or parts of 4 dark bands between levels of arm and leg insertions, or 
theibands: brokenwup Into;spotssossaoe—aoee ease eee ee ee ee 3 
3. Preanal pores small, usually 4 in number (rarely more or less), the pore-bearing 
scales separated medially; maximum snout-vent measurement about 59 
TON se oreo hte i i ca A of brevis (p. 43) 
Preanal pores larger, usually 6 to 10, the series continuous, not separated 
medially; maximum snout-vent measurement 77 mm____-_-___--_------ 4 
4. Usually 4 or less postmental scales touching mental_variegatus slevini (p. 45) 
Usually 5 or more postmental scales touching mental_______________---_-- 5 
5. Preanal pores in males usually 8 or more___--_-_- variegatus bogerti (p. 44) 
Preanalipores nim alespusuallliyg 7c T Css ees ee ee oe 6 
6. Dark transverse body bars in the adults considerably wider than the light 
ANFETSPACESE anes sateen ee is ree ee a reeaeale U 
Dark transverse body bars in adults about equal to or narrower than the 
light interspaces; or bars obsolete and replaced by spotting--.__.________- 8 
7. A middorsal light longitudinal line usually splitting the dorsal body bars in 
Ge ACatS a — e - tee  g  w variegatus sonoriensis (p. 45) 
No middorsal light longitudinal line; adults with longitudinal edges of the 
dark body bars even, with narrow uniform interspaces. 
variegatus peninsularis (p. 44) 
8. Dark body bandsin the adults unicolor; top of head unicolor; nuchal light loop 
MATEOW/ ANC ClORT 320 4 ee, che eee variegatus abbotti (p. 44) 
Dark body bands in the adults with light centers producing a double-barred 
effect, or bars obsolete and replaced by spotting; top of head spotted; nuchal 
light loop irregular or obsolete________-___- variegatus variegatus (p. 44) 
9. Enlarged supranasals 3 separated by a median circular scale. 
elegans nemoralis (p. 43) 
Enlarged supranasals in contact____-._-_._--__-____- elegans elegans (p. 42) 


COLEONYX ELEGANS ELEGANS Gray 


Coleonyx elegans Gray, Ann. Mag. Nat. Hist., vol. 16, 1845, p. 162. 

Coleonyx elegans elegans, KuauBEr, Trans. San Diego Soc. Nat. Hist., vol. 10, 
1945, pp. 191-195, map. 

Glymnodactylus] scapularis A. Dumfrit, in Duméril and Duméril, Catalogue 
méthodique de la collection des reptiles, 1851, pp. 45-46 (Petén, Guatemala, 
here restricted to La Libertad; Mus. Hist. Nat. Paris). 

Gymnodactylus coleonyx A. Dumérit, Arch. Mus. Hist. Nat. Paris, vol. 8, 1858, 
p. 483 (same type as scapularis). 


Type.—Brit. Mus. Nat. Hist., male spec.; Mr. Dyson collector. 


33 Adapted from Klauber, op. cit., pp. 205-206. 

23 We believe the scale above nostril should be designated supranasal rather than prenasal; and those scales 
following the mental should be postmental rather than gular, since the latter term refers to throat or gullet, 
rather than chin. 


CHECKLIST OF REPTILES OF MEXICO 43 


Type locality.—Belize, British Honduras. 

Range.—Atlantic slopes from central Veracruz through the Yucatan 
Peninsula, including Honduras and northern Guatemala; on Pacific 
slopes in Chiapas and undoubtedly adjacent Guatemala. Reported 
from Veracruz: Jalapa, Potrero Viejo, San Lorenzo, Orizaba, Tezonapa, 
Zempoala, Presidio, San Juan Cuatotolapam; Oaxaca: San Cristébal, 
Cosolapa; Tabasco: Teapa, Tenosique; Campeche: Tuxpefia Camp, 
Apasote, Encarnacién; Yucatdén: Chichen Itzi, Mayapan, Puz, Gon- 
gora Caves, Ziz, Xkyc Cave; Chiapas: La Esperanza, Piedra Parada; 
Quintana Roo: Mujeres Island. 

COLEONYX ELEGANS NEMORALIS Klauber 


Coleonyx elegans, Gavow, Proe. Zool. Soe. London (vol. 2), 1905, p. 212. 
Coleonyzx elegans nemoralis KuAuBER, Trans. San Diego Soc. Nat. Hist., vol. 10, 
1945, pp. 195-199. 


Type-—EHT-HMS No. 10509; Hobart M. Smith collector. 

Type locality —Hacienda Paso del Rio, Colima. 

Range.—Colima to southeastern Oaxaca (probably also Michoacan). 
Probably confined to the coastal area of these states. Reported from 
Guerrero: Cocoyul, Agua del Obispo, 4 to 5 miles north of Acapulco; 
Michoacdn: vicinity of Jorullo (?); Oaxaca: (said to be intergrades 
with elegans) Tehuantepec, Mixtequilla, Tapanatepec, etc. 

COLEONYX FASCIATUS (Boulenger) # 

Eublepharis fasciatus BoULENGER, Catalogue of the lizards in the British Museum, 
vol. 1, 1885, p. 423.—Gitnruer, Biologia Centrali-Americana, Reptilia and 
Batrachia, 1893, p. 84, pl. 31, fig. A. 

Coleonyx fasciatus, TayLor, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 203-205. — 
KuavuBeER, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, pp. 182-184. 

Type.—Brit. Mus. Nat. Hist.; A. Forrer collector. 

Type locality — Ventanas, Durango. 

Range.—Known from western Durango, and Sinaloa. The only 
known specific locality other than the type locality is Sinaloa: 10 
miles south of Presidio. 

COLEONYX BREVIS Stejneger 

Coleonyx brevis StEINEGER, North Amer. Fauna, No. 7, 1893, pp. 163-164.— 
Sir, Trans. Kansas Acad. Sci., vol. 36, 1933, pp. 301-314, fig. 1B, pl. 1, 
figs. 2-3.—Kuauser, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, pp. 
184-191. 


Type.—U.S.N.M. No. 18627; Mr. Marnock collector. 
Type locality —Helotes, Bexar County, Tex. 
Range.—The Rio Grande Valley, from northern New Mexico to 
southern Texas, and southward through eastern Chihuahua and 
24 Bogert and Oliver (Bull. Amer. Mus. Nat. Hist., vol. 83, 1945, p. 330) suggest the possibility of inter- 
gradation of this form with variegatus. There is no evidence whatsoever that such isthe case. See Klauber, 


loc. cit. 
861316—50——-4 


44 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


western Tamaulipas to southern Coahuila and extreme eastern 
Durango. Reported from Tamaulipas: Mier; Nuevo Leén: 5 miles 
south of Sabinas Hidalgo, near China, Ciénega de Flores, Mamulique 
Pass; Coahuila: Monclova, Saltillo; Durango: 6 miles northeast of 
Pedricefia. 


COLEONYX VARIEGATUS VARIEGATUS (Baird) 


Stenodactylus variegatus Batrp, Proc. Acad. Nat. Sci. Philadelphia, vol. 10, 1858, 
p. 254. 

Coleonyx variegatus, Corr, Proc. Acad. Nat. Sci. Philadelphia, vol. 18, 1866, pp. 
125, 310.—Smitu, Trans. Kansas Acad. Sci., vol. 36, 1933, pp. 301-314. 
Eublepharis variegatus, BOULENGER (part), Catalogue of the lizards of the British 

Museum, vol. 1, 1885, pp. 233-234. 
Coleonyx variegatus variegatus, KuauBER, Trans. San Diego Soc. Nat. Hist., vol. 
10, 1945, pp. 138-153, map. 


Type.—U.S.N.M. No. 3217 (now lost). 

Type locality—Colorado Desert (fide Klauber, loc. cit.), here 
restricted to Winterhaven, Calif. 

Range.—Southern Nevada south through eastern California and 
western Arizona to northeastern Baja California and northwestern 
Arizona. Reported from Baja California: San Felipe, Colorado River 
Delta, Colorado Desert; Sonora: Paso McDougall, Sierra Blanca, 
Punta Pefiasco, etc. 


COLEONYX VARIEGATUS ABBOTTI Klauber 


Coleonyx variegatus abbotti KuAuBER, Trans. San Diego Soc. Nat. Hist., vol. 10, 
1945, pp. 154-159. 

Type.—No. 34790, collection of L. M. Klauber; William Moore 
collector. 

Type locality—Proctor Valley, between Jamul and Upper Otay 
Reservoir, San Diego County, Calif. 

Range.—Southwestern California and northwestern Baja California. 
Reported from Baja California: Ensenada, 65 miles southeast of 
Tecate, San José (lat. 31°), ? Calmalli, Isla de Cedros. 


COLEONYX VARIEGATUS BOGERTI Klauber 


Coleonyx variegatus bogerti KuauBEeR, Trans. San Diego Soc. Nat. Hist., vol. 10, 
1945, pp. 176-182. 


Type.—lL. M. Klauber No. 32486; Lee W. Arnold collector. 

Type locality Xavier, Pima County, Ariz. 

Range.—Southeastern Arizona and presumably adjacent Sonora 
and New Mexico. Not yet recorded from Mexico. 


COLEONYX VARIEGATUS PENINSULARIS Klauber 


Coleonyx variegatus peninsularis KLAUBER, Trans. San Diego Soc. Nat. Hist., 
vol. 10, 1945, pp. 160-162. 


Type.—Mus. Comp. Zool. No. 37210; Miguel L. Cornejo collector. 


CHECKLIST OF REPTILES OF MEXICO 45 


Type locality —La Paz, Baja California, Mexico. 
Range.—Eastern side of southern Baja California. Reported from 
Baja California: San José del Cabo, between Loreto and Comondt. 


COLEONYX VARIEGATUS SLEVINI Klauber 


Coleonyx variegatus slevini KLAUBER, Trans. San Diego Soc. Nat. Hist., vol. 10, 
1945, pp. 167-171. 
Type.—California Acad. Sci. No. 51697; J. R. Slevin collector. 
Type locality —‘‘South Santa Inez Island” (lat. 27° W.), Gulf of 
California (Baja California coast). 
Range.—Isla Santa Inez and Isla de San Marcos, Gulf of California. 


COLEONYX VARIEGATUS SONORIENSIS Klauber 


Coleonyx variegatus sonoriensis KLAUBER, Trans. San Diego Soe. Nat. Hist., vol. 
10, 1945, pp. 162-166. 


Type.—Mus. Zool. Univ. Michigan No. 72140; Morrow J. Allen 
collector. 

Type locality —F ive miles southeast of Hermosillo, Sonora. 

Range.—Known only from central western Sonora. Reported 
from La Posa 10 miles northwest of Guaymas, Tepoca Bay, Sierra 
Alamo (30 miles west of Caborca), Isla de Tiburén. 


Genus GONATODES Fitzinger 
Gonatodes F1tzINGER, Systema reptilium, 1848, pp. 91-92. 


Genotype.-—Gymnodactylus albogularis Duméril and Bibron =Gona- 
todes albogularis (Duméril and Bibron). 

Range.—Tropical America; West Indies. Species in South Asia, 
Africa, and Malaya formerly regarded as members of the genus are 
now placed in Cnemaspis. 

Species.—About 10, of which only one occurs in Mexico. 


GONATODES FUSCUS (Hallowell) 


Stenodactylus fuscus HALLOWELL, Journ. Acad. Nat. Sci. Philadelphia, ser. 2, 
vol. 3, 1855, p. 33. 

Gymnodactylus fuscus, A. DumBRiL, Arch. Mus., vol. 8, 1856, p. 477.—Bocourt, 
Mission scientifique au Mexique . . ., Etudes sur les reptiles, livr. 2, 1873, 
pp. 48, 49, pl. x, fig. 5, 5a, 5b, 5c. 

[Gonatodes albogularis] fuscus, BOULENGER, Catalogue of the lizards in the British 
Museum, ed. 2, vol. 1, 1885, pp. 59-60. 

Gonatodes fuscus, STEJNEGER, Proc. U. 8. Nat. Mus., vol. 58, 1917, pp. 264-265.— 
Smiru, Copeia, 1944, p. 187. 

Goniodactylus braconnieri O’SuHauGcHNessy, Ann. Mag. Nat. Hist., ser. 4, vol. 16, 
1875, p. 265 (fide Boulenger; type locality, Baranquilla, Colombia; Brit. 
Mus. Nat. Hist.; Mr. Rippon collector). 

Type.—Acad. Nat. Sci. Philadelphia? 


Type locality —Nicaragua, here restricted to Rama. 


46 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Range.—Huixtla, Chiapas (only definite Mexican record), south- 
ward in Central America to northwestern South America; Jamaica 
(probably introduced); Florida (probably introduced). 


Genus PHYLLODACTYLUS Gray 


Phyllodactylus Gray, Spicilegia zoologica, 1830, p. 3. 
Discodactylus Firzincer, Systema reptilium, 1843, p. 95 (type, Phyllodactylus 
pulcher Gray). 

Genotype.—Phyllodactylus pulcher Gray. 

Range.—California to and including most of South America; 
Galapagos Islands, southern Asia, Africa, Madagascar, Australia, 
et cetera. 

Species.—About 48 species are known; 9 are known from Mexico.” 


KEY TO MEXICAN SPECIES OF PHYLLODACTYLUS 


1. Dorsal squamation consisting of minute rounded scales without admixture of 
larger tubercles; no enlarged tubercles on tail; small, length 52 mm. 

unctus (p. 47) 

Dorsal squamation with an admixture of larger tubercles; larger tubercles 


present/or absention alles 28 2S vase ee ee ee ee eee 2 
2. Tail lacking any trace of enlarged tubercles on dorsal surface__________-_-_ 3 
Tail with smaller or larger tubercles intermixed with dorsal lepidosis______- 4 


3. Scales on head between middle of orbits 15-17 %*; on dorsal and lateral surface 
14 to 16 rows of large, very distinct trihedral tubercles; smaller, 44 -+- mm. 
bordai (p. 47) 

Scales between orbits 19-24; dorsal tubercles low, small, rather indistinct, 
arranged in 8 to 12 rows; larger, 90 mm___________-__-- delcampi (p. 47) 

4. Two rows of tubercles on dorsal surface of tail (greatly reduced or absent in 
homolepidurus) 22 oe ee 5 a i ee ee Ee eee 5 
More than two rows of enlarged tubercles, at least on basal third of tail._._ 6 

5. Scales between middle of orbits 22-26; tubercles on back and tail large, well 
CLO Fie Cle a eee a ear ee ees eR cece aa TSN al a NORA a muralis (p. 47) 


Scales between middle of orbits 19-24; tubercles on back much smaller than in 
muralis, the caudal tubercles reduced so that they can be discerned only 
with idificultys.6 7mm 0 os a as ee homolepidurus”’ (p. 48) 


% Phyllodactylus pulcher Gray, Spicilegia Zoologica, from ‘Tropical America,” may be found to occur in 
Mexico. It is distinct from all others listed here by having 20 to 22 longitudinal series of dorsal tubercles 
(rather than 14 to 16), and larger ventral scales, in 22 longitudinal and 53 transverse series (rather than 25 to 
30 and 65, respectively). 

We likewise are unable to place Phyllodactylus mentalis Werner (Jahrb. Hamb. Wiss. Anstalten, vol. 27, 
No. 2, 1910, p. 5; Zool. Mus. Hamburg; L. von Péppinghausen coll.; Central America or Mexico). Mosauer 
(Copeia, 1936, p. 144) includes a few notes on the type. 

36 Not counting scales on vestigial eyelids. 

27 Bogert and Oliver (Bull. Amer. Mus. Nat. Hist., vol. 83, 1945) note that at least nine species of Phyllo- 
dactylus have been described from western Mexico in recent years and that certain of these are allopatric and 
their characters strongly suggest that a more accurate ‘‘indication of the biological conditions that exist’? may be 
obtained by considering them as subspecies or races of the same species. They proceed to place homolepi- 
durus as a subspecies under tuberculosus. Since the latter name cannot be fixed at the present time, we do 
not follow this proposal. Unfortunately, these authors have not examined most of the described forms and 
have overlooked the fact that at least four of the species may occur in the same general locality. They state 
that “the lizards of this genus exhibit considerable variation.’”? These forms are probably no more variable 
than other lizards. For many years most North American species of Phyllodactylus were classified in muse- 
ums as “‘tuberculosus,’’ and examination of such material might create the erroneous idea that ‘‘tuberculosus” 
was an extremely variable species. 


CHECKLIST OF REPTILES OF MEXICO 47 


6. Four distinct longitudinal rows of enlarged tubercles at least on basal fourth of 


tail, usually in rather regular transverse rowsS__~--------------------- a 
At least six distinct rows of enlarged tubercles on basal part of tail, not in- 
variably arranged in regular transverse rows__._.-------------------- 8 


7. Scales between orbits 12-16; dorsal and posterior part of thigh with numerous 


large conical tubercles; color darker above, cream color below; 80 mm. 
lanei (p. 48) 


Similar to above but tubercles on back heavier, higher; head deeper, less 


depressed, color lighter; yellow below; 66 mm. (1 specimen). 
magnatuberculatus (p. 48) 


8: Scales between orbits 25-30; 95 mm-_-__--____-_....._----.- magnus (p. 48) 
Scales between orbits much fewer_._-.._____-_-_------ tuberculosus (p. 49) 


PHYLLODACTYLUS UNCTUS (Cope) 


Diplodactylus unctus Cor®, Proc. Acad. Nat. Sci. Philadelphia, 1863, p. 102. 
Phyllodactylus (Diplodactylus) unctus, Bocourt, Mission scientifique au Mexique 
. . .» Etudes sur les reptiles, livr. 2, 1873, pp. 43-44. 
Phyllodactylus unctus, TAyLor, Univ. Kansas Sci. Bull., vol. 28, 1942, pp. 96-99, 
fig. 2, A—D. 
Type. —U.S.N.M. No. 5304. 
Type locality —Cape San Lucas, Baja California. 
Range.—Southern Baja California. Reported from Baja California: 
San José del Cabo; Isla Partida near Isla Espiritu Santo, Isla Gallina, 
and Isla San Francisco. 


PHYLLODACTYLUS DELCAMPI Mosauer 


Phyllodactylus delcampi Mosavtr, Copeia, 1936, pp. 141-146. 


Type.—Mus. Comp. Zool. No. 41238; W. Mosauer collector. 
Type locality.—Tierra Colorada, Guerrero. 
Range.—Known only from the type locality. 


PHYLLODACTYLUS BORDAIT Taylor 


Phyllodactylus bordai Taytor, Univ. Kansas Sci. Bull., vol. 28, pt. 1, 1942, pp. 93- 
96, fig. 1. 
Type-—EHT-HMS No. 27732; E. H. Taylor collector. 
Type locality.—Six miles north of Taxco, Guerrero. 
Range.—Northern Guerrero; known only from the type locality. 


PHYLLODACTYLUS MURALIS Taylor 


Phyllodactylus muralis Taytor, Univ. Kansas Sci. Bull., vol. 26, 1939 (1940), 
pp. 543-547, fig. 7. 
Type-—EHT-HMS No. 10902; H. M. Smith collector. 
Type locality —Totolapam, Oaxaca. 
Range.—Oaxaca. Reported from Totolapam, Tehuantepec, San 
Gerénimo, Salina Cruz. 


48 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 
PHYLLODACTYLUS HOMOLEPIDURUS Smith 


Phyllodactylus homolepidurus Smitu, Univ, Kansas Sci. Bull., vol. 22, 1935, pp. 
121-125, pl. 25, fig. 2, text fig. 1A. 

Phyllodactylus tuberculosus homolepidurus, BoGERT and Oxtver, Bull. Amer. Mus. 
Nat. Hist., vol. 83, art. 6, 1945, pp. 344-345. 


Type-—EHT-HMS No. 10853 (originally EHT field No. 146); E. 
H. Taylor collector. 

Type locality —Five miles southwest of Hermosillo, Sonora. 

Range.—Southern Sonora and adjacent Chihuahua. Reported from 
Sonora: La Posa, 5 miles southwest of Hermosillo, Guirocoba; Chi- 
huahua: Batopilas. 


PHYLLODACTYLUS MAGNATUBERCULATUS Taylor 


Phyllodactylus magnatuberculatus TAyLor, Univ. Kansas Sci. Bull., vol. 26, 1939 
(1940), pp. 547-549. 
Type-—EHT-HMS No. 10995; H. M. Smith collector. 
Type locality —Acapulco, Guerrero. 
Range.—Known only from the type locality. 


PHYLLODACTYLUS MAGNUS Taylor 


Phyllodactylus magnus Taytor, Univ. Kansas Sci. Bull., vol. 28, 1942, pp. 99- 
108, fig. 3. 

Type-—EHT-HMS No. 21783; E. H. Taylor collector. 

Type locality —Tierra Colorada, Guerrero. 

Range.—Guerrero, Oaxaca, and Chiapas. Reported from Guerrero: 
Garrapatas, El] Ocotito, Tierra Colorada, Agua del Obispo; Oazaca: 
Tehuantepec, Oaxaca, Tres Cruces, Cerro Arenal, Cajén de Piedra, 
Escurana; Chiapas: Tonala. 


PHYLLODACTYLUS LANEI # Smith 


Phyllodactylus lanei Smitu, Univ. Kansas Sci. Bull., vol. 22, 1985, pp. 125-132, 
fig. 1b; pl. 25, fig. 3—Taytor, Univ. Kansas Sci. Bull., vol. 26, No. 15, 
1939 (1940), pp. 541-542, fig. 6. 


Type-—EKHT-HMS No. 10942 (originally EHT field No. 146); E. 
H. Taylor and H. M. Smith collectors. 
Type locality —Near Tierra Colorada, Guerrero, Mexico. 
Range.—Sinaloa and (?) western Durango *” southward to central- 
southern Guerrero, probably confined to the coastal areas up to an 
elevation of about 3,500 feet. Recorded from the states of Sinaloa 
(including the Tres Marias Islands), ? Durango, Nayarit, ? Jalisco, 
Colima, Michoacan, and Guerrero. 
28 The suggestion of common identity of P. lanei and P. tuberculosus by Mosauer is, we believe, unten- 
able (Mosauer, Copeia, 1936, p. 194). 
2 A specimen of Phyllodactylus collected at Durango, Mexico by Dr. T. H. Webb (U.S. N. M. No. 3208) 
and figured by Baird (Reptiles of the Boundary, U. S. and Mexican Boundary Surveys, 1859, p. 12, pl. 


23, figs. 1-8) may or may not belong to this species, since Durango is considerably higher than this species 
is known to range. 


CHECKLIST OF REPTILES OF MEXICO 49 


PHYLLODACTYLUS TUBERCULOSUS Wiegmann * 


Phyllodactylus tuberculosus W1BGMANN, Nova Acta Phys.-Med. Acad. Caes. 
Leop.-Carol., vol. 17, pt. 1, 1835, p. 241, pl. 18, fig. 2. 

? Phyllodactylus Xanti Corn, Proc. Acad. Nat. Sci. Philadelphia, 1863, p. 103 
(Cape San Lucas, Baja California). 


Type.—? Zool. Mus. Berlin; collector unknown. 

Type locality —“Californien.”’ 

Range.—? Extreme southern part of California, south through Baja 
California. 


Genus THECADACTYLUS Gray 
Thecadactylus Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 198. 


Genotype.—Gecko rapicauda Houttuyn. 

Range.—Southern Mexico to northern South America (Peru); 
Antilles; islands north of Australia in Torres Strait. 

Species.—Two. 


THECADACTYLUS RAPICAUDUS (Houttuyn) 


Gekko rapicauda Houtruryn, Verh. Genootsch. wet. Vlissing., vol. 9, 1782, p 
323, pl. 3, fig. 1. 

Thecadactylus rapicauda, Gray, Catalogue of the lizards in the collection of the 
British Museum, 1845, p. 146.—BouLrenaeEr, Catalogue of the lizards in the 
British Museum, vol. 1, 1885, pp. 111-112. 


Type.—Unknown. 

Type locality —American Islands, here restricted to Chichen Itzé4, 
Yucatan. 

Range.—Southward from Yucatén through Central America to 
northwestern South America; Lesser Antilles. Reported from 
Yucatdén: Chichen Itz4; Puz Cave, Oxkutzcab; Chakxix Cave, Tekax. 
The Guadalajara reference of Cope (U. S. Nat. Mus. Bull. 32, 1887, 
p. 28) is very doubtful. 


Genus HEMIDACTYLUS COken 


Hemidactylus OxeEn, Isis, 1817, p. 1183 (based on Cuvier’s Hemidactyle, Régne 
animal, vol. 2, 1817, p. 47). 

Tachybates Firzincer, Systema reptilium, 18438, p. 105 (type, ‘‘Hemidactylus 
mabuya Cuvier’’). 

Pnoépus Firzincer, Systema reptilium, 18438, p. 105 (type, ‘“Hemidactylus javanicus 
Cuvier’’). 


30 We have here followed the idea that Baja California or California is the type locality. However, since 
there are two forms (one undescribed) in Baja California, one with a spiny tail, the other without spines 
on the tail, it is not certain to which the name must apply, inasmuch as the type has a reproduced tail. 
Direct comparison of the two species with the type of tuberculosus will be necessary to determine the proper 
application of the name on the basis of characters not now discernible from the description or figure. 

There are many reports of this form in the literature, and it is impossible to determine to what species 
many of them refer. Although for that reason many localities are thus not here cited, we have included 
every state from which ‘‘twberculosus’’ has been recorded in the records for at least one species, basing our 
allocation either upon actual examination of specimens or a reasonably accurate geographic indication, 
There is one exception: Iziicar de Matamoros, Puebla. 


50 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Genotype.—Gekko tuberculosus Daudin [=Hemidactylus mabouia 
Moreau de Jonnés]. 

Range.—Widespread in Africa, southern Eurasia, tropical America, 
Polynesia. 

Species—About 60 known. Three are found in Mexico. 


KEY TO MEXICAN SPECIES OF HEMIDACTYLUS 


1. Preanal pores 4 to 10, forming a short angular series; no femoral pores; dorsal 
scales minute granules intermixed with large tubercles usually larger than 
interspaces between them; inner digits well developed___-_- turcicus (p. 50) 

Preanal and femoral pores forming a continuous series____._..___-_____- 2 

2. Small granules of the dorsal surface with only a few relatively small, convex 
tubercles, these usually on posterior part of body; pores 15-18 on each side, 
not separated .mesially =... IVICA EE See ota as frenatus (p. 50) 

Small granules of the dorsal surface mixed with very numerous trihedral 
tubercles, irregularly scattered; pores from 15 to 30 on each side. 
mabouia (p. 50) 


HEMIDACTYLUS MABOUIA (Moreau de Jonnés) 


Gecko mabouia MoreEav de Jonnb&s, Bull. Soc. Philom., 1818, p. 138. 
Hemidactylus mabouia, BouLENGER, Catalogue of the lizards of the British 
Museum, vol. 1, 1885, pp. 122-123.—Lovrripasx, Copeia, 1941, p. 248. 


Type.—Mus. Hist. Nat. Paris. 

Type locality —“Antilles,” here restricted to St. Vincent, Lesser 
Antilles. 

Range.—In Mexico known only from ‘Veracruz’ (probably city). 
Widespread in the West Indies, and the northern and eastern coast of 
South America; Madagascar and Africa. The American localities are 


probably due to relatively recent introductions from Africa, or the 
West Indies. 


HEMIDACTYLUS FRENATUS Schlegel 


Hemidactylus frenatus SCHLEGEL, in Duméril and Bibron, Erpétologie générale 

. .«, Vol. 3, 1836, pp. 366-368.—TayLor, Univ. Kansas Sci. Bull., vol. 26, 

1939, pp. 444.—Burt and Myers, Stanford Univ. Publ., biol. sci., vol. 8, 

1942, pp. 285-286.—Smiru, Anal. Inst. Biol., vol. 14, 1948, p. 342.— Eparen, 
Nat. Hist. Misc., No. 55, 1950, p. 1. 


Type.—Leyden Mus. 

Type locality.— Java. 

Range.—Guerrero, in the region about Acapulco. Widespread in 
southern Asia, Pacific Islands, etc. Reported from Guerrero: Aca- 
pulco, Tierra Colorada. 


HEMIDACTYLUS TURCICUS TURCICUS (Linnaeus) 


Lacerta turcica LINNAEUS, Systema naturae, ed. 10, 1758, p. 202. 

Hemidactylus turcicus, SmirH, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 132-138, 
pl. 23, fig. 2.—Granr and Drvst, Herpetologica, vol. 2, 1944, pp. 130-135. 

Hemidactylus turcicus turcicus, LovERIDGE, Copeia, 1941, p. 247. 


CHECKLIST OF REPTILES OF MEXICO 51 


Hemidactylus exsul BARBouUR and Cote, Bull. Mus. Comp. Zool., vol. 50, 1906, 
p. 148 (type locality, Progreso, Yucatén; Mus. Comp. Zool. No. 7039).— 
Stuart, Copeia, 1934, No. 4, p. 185. 

Type.—Unknown. 

Type locality—“Habitat in Oriente,” here restricted to Cairo, 
Egypt. 

Range.—Tamaulipas south to Yucatan, distribution very probably 
discontinuous. Found in the West Indies, Florida, North Africa, 
and southwestern Asia. Reported from Tamaulipas: 12 miles east 
of Llera; Veracruz: no specific locality; Tabasco: Alvaro Obregén, 
Emiliano Zapata; Campeche: Ciudad del Carmen; Yucatdn: Progreso. 
The Mexican records are due to modern introductions, probably from 
northern Africa. 

Genus ARISTELLIGER Cope 


Aristelliger Corr, Proc. Acad. Nat. Sci. Philadelphia, 1861, p. 496. 
Idiodactylus Bocourt, Mission scientifique au Mexique ..., Etudes sur les 
reptiles, livr. 2, 1873, p. 41 (type, Idiodactylus georgeensis Bocourt). 


Genotype.—Aristelliger lar Cope. 

Species.—Six, all West Indian except the one Yucat4n form. 

Range.—West Indies and the peninsula of Yucatan. 
ARISTELLIGER GEORGEENSIS (Bocourt) 


Idiodactylus georgeensis Bocourt, Mission scientifique au Mexique . . ., Etudes 
sur les reptiles, livr. 2, 1873, pp. 41—42, pl. 10, fig. 1. 

Aristelliger georgeensis, Scumipt, Publ. Field Mus. Nat. Hist., zool. ser., vol. 22, 
1941, p. 490. 

Aristelliger praesignis, BoULENGER, Catalogue of the lizards in the British 
Museum, vol. 1, 1885, pp. 146-147 (part). 

Aristelliger irregularis Corr, Proc. Amer. Philos. Soc., vol. 22, 1885, p. 387 
(Cozumel Island; Ridgway coll.; U.S.N.M. No. 13908). 


Type.—Mus. Hist. Nat. Paris. 

Type locality —St. George Island, near Belize, British Honduras. 

Range.—The eastern edge of the Yucatan Peninsula. Recorded in 
Mexico only from Cozumel Island (Quintana Roo). 


Genus PEROPUS Wiegmann?! 


Peropus WirGMann, Nova Acta Acad. Leop.-Carol., vol. 17, pt. 1, 1835, p. 238. 

Dactyloperus FirzincEr, Systema reptilium, 18438, p. 103 (type, Peropus peronti 
Duméril and Bibron= Peropus mutilatus Wiegmann). 

Peripia Gray, Catalogue of the specimens of lizards in the British Museum, 
1845, p. 158 (type, Peripia peronit Duméril and Bibron=Peropus muttlatus 
Wiegmann). 

Chalinocnemis Ducts, La Naturaleza, vol. 6, 1883, p. 312 (type, Hemidactylus 
navarrt Dugés=Peropus mutilatus Wiegmann). 

Spasmocnemis Duais, loc. cit. (type as for preceding). 


Genotype.—Hemidactylus mutilatus Wiegmann. 


31 We are inclined to maintain Peropus and Gehyra based on the different character of the subdigital 
lamellae. 


52 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Range.—Widespread in southern Asia from Ceylon to the Philip- 
pines; south through the Malay Archipelago to New Guinea; Sey- 
chelles; western Mexico. 

Species.—One form occurs in Mexico, probably introduced from the 
Philippines by man; about 10 others are known. 


PEROPUS MUTILATUS (Wiegmann) 


Hemidactylus (Peropus) mutilatus WimGMANN, Nova Acta Acad. Leop.-Carol., 
vol. 17, 1835, p. 238. 

Peropus mutilatus, Smirn and Necker, Anal. Esc. Nac. Cienc. Biol., vol. 3, 1948, 
pp. 197-199, pl. 3, fig. 3. 

Hemidactylus Navarri Ducts, La Naturaleza, vol. 6, 1883, pp. 309-312, pl. 7a 
(type locality, San Blas, Nayarit; 2 types in Alfredo Dugés Museum, Guana- 
juato, Guanajuato.) 

Type.—Zool. Mus. Berlin?. 

Type locality —Manila, Philippine Islands. 

Range.—Seaports of Nayarit and Sinaloa. Widespread in south- 
eastern Asia, Philippine Islands, Dutch Indies, to New Guinea and 
certain Pacific Islands. Reported from Nayarit: San Blas, Tepic; 
Sinaloa: Presidio, Mazatlan. 


Genus SPHAERODACTYLUS Wagler 


Sphaerodactylus (part) WaGurrR, Natiirliches System der Amphibien, 1830, p. 
143.—Barzsour, Mem. Mus. Comp. Zool., vol. 47, 1921, pp. 215-279, pls. 
1-26. 

Genotype.—Sphaerodactylus sputator Cuvier. 
Range.—West Indies, southern Mexico, Central America. 
Species—About 38 forms. Three are known or expected from 
Mexico.” 
KEY TO MEXICAN SPECIES OF SPHAERODACTYLUS 


ieeDorsaléscalesvoranularmkeeledss sss aes ee eee ae eee lineolatus (p. 52) 
Dorsal'scales imbricatessmooths £5228 Uae STE tees ee) PA 2 

2. A single nuchal band; two distinct spots at base of tail, light spots at elbow 
and knee usually idiscernibles= fs! 22 eee sees glaucus glaucus (p. 53) 


A cream-colored nuchal band bordered by black bands of equal width anteriorly 
and posteriorly; two narrow longitudinal light lines from collar to eye; in- 
distinct narrow light bands on back; spots on tail base absent or obsolescent; 
no light spots at knee and elbow in young or adult. 

glaucus torquatus (p. 53) 


SPHAERODACTYLUS LINEOLATUS Lichtenstein 
Sphaerodactylus lineolatus LicHTENSTEIN, Nomenclator reptilium et amphibiorum 


musei zoologici berolinensis, 1856, p. 46——BarBour, Mem. Mus. Comp. 
Zool., vol. 47, 1921, pp. 233-240, pl. 4, figs 1, 2, pl. 14, figs. 1-4. 


Type.—Zool. Mus. Berlin, three cotypes. 


32 Until new material is discovered in Mexico, the Mexican reports of Sphaerodactylus fantasticus, an- 
thracinus, punctatissimus (cinereus), and sputator, are to be regarded as uncertain. The type locality of 
anthracinus is ‘‘Mexico,”’ but possibly in error (see p. 213). It would not beimpossible for a part of these forms 
to occur, as recent introductions in Mexico. Banana fruits serve as a ready means of transportation. 


CHECKLIST OF REPTILES OF MEXICO 53 


Type locality —Veraguas, Panamé. 
Range.—The eastern edge of the Yucatan Peninsula southeastward 
to Colombia. No definite records from Mexico.* 


SPHAERODACTYLUS GLAUCUS GLAUCUS Cope 


Sphaerodactylus glaucus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1865, p. 192.— 
BouLENGER, Catalogue of the lizards in the British Museum, vol. 1, 1885, 
p. 221, pl. 18, fig. 3—BarBour, Mem. Mus. Comp. Zool., vol. 47, 1921, 
pp. 240-241, pl. 14, figs. 5-8.—Taytor, Univ. Kansas Sci. Bull., vol. 31, 
pt. 2, 1947, pp. 305-309, fig. 2. 

Sphaerodactylus glaucus glaucus, Smiru, Journ. Washington Acad. Sci., vol. 39, 
1949, pp. 34-35. 

Sphaerodactylus inornatus Peters, Monatsb. Akad. Wiss. Berlin, 1873, p. 738 
(Zool. Mus. Berlin No. 4589; Uhde coll.; Mexico #4 here restricted to Te- 
huantepec, Oaxaca. 

Type —U.S.N.M. No. 6572, three cotypes (now one, No. 13570, 
in Mus. Comp. Zool.); Arthur Schott collector. 

Type locality — Near Mérida, Yucatan. 

Range.—Known from southern Veracruz to Guatemala and British 
Honduras, and possibly to Costa Rica. Reported from Oaxaca: Te- 
huantepec, Salina Cruz, Guengola Mountain; Tabasco: Teapa; 
Campeche: Tuxpefia Camp, Ciudad del Carmen, Balchacaj, Apazote; 
Veracruz: Pérez, Coatzacoalcos River; Yucatén: Mérida. 


SPHAERODACTYLUS GLAUCUS TORQUATUS Sirauch 


Sphaerodactylus torquatus StraucH, Mem. Acad. Sci. St. Petersburg, ser. 7, vol. 
35, 1887, p. 35.—TayLor, Univ. Kansas Sci. Bull., vol. 31, 1947, pp. 302-305, 
figs. 1-2. 

Sphaerodactylus glaucus torquatus, SmitH, Journ. Washington Acad. Sci., vol. 
39, 1949, p. 35. 


Type—Mus. Petrograd No. 3268, three specimens; Hr. Saimin 
collector. 

Type locality —‘ Mazatlan, Mexico” (= Mazatlan, Oaxaca?). 

Range.—Uncertain. Known from the type locality and Cajén de 
Piedra, Oaxaca.® 


Family IGUANIDAE Gray 
Iguanidae Gray, Philos. Mag., ser. 2, vol. 2, 1827, p. 56. 


Genera.—About 65 genera are commonly recognized; 21 occur in 
Mexico. 


#3 The species is included here on the strength of its occurrence in British Honduras (Schmidt, Publ. 
Field Mus. Nat. Hist., zool. ser., vol. 22, 1941, p. 489), for although it has been cited from ‘“‘Mexico” by 
numerous authors, no definite record of its occurrence in that country is known to us. Its existence in at 
least Quintana Roo is virtually a certainty. 

#4 It now seems most probable that this name was applied to a unicolor intergrade between S. g. glaucus 
and S. g. torquatus such as those known to occur in the vicinity of Tehuantepec, Oaxaca. The entire descrip- 
tion, of both scutellation and coloration, fits glaucus, not lineolatus, although Barbour (op. cit., p. 238) places 
Peters’s name in the synonymy of the latter species. See Smith, loc. cit., and Taylor, loc. cit., for 
discussion.—H. M.8. 

35 The subspecies does not occur in the vicinity of the city of Tehuantepec, but only, so far as we are aware, 
near the coast southwest of Salina Cruz, where a village by the name of Mazatlan does exist. 


54 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Range.—Extreme southwestern Canada and extreme southeastern 
New York, southward throughout the Americas; Bermuda Islands; 
West Indies; Revillagigedos Islands; Galapagos Islands; Madagascar 
(two genera); Fiji and Tonga Islands (one genus). 


KEY TO MEXICAN GENERA OF IGUANIDAE 


Pe NO TEM OTAlGMOLeS sane ees Sere see eee nee eee en eV ec en OT 2, 
Femoral pores present, conspicuous in males, less well defined in females__ 5 

2. Ventral lamellae on next to last phalanx of digits expanded laterally, forming 
a padlike structure; posterior part of head not produced____Anolis (p. 55) 
Ventral lamellae on digits not expanded to produce a pad; posterior part of 


head more or less produced f2 ee ee eee ee ee ee 3 

3. Infradigital lamellae with a single, tubercular keel_____- Laemanctus (p. 69) 
Infradigital lamellae with several sharp keels___.._.__...-.-.-----_---- 4 

4.6 Toes*withyallateralifringe .. 0. Selt Je eee. Bes tees Basiliscus (p. 71) 
INoifiringeron. toes 22. = see eee ee ee Corythophanes (p. 68) 

5. A single, median, dorsal, longitudinal series of enlarged scales._._._._____-_- 6 


Seales in vertebral row not conspicuously if any larger than adjacent scales. 9 
6. Tail with whorls of much-enlarged, spiny scales, usually separated from each 


others byswihorlsyofesm7 all iscsi] esters ae ek en ene are tr eee 7 
Scales on dorsal and lateral surfaces of tail (except the vertebral row) subequal 
infsizessmiallel x 2 ewe eee Be kee UTR. os eee! Ne Aree oe eee 8 


7. A group of much enlarged, spinose, protuberant scales on shank and some- 
times on thigh; basal caudal region (or entire tail in short-tailed species) 
highly modified, sometimes flattened and widened, with whorls of greatly 
enlarged scales separated from each other by no more than 1 row of small 
scales; total length less than 450 mm___________--- Enyaliosaurus (p. 75) 

No conspicuously enlarged scales on shank or thigh; basal caudal area much 
like rest of tail (which is always long), not flattened but cylindrical, with 
whorls of enlarged scales separated from each other by two or more rows 
of small scales throughout most of tail; total length often more than 450 


mim, reachingrat least ol 200immess aoe ee eee eee Ctenosaura (p. 73) 

8. A large, circular shield below tympanum; a well-developed gular append- 
SECUSA RS I TUS a ats eae eee eee Iguana (p. 72) 

No shield below tympanum; no gular appendage__-__-__- Dipsosaurus (p. 77) 

9. Head bearing bony spines posteriorly, or elevated somewhat in a projecting 
BT ees ee enh te Sct a Phrynosoma (p. 94) 
Head normal, not produced posteriorly in a bony ridge or in spines______-- 10 


10. Superciliary scales not imbricate, but juxtaposed, like other scales on head; 
rostral subdivided, with a median suture meeting lip___Sauromalus (p. 79) 
Superciliary scales strongly imbricate, large; rostral not divided, no median 


suture reaching lip: 25s 2 ee ee Ses Ae eee el ee 11 

11. Sutures between supralabials diagonal, not vertical; mental very small, 
bordered posteriorly by a median postmental__..._..-.-_---------- 12 
Sutures between supralabials vertical; mental larger, not bordered posteriorly 

by median’ postmentals 2.2.6. 332.325 eee enn eee ee ee 14 

12 SON OLCAE OTC TING Sec eo Sec: 23a eae ent ey ea Holbrookia (p. 81) 
Asdistinet car opening. = 2. Jee ios ae ae eee ee eg ee yen ee Ree 13 


13. Interparietal much smaller than ear opening, not more than twice as large 
as any other dorsal head scale; several enlarged auricular lobules; a patch 
of enlarged scales with projecting points on rear of femur-_-_-_-_-_- Uma (p. 89) 


CHECKLIST OF REPTILES OF MEXICO 55 


Interparietal nearly as large as ear opening, or larger, much larger than 
any other dorsal head scale; no enlarged auricular lobules; no patch of 
enlarged scales on rear of femur_...........---.----- Callisaurus (p. 85) 

14. No complete transverse gular fold, or if so body and tail compressed and 
dorsal scales nearly uniform, large, no more than 75 from interparietal to 
rear of thighs! 2. 2s bie eee ee eee eee ee ea 15 

A complete transverse gular fold, marked by granular or reduced scales, 
much smaller than those preceding or following fold; dorsal scales very 
small, or at least not uniformly large and as few as 75 from interparietal 
to rear of thighs (except Uta squamata); body and tail flattened, never 
Compressed even SHeipiy ns S25 22 L 2 des 5 a NE a OP Bde 16 

15. A gular fold usually present; lateral scales granular; body and tail somewhat 
ROI pLCONOG seen kas ek eet a eS ee Sator (p. 139) 

Gular fold never present; lateral scales not granular in those species with 

COMMDRESSCG Cail 226 en wos a ee ee ee 2 2 Sceloporus (p. 105) 

16. Interparietal very large, about as large as or larger than ear opening.._._._ 18 
Interparietal very small, a great deal smaller than ear opening_.-.-__.-- 17 

17. Twoscales bordering mental posteriorly between infralabials; head very broad, 
TUS Ease TNE TENG COV eee er ats ae a Ee Crotaphytus (p. 91) 

Three or more scales bordering mental posteriorly between infralabials; head 

elongate, neck not so sharply defined__-.-...-..------- Gambelia (p. 93) 
18. Dorsal scales all small, granular, smooth, very little larger than lateral scales; 
enlarged supraoculars in more than one row; one or more blackish bars 
across shoulders; size frequently over 75 mm. snout to vent__._____-- 19 

Dorsal scales, at least toward middle of back, larger than lateral scales, keeled; 
enlarged supraoculars in one row; no blackish bars across shoulders; size 
OSS mt LL ANN ¢/ UAT STA O LLL Oe VC TN ce es ea 20 

19. Caudal scales smaller than ventrals on body, not mucronate; three or four 
DIAC KEDANGS ACLOSSyDOG yess see eee ee ee ee eee Petrosaurus (p. 90) 

Caudal scales large, mucronate, much larger than ventrals on body; a single 

narrow black band across shoulders__---.---------- Streptosaurus (p. 91) 
20. A longitudinal dorsolateral line or dermal fold, usually with some enlarged 
scales on it; usually a second, lateral fold; no small, rounded blue blotch 


behind axilla; frontal divided or not-_.-.------------ Urosaurus (p. 140) 
No longitudinal dorsolateral dermal fold; often a small, rounded, dark blue 
blotch behind axilla; frontal always divided___.___.___----- Uta (p.147) 


Genus ANOLIS Daudin 


Anolis Daupin, Histoire naturelle générale et particuliére des reptiles, year X 
(1802), pp. 50-51.—Barsour, Bull. Mus. Comp. Zool., vol. 78, 1934, pp. 
121-155. 

Dactyloa WacuER, Natiirliches System der Amphibien, 1830, p. 148 (type, Anolis 
viridis Wied). 

Tropidopilus Firzincer, Systema reptilium, 1848, p. 66 (type, Anolis fusco- 
auratus d’Orbigny, monotype). 

Trachypilus F1rzincER, op. cit., p. 67 (type, Anolis sagraet Duméril and Bibron). 

Pristicercus FirzINGER, op. cit., pp. 67-68 (type, Dactyloa biporcata Wiegmann). 

Ctenocercus F1tzINGER, op. cit., pp. 17, 68 (type, Dactyloa bullarits Wagler = Anolis 
carolinensis Linnaeus). 

Gastrotropis FirzinGER, loc. cit. (type, Dactyloa nebulosa Wiegmann). 

Coccoéssus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1862, p. 178 (type, Anolis 
pentaprion Cope). 


56 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Genotype —Anolis bullaris Daudin (= Anolis carolinensis Voigt). 
Species—Perhaps 350. Thirty-five forms are listed here, but two 


are not known from any definite locality, four are known only from 
“‘Mexico,”’ and the Mexican range of another is unknown. 


Range.—Southeastern United States to Brazil; West Indies. 


KEY TO MEXICAN SPECIES OF ANOLIS 


1. “Lailistrongly compressed 2257-4 se Sees See eee Soman See sto 2 
Tail subcylindrical, no more than slightly compressed_________-________ 4 
2. Two continuous rows of scales along middorsal line on tail__barkeri (p. 58) 
One row of seales ‘along middorsal’ line*of tail 222. 2 ee 3 
3: Ventralsukceleceiuant sea ies heres e es amie Rees wee coe sagrei mayensis (p. 59) 
Ventralsismoouhms ce a. ome Swe. ees ep Aa en een ete ee ae cozumelae (p. 59) 
4. Dorsal scales as large as or larger than ventrals, and both strongly keeled_ 5 


10. 


1, 


12. 


13. 


Dorsal scales smaller than ventrals, or one or both sets of scales smooth; or, 
if body scales as above, occipital much larger than ear opening__-___-_ 8 


. Supraorbital semicircles in contact or separated by a single series of scales. 


megapholidotus (p. 59) 
Supraorbital semicircles separated by 2 or 3 series of scales____________ 6 


. Dorsal and ventral scales subequal in size___.._________-_ metallicus (p. 59) 


Dorsal scales distinctly larger than ventrals____.._._.....-.___1.-.202 a 


. Tibia 75 percent to 85 percent length of head (to anterior border of ear) ; adult 


size smaller, about 36 mm.; dewlap brilliant red with a large purple spot 
BUCO ASE LIT Uitesy 2 ae eat ee ee ee ae ae humilis uniformis (p. 60) 
Tibia 90 percent length of head, or longer; adult size larger, about 50 mm.; 
dewlap red, without a purple spot_________..______ tropidonotus (p. 60) 
Pe Ventrals SMmOOUR ts 2 cn acs ales rece i ee) Rea en oe 9 
Ventralstikecled 25 t2ties 20.8 in 25 nh etrghye ah hee  O e eeee 25 
. Supraorbital semicircles in contact and a single row of only 4 or fewer large 
SUPraocWlars: 22 noe She Se eee eee aI el ee ee ee ee ae 10 
Supraorbital semicircles separated or, if in contact, supraoculars numerous (8 
Or7MmOre CHlareed oe ss ye Ae ae ne ee es ee ee a 14 
Two rows of loreals; female with a large gular “‘pouch’’; all scales on body 
flat and granular, small and equal on back and sides, larger on belly. 
impetigosus (p. 60) 
Four to six rows of loreals; females (not known in schmidti) lacking a large 
gular pouch; dorsal scales keeled, at least faintly larger than lateral scales. 
11 
Ventrals and dorsals subequal in size; enlarged supraoculars three-fourths 
widthvef supraccilararca_ oe -6 2= ses oe ee schmidti (p. 60) 
Ventrals distinctly larger than (at least 144 times as large as) dorsals; enlarged 
supraoculars narrower, half width of supraocular area or less________ 12 
Two gulars in contact with mental between chin shields; 9 or fewer scales 
across snout between second canthals (counting as the first canthal that 
which  bordersithe:superciliaries) = 3-= eeu ose. 2 ee ee 13 
Three or (generally) 4 gulars in contact with mental between chin shields; 
10 to 12 scales across snout between second canthals_-_____ gadovii (p. 61) 
Six scales between nasals; 5 scales in contact with rostral between supralabials; 
ventralsvlangen: strech) ae sates ee ee dunni (p. 61) 


36 In six species (cymbops, laeviventris, limifrons rodriguezii, milleri, schiedii, and schmidti) the ventrals may 
appear either smooth or keeled; these are keyed out under both categories. All others are uniformly and 
plainly either smooth or keeled. 


14. 


15. 


16. 


17s 


18. 


19. 


20. 


21. 


22. 


23. 


24. 


25. 


CHECKLIST OF REPTILES OF MEXICO 57 


Seven or 8 scales between nasals; 6 scales in contact with rostral between 
supralabiais: ventrals smaller: 8... 2 aa ee Sk taylori (p. 61) 
All dorsals and laterals equal in size, minute, granular; tibia 85 percent length 
of head (to anterior border of ear); scales of supraorbital semicircles en- 
larged; head.scales:feebly keeled2io-22:.2228_hs5o22 22 damulus (p. 61) 
At least 2 or more vertebral rows of slightly enlarged scales, or, if all dorsal 
scales uniform, tibia less than 70 percent length of head, and either scales of 
supraorbital semicircles not enlarged, or head scales perfectly smooth__ 15 
Scales of supraorbital semicircles, at least in the frontal and frontoparietal 
area, not or scarcely larger than scales between; tail compressed, sub- 
triangular in section with a serrate crest; tibia much shorter than head 
(70 percent or less) all dorsal and lateral scales equal in size. 
pentaprion (p. 61) 
Seales of supraorbital semicircles distinctly larger than scales between; tail not 
or only slightly compressed, with little or no crest____-------------- 16 
Supraorbital semicircles either in contact or separated by a single series of 
scales; postanals enlarged in males (males not known in utowanae, cymbops, 


beckeri-.and. baccatus) =... ts ew 2) ules Sol een Deeks 17 
Supraorbital semicircles separated by 2 or 3 rows of scales; postanals not 

enlarged in males: (not. known in gintheriz)=— «2222-42552 24-222 SSL 22 
interparietal smaller than ear openings: i222 ivsi_-Ss22b52 J2es--5- 55428 18 
imbenparictalalarcersthan\cariopenin ge a= eee eee Sees eee 19 
wRhreeulorealerows=,2- <5. seis ss Saline oli eeu utowanae (p. 62) 
Seremslorenrows:.<..-2-cb Seseed ees eens ee ae ey cymbops (p. 62) 


All dorsal and lateral scales equal in size, minute, granular, smooth; tibia only 
60 to 65 percent head length; all dorsal head scales smooth. 

beckeri (p. 62) 

At least a few vertebral scales slightly enlarged, keeled; tibia longer in propor- 


tion to head length; dorsal head scales smooth or not__.------------ 20 
Wpper head scales tricarinates = 2522 eee eee baccatus (p. 62) 
Upper head scales smooth, slightly rugose, or feebly unicarinate; postanals 

enlanged.im males 5 oe en ee I 2 UE ak 21 
One series of 3 enlarged supraoculars; male lacking first digit on all four feet *’; 

dewlspere dnt aa nae Ph ee eee liogaster (p. 62) 
Two series of 7 to 9 enlarged supraoculars; males with 5 digits on each foot; 

dewlapr yellows ass eee eee eee ee eee aD laeviventris (p. 62) 
Tibia distinctly longer than head (15.5 mm. and 13.5 mm. respectively in the 

typemnGronly, known specimen) i2-- 252" se "sesso eee schiedii (p. 63) 
Head (snout to anterior border of ear opening) at least a little longer than 

DES ERR a: pik ep de pa ent laa se we OE BOE 23 
Interparietal plate twice as long as ear opening, much larger than adjacent, 

very smallcheadiscalests Se) Naisow st jeu Sie ee guntherii (p. 63) 
Interparietal plate not more than 1!4 times as long as ear opening, not more 

than<4tinieg as large as adyacent scales ses 22225009. Bee oe es ee 24 


Ventral scales only slightly larger than dorsals; one large scale in anterolateral 
corner of supraocular area, in contact with superciliaries and separated from 


enlarged supraoculars by one row of small scales___------- milleri (p. 64) 
Ventral scales much larger than dorsals_-_----- limifrons rodriguezii (p. 64) 
infead shorter thandipias 00. ooo eet ee eee ee 26 


Head longer than tibia 


37 Materia Irecently examined demonstrates that this species like others of Anolis possesses five digits on 
both limbs in both sexes. 


58 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


26. Ventrals very strongly keeled; dorsal scales smooth, no larger than laterals; 


head\ broad... 282205 22k ee ee ee ee See capito (p. 65) 
Ventrals very weakly keeled or smooth; dorsal scales keeled, somewhat larger 
thanlaterals; head longers4252 52 322 26 Wasted ese sk schiedii (p. 63) 


27. Lamellae under second and third phalanges of fourth toe, about 27; inter- 
parietal scarcely enlarged, much smaller than ear opening_--petersii (p. 65) 
Lamellae under second and third phalanges of fourth toe, 23 or less; inter- 


parietal: variables.220/4 Seen ee See on Fe eee See eae 28 

28. Two or 3 rows of scales between supraorbital semicircles__._.__.._..------- 29 
Supraorbital semicircles in contact or separated from each other by no more 
than. T4rowsOfiseales sets © eee ee een Ae ee te eee cae ee 30 

29. Ventrals very strongly keeled; 22 or 23 lamellae under phalanges 2 and 3 of 
fourth oem. 5.2. -sos2cessuse sess esec ese ne ssa biporcatus (p. 65) 
Ventrals very weakly keeled; not over 16 or 17 lamellae________.-__--- 24 

30. A single row of not more than 5 large supraoculars______-_-.--------- 31 
Two or more rows of 7 or more enlarged supraoculars___.-__-.-------- 35 

31. Ventrals very weakly keeled; postanals enlarged and dewlap yellow in males; 
supraorbital semicircles broadly in contact__----_------- schmidti (p. 60) 
Ventrais: very strongly keeled. 5 ek Seas eee. ee eee 32 

32. Males with enlarged postanals and dewlap more or less unicolor____---- 33 
Males without enlarged postanals and dewlap with a conspicuous central 
dark. blue:spotsss-2s65225 26. 5 See Ee See eee 34 

33. Dorsal scales very slightly smaller than ventrals_____..---- nebulosus (p. 65) 
Dorsal scales markedly smaller than ventrals_-__-------- nebuloides (p. 66) 


34. Supraorbital semicircles in contact with each other and with interparietal. 
kidderi (p. 66) 
Supraorbital semicircles separated from each other (generally) and from inter- 


parietal«(Ginvariably)iz.2 sex tute eee o Ss eee. Ses eee ustus (p. 66) 

35. Interparietal little if any larger than ear opening; dewlap in males without a 
centralublue spotisso << pans ca ce et ea ee ene rs ee ee 36 
Interparietal nearly or fully twice as long as ear opening; dewlap in males with 

a, central bluevspotiornol: 2-2-2 225-252 ee oe re 38 

36. Ventral scales indistinctly keeled; dorsal scales granular except two vertebral 
rows OlMabruptly enlarged scaless=-2 22252 >see ease ee cymbops (p. 62) 
Ventral scales strongly keeled; dorsal scales uniform in size in both vertebral 
andiparavertebral areas: A222 oc 2220 $2523 soo eee ee 37 


37. Dorsals very small, much smaller than ventrals; 17-19 lamellae under pha- 
langes 2 and 8 of fourth toe; 6-8 rows of loreals_-_lemurinus bourgeaei (p. 66) 
Dorsals rather large, distinctly smaller than ventrals; 15 or 16 lamellae; 4-6 
LOWS OL lOrealst< on sath sae Se ee ae eee cumingii (p. 67) 

88. Dorsals very slightly smaller than ventrals, in about 14 rows; a purple spot in 
center of gular fan in males; postanals not enlarged___-heliactin (p. 67) 
Dorsals smaller; a purple spot or not; postanals enlarged or not__-_---- Mri. 39 

39. Both dorsals and ventrals smaller; no purple spot in center of dewlap; post- 
analsienlanged unpmaless 22222022 eee eee laeviventris (p. 62) 
Dorsals and ventrals larger; a purple spot in center of dewlap; postanals not 
enlarged: in. males.” i252 =i Sc See et eee sericeus (p. 67) 


ANOLIS BARKERI Schmidt 
Anolis barkert Scumipt, Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, 1939, 
pp. 7-9, fig. 2. 
Type.—Brit. Mus. Nat. Hist. No. 36.6.6.12; R. Wright Barker 
collector. 


CHECKLIST OF REPTILES OF MEXICO 59 


Type locality —Cascajal, upper Uzpanapa River, Veracruz. 
Range.—Known only from the type locality. 


ANOLIS SAGREI MAYENSIS Smith and Burger 


Anolis sagrei mayensis SMiTH and BurceEr, Anal. Inst. Biol., vol. 20, 1949, p. 407, 


Type.—K. H. Taylor No. 11523; H. M. Smith collector. 

Type locality—Panlao, Campeche (a small island at mouth of Rio 
Mamantel, Laguna de Términos). 

Range.—The Atlantic coast from central Mexico to northern South 
America. Recorded in Mexico from the states of Yucatdn: Progreso, 
Mérida, Chichen Itzi; Campeche: Balchacaj, Ciudad del Carmen, 
Panlao; Tabasco: San Juan Bautista; Quintana Roo: Cozumel Island. 


ANOLIS COZUMELAE Smith 


Anolis cozumelae Smitru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, 1939, 
pp. 19-20, fig. 4 (at left). 
Type.—Chicago Nat. Hist. Mus. No. 751; male; C. F. Millspaugh 
collector. 
Type locality Cozumel Island, Yucatén Peninsula. 
Range.—Cozumel Island, Quintana Roo. 


ANOLIS METALLICUS Bocourt *% 


Anolis metallicus Bocourt, Ann. Sci, Nat., ser. 5, vol. 17, art. 2, 1873, p. 1; 
Mission scientifique au Mexique . . ., Etudes sur les reptiles, livr. 2, 1878, 
pl. 17 bis, fig. 1. -BouLenaER, Catalogue of the lizards in the British Museum, 
vol. 2, 1885, p. 84. 

Anolis uniformis, BarBour (part), Bull. Mus. Comp. Zool., vol. 77, No, 4, 1934, 
p. 1538. 

Anolis tropidonotus, Stuart (part), Occ. Pap, Mus. Zool. Univ. Michigan, No. 464, 
1942, p. 2. 


Type.—Mus. Hist. Nat. Paris. 


Type locality —Mexico. 
Range.—Unknown. 


ANOLIS MEGAPHOLIDOTUS Smith 
Anolis megapholidotus Smiru, Trans. Kansas Acad. Sci., vol. 36, 1933, pp. 318-320. 


Type-—EHT-HMS No. 11149 (originally EHT-HMS field No. 
1509); E. H. Taylor and H. M. Smith collectors. 

Type locality—Between Rincén and Cajones (about 40-45 km. 
south of Chilpancingo), Guerrero. 

Range.—Known only from the type locality; presumably ranges 
throughout the Sierra Madre del Sur in Guerrero. 

88 Barbour (loc. cit.) synonymizes this with A. h. wniformis, and Stuart (Joc. cit.) suggests synonymy with 

A. tropidonotus. Inasmuch as both the original description and the subsequent illustration indicate the 


possession by the type of relatively small dorsal scales equal in size to the ventrals, while in both tropidonotus 
and wniformis the dorsals are much larger, we regard metallicus as distinct from either. 


861316—50—_5 


60 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


ANOLIS HUMILIS UNIFORMIS Cope 


Anolis uniformis Corr, Proc. Amer. Philos. Soc., vol. 22, 1885, p. 392.— 
BouLenceER, Catalogue of the lizards in the British Museum, vol. 2, 1885, 
p. 84.—Srvuart, Occ. Pap. Mus. Zool. Univ. Michigan, No. 464, 1942, pp. 1-2. 

Anolis humilis uniformis, Stuart, Mise. Publ. Mus. Zool. Univ. Michigan, No. 
69, 1948, p. 48. 

Anolis ruthveni Stuart, Occ. Pap. Mus. Zool. Univ. Michigan, No. 310, 1935, 
pp. 1-4 (type, Univ. Michigan Mus. Zool. No. 76622, 2 miles north of Santa 
Teresa, El Petén, Guatemala). 

Type.—U.S.N.M., Nos. 6774, 2473448, 24750 (Guatemala), 24859 
(Yucatén), cotypes; Mus. Comp. Zool. No. 10933 (Guatemala). 

Type locality —Yucatin and Guatemala, here restricted to 2 miles 
north of Santa Teresa, Guatemala. 

Range.—The southern portion of the Yucatén Peninsula, from 
Chiapas and probably Tabasco to El Petén and British Honduras. 
Known in Mexico from Chiapas: Usumacinta River (across from 
Piedras Negras, El] Petén, Guatemala), Tlacotalpa, Mineral de Santa 
Fé; Tabasco: Teapa. 

ANOLIS TROPIDONOTUS Peters 

Anolis tropidonotus PETERS, Monatsb. Akad. Wiss. Berlin, 1863, p. 135.—Bocourrt, 
Mission scientifique au Mexique . . ., Etudes sur les reptiles, livr. 2. 1873, 
pp. 103-105, pl. 16, fig. 30.—BouLENGER, Catalogue of the lizards in the 
British Museum, vol. 2, 1885, pp. 83-84. 

Norops tropidonotus O’SHAuGHNEssy, Ann. Mag. Nat. Hist., ser. 4, vol. 4, 1869, 

. 273, 

sige yucatanicus BARBouR and Co.x, Bull. Mus. Comp. Zool., vol. 50, 1906, 
p. 149 (type, Mus. Comp. Zool. No. 7036, Chichen Itz4, Yucatdn). 

Type.—Zool. Mus. Berlin, 6 specimens; Dr. Hille collector. 

Type locality —Huanuco (sic) = probably, Huatusco, Veracruz. 

Range.—Atlantic slopes from Veracruz to Nicaragua, including the 
Yucatén Peninsula. Recorded from the states of Veracruz, Tabasco, 
Oaxaca, Campeche, Yucatén, and Quintana Roo (Cob4). 


ANOLIS IMPETIGOSUS Cope 


Anolis impetigosus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 174.— 
BouLENGER, Catalogue of the lizards in the British Museum .. ., ed. 2, 
1885, p. 55, pl. 2, fig. 3. 


Type.—Brit. Mus. Nat. Hist. 
Type locality —Unknown. 
Range.—Unknown. 


ANOLIS SCHMIDTI Smith 
Anolis schmidti Smitu, Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, No. 4, 
1939, pp. 21-23, fig. 4 (at right). 
Type.—Chicago Nat. Hist. Mus. No. 1667; C. H. T. Townsend 
collector. 
Type locality—Manzanillo, Colima. 
Range.—Known only from the type locality. 


CHECKLIST OF REPTILES OF MEXICO 61 
ANOLIS GADOVII Boulenger 


Anolis gadovit BouLENGER, Proc. Zool. Soc. London, 1905, vol. 2, p. 245, pl. 6, 
fig. 1—MosauvEr, Herpetologica, vol. 1, no. 2, 1936, pp. 61-63.—Smiru and 
SPreLER, Copeia, 1945, pp. 165-168. 

Type.—Brit. Mus. Nat. Hist.; Hans Gadow collector. 
Type locality —Tierra Colorada, Guerrero. 
Range.—Known only from the type locality. 


ANOLIS DUNNI Smith 


Anolis gadovii, SmitH (part), Trans. Kansas Acad. Sci., vol. 36, 1933, pp. 316-318. 
Anolis dunni Smiru, Copeia, 1936, p. 9.—SmitrH and Sprever, Copeia, 1945, pp. 
165-168. 
Type.—EHT-HMS No. 1506; E. H. Taylor collector. 
Type locality—Between Rincén and Cajones, Guerrero (a place 
now called Agua del Obispo). 
Range.—Known only from the type locality. 


ANOLIS TAYLORI Smith and Spieler 


Anolis taylort SMitH and SpreLEerR, Copeia, 1945, pp. 165-168. 


Type.—U.S.N.M. (H. M. Smith field No. 10085); E. H. Taylor 
collector. 

Type locality —One mile north of Acapulco, Guerrero. 

Range.—Known only from the type locality. 


ANOLIS DAMULUS Cope 


Anolis damulus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 169.—Bov- 
LENGER, Catalogue of the lizards in the British Museum . . ., ed. 2, vol. 2, 
1885, pp. 47-48, pl. 2, fig. 2. 


Type.—Brit. Mus. Nat. Hist. 
Type locality Unknown. 
Range.—Unknown. 

ANOLIS PENTAPRION Cope 


Anolis (Coccoéssus) pentaprion Corr, Proc. Acad. Nat. Sci. Philadelphia, 1862, 
p. 178. 

Anolis pentaprion, BOULENGER, Catalogue of the lizards in the British Museum, 
ed. 2, vol. 2, 1885, p. 64. 

Anolis sulcifrons Core, Sci. Bull. Philadelphia Mus., No. 1, 1899, p. 6, pl. 2, fig. 1 
(type, Amer. Mus. Nat. Hist. No. 38750; Baranquilla, Colombia). 

Anolis panamensis BOULENGER, Proc. Zool. Soc. London, 1890, p. 81, pl. 13, fig. 2 
(type, Brit. Mus. Nat. Hist. No. 89-17—2-31, 2 cotypes; Panama). 


Type.—F¥ormerly in U.S.N.M., now lost; A. Schott collector. 

Type locality —Truando River, Colombia. 

Range.—Atlantic slopes from the Isthmus of Tehuantepec to 
Colombia. Known in Mexico only from Chiapas: Palenque. 


62 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 
ANOLIS UTOWANAE Barbour 


Anolis utowanae BArRour, Copeia, 1932, p. 11. 


Type.—Mus. Comp. Zool. No. 31035; Thomas Barbour collector 
(purchased from Indian). 

Type locality—Ten miles north of Mazatlan, Sinaloa. 

Range.—Coastal Sinaloa, known only from the type locality. 


ANOLIS CYMBOPS Cope 


Anolis cymbops Corr, Proc. Acad. Nat. Sci. Philadelphia, 1864, pp. 173-174.— 
Bovu.LenceEr, Catalogue of the lizards in the British Museum, vol. 2, 1885, 
pp. 73-75. 
Type.—Brit. Mus. Nat. Hist. 
Type locality.—‘‘ Veracruz,’ Mexico. 
Range.—Known only from the type locality. 


ANOLIS BECKERI Boulenger 


Anolis beckeri BoULENGER, Proc. Zool. Soc. London, 1881, pp. 921-922; Cata- 
logue of the lizards in the British Museum, vol. 2, 1885, pp. 46-47. 
Type.—Royal Belg. Mus., two cotypes; A. Boucard, collector. 
Type locality.— Yucatan. 
Range.—Yucatan Peninsula, including British Honduras. Re- 
corded in Mexico only from Yucatén: Chichen Itza. 





ANOLIS BACCATUS Bocourt 


Anolis baccatus Bocourt, Mission scientifique au Mexique . . ., Etudes sur les 
reptiles, livr. 2, 1873, pp. 59-61, pl. 14, fig. 14.—BouLrencEr, Catalogue of 
the lizards in the British Museum, vol. 2, 1885, p. 54. 

Type —Mus. Hist. Nat. Paris; Auguste Sallé collector. 

Type locality Mexico. 

Range.—Uncertain. The specimen reported from Sepaquite, Alta 
Verapaz, Guatemala (Barbour, Bull. Mus. Comp. Zool., vol. 77, 1934, 
p. 124), is actually A. sericeus (fide L. C. Stuart, in litt.). No definite 
Mexican records are known. 


ANOLIS LIOGASTER Boulenger 


Anolis liogaster BOULENGER, Proc. Zool. Soc. London, 1905, vol. 2, pp. 245-246, 
pl. 6, fig. 2. 


Type.—Brit. Mus. Nat. Hist., two cotypes; Hans Gadow collector. 
Type locality —Omilteme, Guerrero (7,600 feet). 
Range.—Known only from the type locality.” 


ANOLIS LAEVIVENTRIS (Wiegmann) 
D{actyloa] (A[nolis]) laeviventris W1racMann, Herpetologia Mexicana, 1834, p. 47. 


39 Gadow (Proc. Zool. Soe. London, 1905, pp. 195, 216) suggests also the locality Nevado de Colima, Jalisco. 
The record is undoubtedly unacceptable. 


CHECKLIST OF REPTILES OF MEXICO 63 


Anolis laeviventris, Bocourt, Mission scientifique au Mexique . . ., Etudes sur 
les reptiles, livr. 2, 1873, pp. 87-88; livr. 3, 1874, pl. 16, fig. 18, 18a.—Bov- 
LENGER, Catalogue of the lizards in the British Museum, vol. 2, 1885, p. 76.— 
Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, 1939, pp. 20-21. 

Anolis wiegmanni Firzincer, Systema reptilium, 1843, p. 67 (substitute name for 
Dactyloa laeviventris Wiegmann). 

Anolis nannodes Cope (part only; a lectoparatype), Proc. Acad. Nat. Sci. Phila- 
delphia, vol. 16, 1864, p. 173 (lectotype, U.S.N.M. No. 12200, Arriba, Costa 
Rica). 

Anolis intermedius, BoULENGER, Catalogue of the lizards in the British Museum, 
ed. 2, vol. 2, 1885, pp. 78-79 (part). 


Type.—Zool. Mus. Berl.; F. Deppe collector. 

Type locality.— Mexico, here restricted to Jalapa, Veracruz. 

Range.—Atlantic slopes from Central Veracruz to the Isthmus of 
Tehuantepec. Recorded from the states of Veracruz: Jalapa, 
Orizaba, Mirador, Actopam, Jicaltepec, Xico; and Tabasco." 


ANOLIS SCHIEDII (Wiegmann) 


D{actyloa] schiedii W1EGMANN, Herpetologia Mexicana, 1834, p. 48. 

Anolis schiediit, LICHTENSTEIN and Martens, Nomenclator reptilium et amphib- 
iorum Musei Zoologici Berolinensis, 1856, p. 8—Bocourt, Mission scienti- 
fique au Mexique . . ., Etudes sur les reptiles, livr. 2, 1873, pp. 64-65 
(part)*2, pl. 15, fig. 19 (of type).—?BouLENGER, Catalogue of the lizards in 
the British Museum, vol. 2, 1885, p. 52. 


Type.—Zool. Mus. Berl.; Schiede collector. 
Type locality—M exico (by inference). 
Range.—Unknown.* 


ANOLIS GUNTHERII Bocourt 


Anolis guntherit Bocourt, Mission scientifique au Mexique .. ., Etudes sur les 
reptiles, livr. 2, 1873, pp. 61-62, pl. 15, fig. 15. 


40 The original type series included two, a male and a female, in the Brit. Mus. Nat. Hist., from Cobén, 
Guatemala; one, U.S.N.M. No. 12206, from Arriba, Costa Rica, and one, U.S.N.M. No. 6117, now lost, from 
Jalapa, Veracruz. Dunn (Proc. New England Zool. Club, vol. 12, 1930, p. 18), in discussing A. intermedius 
of Central America, mentions “‘the type of nannodes, U.S.N.M. No. 12206, Costa Rica,”’ stating that it is the 
same as intermedius. We regard this action as constituting a restriction of the name to zntermedius and 
rendering U.S.N.M. No. 12206 the lectotype of nannodes. The Mexican specimen obviously represents 
laeviventris, while the Guatemalan species represented by the Cobén specimens remains unnamed. Stuart 
(Mise. Publ. Mus. Zool. Univ. Michigan, No. 69, 1948, p. 50) restricts the name nannodes to the Guatemalan 
species, inasmuch as a name is available for each of the other two species (now laeviventris of Mexico, inter- 
medius of Costa Rica) represented in the cotypes. This procedure certainly is the most desirable, but in 
view of the fact that Dunn previously restricted the name to the Costa Rican species, Stuart’s course can- 
not, apparently, be maintained. Thename Anolis stuarti Smith, new name, is proposed for the Guatemalan 
species represented by the two cotypes of nannodes in the British Museum of Natural History, from Coban, 
Alta Verapaz. For a description of the types, see Boulenger, Cat. Liz. Brit. Mus., vol. 2, 1885, p. 78. 

The Anolis nannodes (auct.) from numerous localities in Mexico is the species laeviventris —H. M. 8. 

41In various papers (first in Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 123) Cope records this species 
from Yucatén and from ‘‘Tehuantepec”’; Dugés (La Naturaleza, ser. 2, vol. 2, 1896, p. 479) records it from 
Tangancicuaro (Michoacin), Guadalajara (Jalisco), and Cerro de los Amoles (state?); Mocquard (Bull. 
Soc. Philom. Paris, ser. 9, vol. 1, 1899, pp. 155-156) records it from Guadalajara, Jalisco; and Peters (Monatsb. 
Berl. Acad., 1869, p. 874) records it from Matamoros, Puebla. We believe all these records refer to other 
species with smooth ventrals. We are unable at this time to place them accurately. 

42 Bocourt possessed for his description the type of schiedii and a series of specimens from Cobén, Guate- 
mala; all the latter are, however, apparently referable to A. cobanensis Stuart. 

43 Cope’s record (U.S. Nat. Mus. Bull. 32, 1887, p. 31) for Jalapa, Veracruz, is highly questionable. 


64 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Anolis guentherit, BouLENGER, Catalogue of the lizards in the British Museum, 
vol. 2, 1885, pp. 54-55. 


Type.—Mus. Milano. 
Type locality —Mexico. 
Range.—Unknown. 


ANOLIS MILLERI Smith, new species “4 


Anolis schiedit, Smirw and Laure (nec Wiegmann), Herpetologica, vol. 3, 1945, 
pp. 2-4. 


Type.—U.S.N.M. No. 120957; Walter S. Miller collector. 
Type locality —Quetzaltepec, Oaxaca. 
Range.—Known only from the type locality. 


ANOLIS LIMIFRONS RODRIGUEZII Bocourt 


Anolis rodriguezit Bocourt, Mission scientifique au Mexique . . ., Etudes sur 
les reptiles, livr. 2, 1873, pp. 62-63, pl. 13, fig. 1. 

Anolis limifrons rodriguezii,4® Stuart, Mise. Publ. Mus. Zool. Univ. Michigan, 
No. 69, 1948, pp. 49-50. 

Anolis rubigenosus Bocourt, Ann. Sci. Nat., ser. 5, art. 2, 1873, p. 1; Mission 
scientifique au Mexique . . ., Etudes sur les reptiles, livr. 3, 1874, pl. 17 bis, 
fig. 2 (type, Mus. Hist. Nat. Paris; Oaxaca [state]) .46 

Anolis aureolus Corr, Proc. Amer. Philos. Soc., 1885, pp. 390-391 (Guatemala, 
2 cotypes, U.S.N.M. Nos. 24850-1; Yucatan, 3 cotypes U.S.N.M. Nos. 25854— 
5, 25857, one cotype Mus. Comp. Zool. No. 10929, type locality here restricted 
to Chichen Itz4, Yucatan). 

Anolis acutirostris Ives, Proc. Acad. Nat. Sci. Philadelphia, 1891, pp. 459-460 
(type, Acad. Nat. Sci. Philadelphia No. 7889; Citilpech, Yucatdn). 


Type.—Mus. Hist. Nat. Paris; collected by Commission Scientifique 
du Mexique. 

Type locality —Panzés, Alta Verapaz, Guatemala. 

Range.—Atlantic slopes from the Isthmus of Tehuantepec to 
Honduras, including the peninsula of Yucat&n except its semiarid 
northwestern coast. Recorded in Mexico from Oazaca (no locality) ; 
Chiapas: Palenque; Campeche: Balchacaj, Champotén, Tres Brazos, 
Encarnacién; Yucatdn: Citilpech, Chichen Itz4, Mérida; and Quintana 
Roo: Cozumel Island. 


44 Diagnosis.—As in the accompanying key (p. 57). Differs from schiedii in two important respects: The 
head length from snout to ear (15.8 mm.) is distinctly greater than tibia length (13.6 mm.), and the inter- 
parietal is much larger, its length equaling the diameter of ear opening (the tibia, 15.5 mm., is longer than 
head, 13.5 mm., and length of the interparietal is little more than half the greatest diameter of the ear open- 
ing, in schiedii). A. schiedii is unique, in its group of the genus, in its very long tibia. 

Description—A lengthy description of the type was published in Smith and Laufe (Joc. cit.) and needs no 
repetition.—H. M.S. 

45 Spelled ridriguezii by error. 

46 The type, according to Bocourt’s figure and description, does not differ from the form most authors in the 
past have called aureolus, except perhaps in size of the interparietal. It appears probable to us that the 
deceptive marking on the type of rodriguezii (a broad vertebral light stripe), which Bocourt apparently 
thought was constant, may have been responsible for his failure to consider rodriguezii and the type of his 
rubigenosus as conspecific. 

Although no other record indicates the occurrence of rodriguezii in Oaxaca, the known distribution else- 
where strongly implies occurrence on Atlantic slopes of that state near or at the Isthmus of Tehuantepec. 


CHECKLIST OF REPTILES OF MEXICO 65 


ANOLIS CAPITO Peters 


Anolis (Draconura) capito Peters, Monatsb. Acad. Wiss. Berlin, 1863, p. 142. 

Anolis capito, Bocourt, Mission scientifique au Mexique . . ., Etudes sur les 
reptiles, livr. 2, 1873, pl. 16, fig. 27—BouLEenesErR, Catalogue of the lizards 
in the British Museum, vol. 2, 1885, pp. 94-95. 

Anolis carneus Coreg, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 169 (Brit. Mus. 
Nat. Hist., two cotypes; lower Verapaz forest). 


Type.—Zool. Mus. Ber. No. 4086, two cotypes; C. Hoffman collector. 
Type locality Costa Rica, here restricted to Palmar. 
Range.——Tabasco to Panama. Recorded in Mexico only from 
“Tabasco.” 
ANOLIS PETERSII Bocourt 


Anolis petersii Bocourt, Mission scientifique au Mexique . . ., Etudes sur les 
reptiles, livr. 2, 1873, pp. 79-80, pl. 13, fig. 2, and pl. 15, figs. 11, lla.— 
BouLENGER, Catalogue of the lizards in the British Museum, vol. 2, 1885, pp. 
66-67. 

Anolis petersii bivittatus WERNER, Verh. zool.-bot. Ges. Wien, vol. 46, 1898, p. 351 
(Munich Mus., two cotypes; Guatemala, here restricted to Cobdn). 


Type.—Mus. Hist. Nat. Paris No. 1641 8, two specimens; collected 
by Commission Scientifique du Mexique. 

Type locality—Alta Verapaz, Guatemala, here restricted to Finca 
Samac. 

Range.—Atlantic slopes from central Veracruz to Alta Verapaz in 
Guatemala. Recorded in Mexico only from Veracruz: Cuautlapan, 
Mirador, Orizaba; San Luis Potosi: Xilitla. 


ANOLIS BIPORCATUS (Wiegmann) 


D.{actyloa] biporcata WiEGMANN, Herpetologia Mexicana, 1834, p. 47. 

Anolis biporcatus, ScuHmiptT, Publ. Field Mus. Nat. Hist., zool. ser., vol. 22, 1941, 
p. 491.—Smiru, Rev. Soc. Mex. Hist. Nat., vol. 7, 1946, pp. 66-67.—StTuart, 
Misc. Publ. Mus. Zool. Univ. Michigan, No. 69, 1948, pp. 46-47. 

Anolis copei Bocourt, Mission scientifique au Mexique. . ., Etudes sur les 
reptiles, livr. 2, 1873, pp. 77-78, pl. 15, fig. 10 (Panzos, Alta Verapaz, Guate- 
mala; Mus. Hist. Nat. Paris) —BouLENGER, Catalogue of the lizards in the 
British Museum, vol. 2, 1885, pp. 65-66. 


Type.—Zool. Mus. Berlin; F. Deppe collector. 

Type locality —Mexico, here restricted to Piedra Parada, Chiapas. 

Range.—Chiapas to Panama. Known in Mexico only from Chiapas 
(Piedra Parada). 


ANOLIS NEBULOSUS (Wiegmann) 


D{actyloa] nebulosa WrEGMANN, Herpetologia Mexicana, 1834, p. 47. 

Anolis nebulosus, Bocourt, Mission scientifique au Mexique . . . , Etudes sur les 
reptiles, livr. 2, 1873, pp. 68-69.—BouLENGER, Catalogue of the lizards in 
the British Museum, vol. 2, 1885, pp. 76-77. 

Anolis boulengerianus THominot, Bull. Soc. Philom., ser. 7, vol. 11, 1887 (Mus. 
Hist. Nat. Paris, three cotypes, Isthmus of Tehuantepec, F. Sumichrast 
collector). 


Type.—Zool. Mus. Berlin, two cotypes; F. Deppe collector. 


66 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type locality.—Mexico (by inference), here restricted to Mazatlan, 
Sinaloa. 

Range.—Western Mexico, from Sinaloa to the Isthmus of Te- 
huantepec. Known from the states of Morelos, Puebla, Nayarit, 
Colima, Oaxaca, Guerrero, Michoacan, Sinaloa, Jalisco, Durango, 
and the Tres Marias Islands.” 


ANOLIS NEBULOIDES Bocourt 


Anolis nebuloides Bocourt, Mission scientifique au Mexique . . ., Etudes sur les 
reptiles, livr. 2, 1873, pp. 74-75, pl.13, fig. 10.—BouLencrer, Catalogue of 
the lizards of the British Museum, vol. 2, 1885, p. 77. 

Type.—Mus. Hist. Nat. Paris, four cotypes; collected by (?) Com- 
mission Scientifique du Mexique. 

Type locality —Putla, Oaxaca. 

Range.—Western Mexico from Chihuahua to Oaxaca. Recorded 
from the states of Chihuahua, Sonora, Colima, Guerrero, Puebla, 
and Oaxaca.* 

ANOLIS KIDDERI Ruthven 
Anolis kiddert Ruruven, Occ. Pap. Mus. Zool. Univ. Michigan, No. 257, 1933, 
pp. 1-2. 

Type-—Mus. Zool. Univ. Michigan No. 72851; Edwin P. Creaser 
collector. 

Type locality.—Mérida, Yucatan. 

Range.—The western part of the Yucatan Peninsula. Recorded 
from Yucatin and Campeche. 


ANOLIS USTUS Cope 


Anolis ustus Cops, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 172.— BouLENGER, 
Catalogue of the lizards in the British Museum, vol. 2, 1885, p. 73.—Scumint, 
Publ. Field Mus. Nat. Hist., zool. ser., vol. 22, No. 8, 1941, p. 493.—Srvart, 
Mise. Publ. Mus. Zool. Univ. Michigan, No. 69, 1948, pp. 49, 51. 


Type.—Brit. Mus. Nat. Hist., two cotypes. 

Type locality—Belize, British Honduras. 

Range.—The Yucatan Peninsula and the area at its base. Recorded 
in Mexico from Yucatan and Campeche. 


ANOLIS LEMURINUS BOURGEAEI Bocourt 


Anolis bourgeaei Bocourt, Mission scientifique au Mexique . . ., Etudes sur les 
reptiles, livr. 2, 1873, pp. 76-77, pl. 15, fig. 9 -BouLENGER, Catalogue of the 
lizards in the British Museum, vol. 2, 1885, pp. 74-75. 

Anolis limifrons bourgeaei, Stuart, Misc. Publ. Univ. Michigan Mus. Zool., 
No. 69, 1948, p. 49. 


47 Records from the states of Veracruz and Yucat&n are probably erroneous, and that from Chihuahua is 
said to be referable to A. nebuloides. Virtually all records for both species are, for that matter, open to 
question. The two forms have never been adequately characterized, and very likely more than two species 
are involved. 

48 Undoubtedly some records of nebulosus belong here, and perhaps vice versa. A record for Veracruz is 
probably in error. 


CHECKLIST OF REPTILES OF MEXICO 67 


Anolis biporcatus, Bocourt*® (nec Wiegmann), Mission scientifique au Mexique..., 
Etudes sur les reptiles, livr. 2, 1873, pp. 98-99, pl. 15, fig. 8 —BouULENGER, 
op. cit., pp. 88-89. 

Anolis ustus veraepacis BARBouR, Proc. New England Zool. Club, vol. 12, 1932, 
p-. 98 (type, Mus. Comp. Zool. No. 32324; A. W. Anthony coll.; Hacienda 
Chimoxdn, Alta Verapaz, Guatemala). 


Type—Mus. Hist. Nat. Paris, one female (Orizaba, collected by 
Bourgeau); Zool. Mus. Berlin, one male (Huatusco, collector?). 

Type locality—Huatusco and Orizaba, Veracruz, here restricted to 
Huatusco. 

Range.—Atlantic slopes from Veracruz to Guatemala and British 
Honduras. Recorded in Mexico from the states of Veracruz, Tabasco, 
Campeche, and Yucatan. 


ANOLIS CUMINGII Peters 


Anolis cumingii Prrrers, Monatsb. Akad. Wiss. Berlin, 1863, p. 140.—Bocourt, 
Mission scientifique au Mexique . . . , Etudes sur les reptiles, livr. 2, 1873, 
p. 89, pl. 16, fig. 20——BouLencsmr, Catalogue of the lizards in the British 
Museum, vol. 2, 1885, p. 80.—Burr and Mysrs, Stanford Univ. Publ., 
biol. sci., vol. 8, 1942, p. 291.—Smirn, Anal. Inst. Biol. México, vol. 14, 1942, 
p. 342. 


Type.—Zool. Mus. Berlin; Herr Cuming collector. 

Type locality.—Mexico. 

Range.—Uncertain; possibly the upper Balsas Basin. Recorded 
only from La Paz (Puebla?). 


ANOLIS HELIACTIN Cope 


Anolis heliactin Corr, Proc. Acad. Nat. Sci. Philadelphia, 1864, pp. 172-173.— 
Bocourt, Mission scientifique au Mexique . . . , Etudes sur les reptiles, 
livr. 2, 1873, pp. 106-108, pl. 18, figs. 4, 4a-c (in color), pl. 16, fig. 32.— 
BouLENGER, Catalogue of the lizards in the British Museum, vol. 2, 1885, 
p. 72. 


Type.—Acad. Nat. Sci. Philadelphia No. 7914. 
Type locality —Mexico. 
Range.—Recorded only from ‘‘Oaxaca.”’ 


ANOLIS SERICEUS Hallowell 


Anolis sericeus HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, vol. 8, 1856, 
pp. 227—228.—Barsour, Bull. Mus. Comp. Zool., vol. 77, 1934, pp. 149-150.— 
Scumipt, Publ. Field Mus. Nat. Hist., zool. ser., vol. 22, 1941, p. 492. 

Anolis sallaei GiNTHER, Proc. Zool. Soc. London, 1859, p. 421 (Brit. Mus. Nat. 
Hist.; Mexico; Salle coll.).— Bocourt, Mission scientifique au Mexique . . . , 
Etudes sur les reptiles, livr. 2, 1873, p. 90, pl. 13, fig. 3, pl. 16, fig. 21.— 
BouLENGER, Catalogue of the lizards in the British Museum, vol. 2, 1885, 
pp. 79-80. 


49 And of other authors prior to Schmidt, 1941; see synonymy of A. biporcatus. 
8 According to Stuart (op. cit., 1948, p. 51) the type and several of the paratypes of A. wu. veraepacis Barbour 
are referable to A. bourgeaei while the remaining paratypes are A. sericeus. 


68 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Anolis jacobi Bocourt, op. cit., p. 74, pl. 18, fig. 8 (Mus. Hist. Nat. Paris; Vera- 
cruz, here restricted to Veracruz). 


Type.—Acad. Nat. Sci. Philadelphia, now lost; Mr. Pease donor. 

Type locality —E| Encero de Jalapa, Veracruz. 

Range.—Atlantic slopes from Tamaulipas and Pacific slopes from 
the Isthmus of Tehuantepec south to Nicaragua. Recorded in Mexico 
from the states of Tamaulipas, San Luis Potosi, Veracruz, Oaxaca, 
Tabasco, and Chiapas. 


Genus CORYTHOPHANES Boie 


Cory tophanes {|sic] Boin, in Schlegel, Isis von Oken, vol. 20, pt. 3, 1826, p. 290. 

Chamaeleopsis WIEGMANN, in Gray, in Griffith, The animal kingdom ... by 
the Baron Cuvier ... , vol. 9, 1831, Synopsis, p. 45 (type, Chamaeleopsis 
hernandezii Wiegmann) .*! 


Genotype.—Agama cristata Merrem. 

Species.—Three. 

Range.—Central Veracruz and the Pacific coast of Chiapas south- 
eastward to Costa Rica. 


KEY TO SPECIES OF CORYTHOPHANES 


1. Nuchal crest interrupted, not continuous with dorsal crest_hernandezii (p. 68) 
Nuchal crest continuous with dorsal, although sometimes low where the two 
TCC Ue pay see ea ee en 2 

2. Parietal crests joining very near extremity of helmet; upper head scales keeled 
OLAT UL OSC as See eee ee Ree ek ge one arene page a. Peete percarinatus (p. 69) 
Parietal crests joining about halfway from orbits to extremity of helmet; upper 
headiscalesssmoothes ss a. se nem Sree eee Sm ee ee cristatus (p. 69) 


CORYTHOPHANES HERNANDEZII (Wiegmann) 


Chamaeleopsis hernandezit W1EGMANN, in Gray, in Griffith, The animal kingdom 
. . .. by the Baron Cuvier . . ., vol. 9, 1831, Synopsis, p. 45; Isis von Oken, 
1831, p. 298; Herpetologia Mexicana, pt. 1, 1834, pp. 15, 37, pl. 6. 

Corythophanes hernandezii, LicHTENSTEIN and Martens, Nomenclator reptilium 
et amphibiorum Musei Zoologici Berolinensis, 1856, p. 8.—BouULENGER, 
Catalogue of the lizards in the British Museum, vol. 2, 1885, pp. 103-104. 

Corythophanes chamaeleopsis DumM®&RIL and Brsron, Erpétologie générale, vol. 4, 
1837, p. 175-177 (substitute name for Chamaeleopsis hernandezii Gray). 

Corythophanes mexicanus Bocourt, Mission scientifique au Mexique . . ., Etudes 
sur les reptiles, livr. 3, 1874, pp. 122-123, pl. 17, fig. 1 (substitute name for 
Chamaeleopsis hernandezii Wiegmann, adopting Herndndez’s pre-Linnaean 
name Chamaeleo Mexicanus) .* 





5t The author of this generic name, and of the specific name as well, has been accepted as Gray by some 
authors, as Wiegmann by others. Wiegmann published his own description in 1831, in great detail; Gray 
published but a few words, also in 1831, but attributed both names to ‘‘Wiedemann, MSS” (in error; = 
Wiegmann). Even though Gray’s description may have preceded Wiegmann’s, it seems obvious that 
Wiegmann was ethically the author, having provided the only reasonable description, and his manuscript 
having been acknowledged by Gray as the source of the latter’s information. It is true that Gray stated 
specimens were in the British Museum (Boulenger cites one purchased by Gray), so that Gray could well 
have drawn up his description from them and not from Wiegmann’s manuscript. The correct authorship 
thus remains debatable. 

82 Wiegmann cited Chamaeleo Mezicanus of Hernfndez in both of his papers (1831 and 1834) but did not 
adopt the name. 


CHECKLIST OF REPTILES OF MEXICO 69 


Type.—Zool. Mus. Berlin; F. Deppe collector. 

Type locality —Mexico, here restricted to Jalapa, Veracruz. 

Range.—Atlantic slopes from central Veracruz south and east to 
Guatemala. Recorded in Mexico from various localities in the states 
of Veracruz, San Luis Potosi (Xilitla), Oaxaca, Chiapas, Yucatén, 
and Quintana Roo (Cobé). 


CORYTHOPHANES CRISTATUS (Merrem) 


Agama cristata MrrreM, Tentamen systematis amphibiorum, 1821, p. 50. 

Corythophanes cristatus,3 GRAVENHORST, Acta Acad. Leop.-Carol., vol. 16, 1833, 
p. 938, pl. 65, figs. 6-10.—Dumé&rin, Arch. Mus. Hist. Nat. Paris, vol. 8, 
1856, p. 517, pl. 20, fig. 1.—BovuLEN@ER, Catalogue of the lizards in the 
British Museum, vol. 2, 1885, pp. 101-102. 


Type.—Unknown. 
Type locality—Ceylon (in error), here restricted to Orizaba, Veracruz. 
Range.—Atlantic slopes from central Veracruz to Costa Rica. 
Reported in Mexico from the states of Veracruz, Campeche, Yucatan, 
and Chiapas (Usumacinta River across from Piedras Negras, Petén, 
Guatemala). 
CORYTHOPHANES PERCARINATUS Duméril 


Corythophanes percarinatus Dumf&rit, Arch. Mus. Hist. Nat. Paris, vol. 8, 1856, 
p. 518, pl. 20, fig. 3—Bocourt, Mission scientifique au Mexique .. ., 
Etudes sur les reptiles, livr. 3, 1874, pp. 120-122, pl. 17, fig. 2—Smiru, Journ. 
Washington Acad. Sci., vol. 39, 1949, p. 34. 

Type.——Mus. Hist. Nat. Paris. 

Type locality —Escuintla, Guatemala. 

Range.—Pacific slopes of Guatemala and Chiapas, and Atlantic 
slopes in Alta Verapaz, Guatemala. Known in Mexico only from Col- 
onia Hidalgo, 8 kilometers north of La Esperanza, Chiapas. 


Genus LAEMANCTUS Wiegmann 


Laemanctus (part) W1rEGMANN, Herpetologia Mexicana, pt. 1, 1834, pp. 45-46. 


Genotype—Laemanctus longipes Wiegmann. 

Range.—Lowlands from San Luis Potosi south to and including 
Guatemala, and British Honduras; ? Colima. 

Species.—Four, all occurring in Mexico. 


KEY TO MEXICAN SPECIES OF LAEMANCTUS 


1. Scales on upper surface of snout much larger than those on occipital region_. 2 
Seales on upper surface of snout not distinctly larger than those on occipital 
region; no dorsal crest; part of scales on body bicarinate or tricarinate; head 
yellowish above, the posterior contour black; body above dark purple or 


88 While Boie designated Merrem’s species as type of his genus Corytophanes, he did not actually print the 
combination. That it was his intent so to spell the name is indicated in his article in vol. 21 (1828) of Isis 
von Oken, p. 363, where he (Boie) spells the name Corytophanes. Kaup (Isis von Oken, vol. 21, 1828, p. 
1147) does likewise. 


70 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


dark green, with blackish cross bands; a light lateral streak; lilac below; 690 


MAW So oe ee i Ee es ser ee deborrei (p. 71) 
2. A series of projecting triangular scales on the posterior contour of the head; a 
Peele Ors CLES 2 Beem pa ae a a a ee 3 


No distinct serration on the posterior contour of head; no dorsal crest present; 
reddish or olive above with darker bands across back; a broad dark band 
from the eye to foreleg, passing through tympanum, bordered by a yellowish 
streak inferiorly; a yellowish streak from axilla to groin sometimes broken; 
a yellow spot on each side of base of tail; lower surfaces pink; 730 mm. 

longipes (p. 71) 

3. Scales around middle of body, 57-61; reddish or purplish above with dark brown 
bands across body, most distinct in the vertebral region; a dark brown band 
from eye to tympanum, sometimes continued along side of back; a white 
streak from below the eye to forelimb, and from axilla to groin; a white spot 
OMREACHYSICE MO Lally ASE O tant a Tle eee ee eee ee serratus (p. 70) 

Seales round middle of body, 45-51; gulars elongate; a less distinct vertebral 
serration; a white spot in front of and one behind thigh. 

alticoronatus * (p. 70) 


LAEMANCTUS ALTICORONATUS Cope 


Laemanctus alticoronatus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1865, p. 192; 
Proc. Amer. Philos. Soec., 1869, pl. 11.—BouLEncErR, Catalogue of the 
lizards of the British Museum, vol. 2, 1885, p. 105. 


Type.—U.S.N.M. No. 308; Arthur Schott collector. 
Type locality —Mérida, Yucatan. 
Range.—Yucatan. Reported from Chichen Itz& and Mérida. 


LAEMANCTUS SERRATUS Cope 


Laemanctus longipes, Dumérit, Arch. Mus. Hist. Nat. Paris, vol. 8, 1856, p. 512, 
pl. 21, fig. 4—Bocourt, Mission scientifique au Mexique... , Etudes sur 
les reptiles, livr. 3, 1874, pp. 114-116, pl. 17, fig. 4 (non Wiegmann, fide 
Cope loc. cit.). 

Laemanctus serratus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 176.— 
BouLENGER, Bull. Soc. Zool. France, vol. 2, 1877, p. 463, pl. 7, fig. 3.— 
Smitu and Laure, Trans. Kansas Acad. Sci., vol. 48, 1945, pp. 229-231.— 
Peters, Chicago Acad. Sci. Nat. Hist. Misc., No. 27, 1948, pp. 1-3, fig. 1. 


Type.—Mus. Leyden. 

Type locality —“Orizaba Valley, Mexico.” 

Range.—Lowlands from extreme eastern Guanajuato to Yucatan. 
Reported from Yucatén: Chichen Itz4; Oaxaca: “Oaxaca” (no specific 
locality), Tlacolula; Veracruz: Boca del Rio, Jicaltepec, Misantla, 
Orizaba; Guanajuato: Huasteca Potosina; Hidalgo: Zacualtipan; Cam- 
peche: Champotén, Oxpemul; San Luis Potosi: Xilitla; Tamaulipas: 
near Gémez Farias. 


& Mrs. Helen T. Gaige suggests the possibility that serratus and alticoronatus may be identical since speci- 
mens from Yucatén and Campeche seem to show the characters of both (Carnegie Inst. Washington Publ. 
No. 457, 1936, p. 296). 


CHECKLIST OF REPTILES OF MEXICO 71 


LAEMANCTUS DEBORREI Boulenger 


Laemanctus deborret BOULENGER, Bull. Soc. Zool. France, vol. 2, 1877, pp. 462- 
463, 465-466, pl. 7, fig. 1; Catalogue of the lizards in the British Museum, 
vol. 2, 1885, p. 106.—Smiru, Journ. Washington Acad. Sci., vol. 34, 1944, pp. 
155-156.—WEBER, Nat. Geogr. Mag., vol. 87, No. 2, 1945, pp. 209, 210, pl. 
14 (col.). 

Type.—Museum de Bruxelles, two adults; M. Linden collector (?). 
Type locality.—Tabasco. 
Range.—Atlantic slopes from Tabasco to British Honduras. 

Reported from Tabasco: La Venta; Oaxaca: Mountains near Santo 

Domingo (U.S. Nat. Mus.). 


LAEMANCTUS LONGIPES Wiegmann 


Laemanctus longipes WIEGMANN, Herpetologia Mexicana, pt. 1, 1834, p. 46, pl. 
4.—Bou.EncER, Bull. Soc. Zool. France, vol. 2, 1877, pp. 464-465, pl. 7, 
fig. 2; Catalogue of the lizards in the British Museum, vol. 2, 1885, pp. 105- 
106. 


Type.—Zool. Mus. Berlin; F. Deppe collector. 

Type locality —Jalapa, Veracruz. 

Range.—Eastern slopes of the plateau from Veracruz to Oaxaca, and 
(?) western slopes in Colima. Reported from Veracruz: Actopam, 
Jalapa, ‘‘near Veracruz,’’ Mirador, Potrero Viejo; ? Colima: Paso del 
Rio; Oaxaca: Oaxaca. 


Genus BASILISCUS Laurenti 


Basiliscus LAURENTI, Specimen medicum exhibens synopsin reptilium, 1768, p. 50. 

Corythaeolus Kaup, Isis von Oken, vol. 21, 1828, p. 1147 (type, Basiliscus vittatus 
Wiegmann). 

Oedicoryphus WactzEr, Natiirliches System der Amphibien, 1830, p. 148 (type, 
Basiliscus vittatus Wiegmann). 

Cristasaura Gray, Ann. Mag. Nat. Hist., ser. 2, vol. 10, 1852, p. 489 (type, 
Cristasaura mitrella Gray = Basiliscus vittatus Wiegmann). 

Paraloma Core, Proc. Acad. Nat. Sci. Philadelphia, 1867, p. 181 (type, Dacon- 
ura bivittata Hallowell= Basiliscus vittatus Wiegmann). 


Genotype.—Basiliscus americanus Laurenti=(Basiliscus basiliscus 
(Linnaeus)). 

Range.—Lowlands of Mexico south through Jalisco and Tamaulipas, 
through Central America to northwestern South America. 

Species.—¥F ive; one occurs in Mexico. 


BASILISCUS VITTATUS Wiegmann 


Basiliscus vittatus W1nGMANN, Isis von Oken, vol. 21, 1828, p. 373.—Dum&mrit, 
Arch. Mus. Hist. Nat. Paris, vol. 8, 1856, pp. 522-524, pl. 21, figs. 2, 3.— 
Bocourt, Mission scientifique au Mexique . . . , Etudes sur les reptiles, livr. 
3, 1874, pp. 129-131, pl. 17, fig. 3—BouLrencmrR, Catalogue of the lizards of 
the British Museum, vol. 2, 1885, pp. 109-110. 

Cristasaura mitrella Gray, Ann. Mag. Nat. Hist., ser. 2, vol. 10, 1852, p. 439 
(Brit. Mus. Nat. Hist.; Dyson coll.; Honduras). 


72 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Daconura bivittata HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, 1860, p. 482 
(Acad. Nat. Sci. Philadelphia; Nicaragua, here restricted to Greytown). 
Basiliscus (Cristasaura) nuchalis Corr, Proc. Acad. Nat. Sci., Philadelphia, 1862, 
p. 181 (U.S.N.M. No. 5845; Caldwell collector; Greytown, Nicaragua). 

Type.—Zool. Mus. Berlin; F. Deppe collector. 

Type locality—Mexico, here restricted to Veracruz, Veracruz. 

Range.—The coasts from Jalisco and southern Tamaulipas, south 
to Nicaragua. Reported from numerous localities in the states of 
Tamaulipas, Veracruz, Tabasco, Campeche, Yucatan, Quintana Roo, 
Chiapas, Oaxaca, Guerrero, Michoacan, Colima, and Jalisco. 


Genus IGUANA Laurenti 


Iguana LavRENTI, Specimen medicum exhibens synopsin reptilium, 1768, p. 47. 
Hypsilophus WacurEr, Natiirliches System der Amphibien, 1880, p. 147 (type, 
Lacerta iguana Linnaeus). 


Genotype.—Lacerta 1guana Linnaeus. 

Range.—Mexico on the Pacific coast from Sinaloa to Chiapas, on 
the eastern coast north to central Veracruz. Absent on the higher 
parts of the plateau. Lowlands of Central America, and central and 
northern South America. 

Species.—Two, one with two subspecies. Extralimital are Iguana 
iguana iguana and Iguana delicatissima of South America. 


IGUANA IGUANA RHINOLOPHA Wiegmann 


I[guana] H{ypsilophus)] rhinolophus WieGMaAnn, Herpetologia Mexicana, pt. 1, 
1834, pp. 44-45. 

Iguana tuberculata var. rhinolopha, BouLENGER, Catalogue of the lizards in the 
British Museum, vol. 2, 1885, pp. 190-191. 

Iguana iguana rhinolopha, VAN DenBuRGH, Proc. Acad. Nat. Sci. Philadelphia, 
1897, p. 461.—Dunn, Copeia, 1934, p. 1. 

?Iguana Hernandesii Jan, Indice sistematico dei rettili ed anfibi esposti nel 
medesino, 1857, p. 58 (nomen nudum). 


Type.—Zool. Mus. Berlin; F. Deppe collector. 

Type locality Mexico, by inference here restricted to Cérdoba, 
Veracruz. 

Range.—Veracruz and Sinaloa southward along both coasts to 
Panama. Reported from Sinaloa: Mazatlan, Presidio; Nayarit: 
San Blas; Colima: Manzanillo, Tecom4n, Colima; Jalisco: ?San Blas 
(possibly Nayarit); Michoacdn: Hacienda California, Chacan River 
Sierra Madre, Jorullo; Guerrero: Pie de la Cuesta, Acapulco; Puebla: 
Chiautla; Oaxaca: Tehuantepec, Rio Coatzacoalcos, ?Tetela, Agua 
Fria, Tequesixtlan, Totolapan; Veracruz: Cérdoba, Vega de Alatorre, 
vicinity of Cuatotolapam, Matacabestro, Puente Nacional; Tabasco: 
Tlacotalpan, Tenosique; Campeche: Ciudad del Carmen; Quintana 
Roo: Cozumel Island; Chiapas: La Esperanza. 


CHECKLIST OF REPTILES OF MEXICO Ge 


Genus CTENOSAURA Wiegmann 


Ctenosaura W1EGMANN, Isis von Oken, 1828, p. 371.—Bat.ey, Proc. U. S. Nat. 
Mus., vol. 73, art. 12, 1928, pp. 1-58, pls. 1-30 (part) .5 

Genotype.—Ctenosaura cycluroides Wiegmann [= C. acanthura 
(Shaw)]. 

Species.—Five, all but one of which (C. bakeri of Utilla Island, 
Honduras) occur in Mexico. 

Range.—Baja California, Sonora, and perhaps southern Arizona 
on the west, and Tamaulipas on the east, south along both coasts 


to Panama. 
KEY TO MEXICAN SPECIES OF CTENOSAURA 


1. Small scales between whorls of enlarged scales reduced to 1 row (and no more 
than 1 row) between at least a few whorls (generally near middle of tail), 


oftenonentire distal/half ormoreiof taili2=siee2 2 ete a ee eee 2 
Small scales in at least one complete and another incomplete row between all 
whoris'of enlarged scales‘on' tail. 25 2258. ws. 2 Sa hee  e eeee 3 

2. Fifth whorl of enlarged scales preceded by more than a single row of small 
S Calle g BEE SANE & SE) Sheesh Oy Eee eye ict peta 2 Ye ee acanthura (p. 74) 
Fifth whorl of enlarged scales preceded by one row and one row only (no in- 
complete row) Of‘small scales. 2. se eee ee hemilopha (p. 75) 

3. Three rows of small scales, complete or incomplete, preceding each of the 
proximal 5 whorls of enlarged caudal scales___________-_-_ pectinata (p. 75) 
Small scales reduced to only two rows preceding one or more of the proximal 
fivekcaudal}whorls ai! {tees oe Ae ae ete eee cee OE eek similis (p. 73) 


CTENOSAURA SIMILIS SIMILIS (Gray) 


Iguana (Ctenosaura) similis Gray, in Griffith, Animal Kingdom, by the Baron 
Cuvier, vol. 9, Synopsis . . . , 1831, p. 38. 

Cyclura (Ctenosaura) similis, WIEGMANN, Herpetologia Mexicana, 1834, p. 42. 

Ctenosaura similis, Battey, Proc. U. 8S. Nat. Mus., vol. 73, 1928, pp. 32-37, pls. 
16-—20.—Smiru, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 139-140; Journ. 
Washington Acad. Sci., vol. 39, 1949, pp. 35-36. 

Ctenosaura similis [similis], BARBOUR and SHREVE, Occ. Pap. Boston Soc. Nat. 
Hist., vol. 8, 1934, p. 197.—Smiru, Ann. Carnegie Mus., vol. 30, 1944, p. 89. 

Ctenosaura completa Bocourt, Mission scientifique au Mexique .. ., Etudes 
sur les reptiles, livr. 3, 1874, pp. 145-146 (Mus. Hist. Nat. Paris Nos. 2251- 
2252, La Unién, El Salvador; Mus. Comp. Zool., 1 spec., ‘‘Mexico’’; Mus. 
Hist. Nat. Paris, 2 cotypes, ‘‘Guatemala’’) .56 


Type.—Lost. 
Type locality.—Restricted (by Bailey) to Tela, Honduras. 
Range.—Coastal areas from the Isthmus of Tehuantepec on the 


55 We are not totally in agreement with Bailey’s treatment of the species in this genus, nor with some of 
the more recent records no doubt influenced by his work. The four forms we recognize are more or less 
completely allopatric; at least such suggestions as of C. acanthura in Chihuahua and C. similis in Guerrero 
appear to us wholly unwarranted. The only extensive overlapping of ranges of which we are aware occurs 
on the Atlantic coast and involves C. acanthura and C. similis. 

56 Bocourt (loc. cit.) states that the types are from ‘“‘Guatemala” and ‘‘La Union,’”’ E] Salvador. Barbour 
and Loveridge (Bull. Mus. Comp. Zool., vol. 69, 1929, p. 247), however, state that a cotype in the Mus. 
Comp. Zool. is from ‘‘Mexico.’’ The type locality is here restricted to La Uni6én. 


74 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Pacific and central Veracruz on the Atlantic south to Panama. 
Recorded in Mexico from the states of Veracruz, Tabasco, Campeche, 
Yucatan, Quintana Roo (including Mujeres and Cozumel Islands), 
Oaxaca, and Chiapas (Tonal, Palenque).°” 


CTENOSAURA ACANTHURA (Shaw) 


Lacerta acanthura SHaw, General zoology, vol. 3, 1802, pp. 216-217. 

Ctenosaura acanthura, Gray, Catalogue of the lizards in the collection of the 
British Museum, 1845, p. 191.—Baiey, Proc. U. 8. Nat. Mus., vol. 73, 1928, 
pp. 9-16, pls. 1-4.—Smiru, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 137-139; 
Journ. Washington Acad. Sci., vol. 39, 1949, p. 36. 

Cyclura teres HARLAN, Journ. Acad. Nat. Sci. Philadelphia, vol. 4, 1825, pp. 242— 
251, pl. 26 (type lost; Tampico, Tamaulipas). 

Ctenosaura cycluriodes WIEGMANN, Isis von Oken, 1828, vol. 21, p. 371 (Zool. Mus. 
Berl. Nos. 576, 578, Mus. Comp. Zool. No. 2253, cotypes; Mexico, here 
restricted to Veracruz, Veracruz). 

Iguana (Ctenosaura) armata Gray, in Griffith, Animal Kingdom, by the Baron 
Cuvier, vol. 9, Synopsis . . . 1831, p. 38 (type lost; no type locality cited, 
here restricted to Tampico, Tamaulipas). 

Cyclura shawii Gray, loc. cit. (synonymic name for Iguana (Ctenosaura) acanthura 
Gray). 

Iguana (Ctenosaura) lanceolata Gray, loc. cit. (type lost; no type locality cited, 
here restricted to Tampico, Tamaulipas). 

Iguana (Ctenosaura) bellii Gray, loc. cit. (type lost; no type locality cited, here 
restricted to Tampico, Tamaulipas). 

Cyclura articulata W1neMANN, Herpetologia Mexicana, 1834, p. 43 (types unknown; 
type locality, Mexico). 

Cyclura denticulata WIEGMANN, ibid., pp. 43—44 (substitute name for Ctenosaura 
cycluroides Wiegmann). 

Cyclura semicristata FirziNcER, Systema reptilium, 1843, p. 56 (synonymic name 
for Cyclura denticulata Wiegmann).®8 

Ctenosaura multispinis Corr, Proc. Amer. Philos. Soc., vol. 238, 1886, p. 197 
(U.S.N.M. No. 72737; Captain Dallas collector; Dondominguillo, Oaxaca). 


Type.—Brit. Mus. Nat. Hist. No. XXII, 20-a (fide Bailey) ; collector 
unknown. 

Type locality —Not given by Shaw. Boulenger (Catalogue of the 
lizards in the British Museum, vol. 2, 1885, p. 196) gives ‘‘California,”’ 
which probably means Baja California. Since the species does not 
occur there we accept Bailey’s restriction to Tampico, Tamaulipas. 

Range.—Atlantic slopes from central Tamaulipas south to the 
Isthmus of Tehuantepec. Reliably reported only from the states of 
Tamaulipas, San Luis Potosi (Pujal), Veracruz, and Oaxaca.” 


‘7 Burt and Myers (Stanford Univ. Publ., biol. ser., vol. 8, 1942, p. 299) record the species, in error, from 
Acapulco, Guerrero. 

88 Fitzinger (Joc. cit.) attributes this name to Wiegmann, as do also Lichtenstein and Martens (Nomen- 
clator . . . Berolinensis, 1856, p. 8); we are, however, unable to locate Wiegmann’s use of the name and 
attribute it to Fitzinger, as did Bailey, Proc. U. 8. Nat. Mus., vol. 73, 1928, p. 10). 

89 Reexamination of specimens in the U. S. Nat. Mus. referred by Bailey (loc. cit.) to C. acanthura reveals 
that two other species were confused with C. acanthura. One specimen from Mirador, Veracruz, is a C. 
similis; and all from Uruapan, Michoac4n; Tehuantepec, Oaxaca (except Nos. 26341, 58498); Isabel Island; 
Tres Marfas Islands; Cuicatlan, Oaxaca; Tlapa and Balsas, Guerrero, are C. pectinata. That all other west- 
coast specimens in his list, from other museums, are also C. pectinata we believe is virtually a certainty. 


CHECKLIST OF REPTILES OF MEXICO 75 


CTENOSAURA PECTINATA (Wiegmann) 


Cyclura pectinata WIEGMANN, Herpetologia Mexicana, 1834, p. 42, pl. 2. 

Ctenosaura pectinata, Gray, Catalogue of the specimens of lizards in the collection 
of the British Museum, 1845, p. 191.—Barury, Proc. U.S. Nat. Mus., vol. 73, 
1928, pp. 24-27, pls. 7-11.—Smiru, Univ. Kansas Sci. Bull., vol. 22, 1935, 
pp. 134-137; Journ. Washington Acad. Sci., vol. 39, 1949, pp. 36, 37. 

Ctenosaura brevirostris Cope, Proc. Amer. Philos. Soc., vol. 23, 1886, pp. 266-268 
(type, U.S.N.M. No. 24709; John Xantus collector; Colima, Colima).— 
BAILEY, op. cit., pp. 27-29, pls. 12, 18, 15. 

Ctenosaura teres brachylopha Corer, Proc. Amer. Philos. Soc., voi. 28, 1886, p. 269 
(type, U.S.N.M. Nos. 7180-3; Bischoff collector; Mazatlan, Sinaloa).— 
Balitey, Proc. U. 8. Nat. Mus., vol. 73, art. 12, pp. 22-24, pl. 6. 

Ctenosaura parkeri Battey, Proc. U. S. Nat. Mus., vol. 73, art. 12, pp. 29-32, 
pls. 14, 15 (type, U.S.N.M. No. 18967; P. L. Jouy collector; Barranca 
Ibarra, Jalisco). 


Type—Mus. Zool. Berlin. No. 574; F. Deppe collector. 

Type locality Restricted to Colima, Colima. 

Range.—Pacific coast from southern Sinaloa southward to the 
Isthmus of Tehuantepec. Recorded from Isabel and the Tres Marias 
Islands and the states of Sinaloa, Durango (Ciudad, Ventanas), 
Nayarit, Jalisco, Colima, Michoacan, Morelos, Guerrero, Puebla, and 
Oaxaca. A record for the state of Guanajuato is unacceptable. 


CTENOSAURA HEMILOPHA (Cope) 


Cyclura (Ctenosaura) hemilopha Corn, Proc. Acad. Nat. Sci. Philadelphia, 1863, 
pp. 105-106. 

Ctenosaura hemilopha, Corn, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 312.— 
Van Densuren, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 39, 
66—71.—-Battey, Proc. U. 8S. Nat. Mus., vol. 73, 1928, pp. 17-22, pl. 5.— 
Smitu, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 140-142, pl. 23, fig. 1. 

Ctenosaura interrupta Bocourt, Le Naturaliste, vol. 2, 1882, p. 47 (Mus. Hist. 
Nat. Paris Nos. 2243, 2245, 2848, Brit. Mus. Nat. Hist. 1 spec., cotypes, 
M. Botta collector; ‘‘Baja California’, here restricted to Cape San Lucas). 

Ctenosaura conspicuose DICKERSON, Bull. Amer. Nat. Hist., vol. 41, 1919, p. 
461 (U.S.N.M. No. 64440; C. H. Townsend collector; San Esteban Island, 
Sonora). 

Ctenosaura insulana DicKERSON, op. cit., pp. 462-463 (U.S.N.M. No. 64439; 
C. H. Townsend collector; Cerralvo Islands, Baja California). 


Type—U.S.N.M. No. 529, four cotypes; John Xantus collector. 

Type locality —Cape San Lucas, Baja California. 

Range.—The southern two-thirds of Baja California, central 
Sonora (including San Pedro Nolasco Island) south to northern 
Sinaloa, and southwestern Chihuahua. Recorded only from the 
states cited. It is possible that the species extends as far north as 
southern Arizona. 

Genus ENYALIOSAURUS Gray 


Enyaliosaurus Gray, Catalogue of the specimens of lizards in the collection of 
the British Museum, 1845, p. 192. 
861316—50—6 


76 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Cachryz Corn, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 124 (genotype, Cteno- 
saura erythromelas Boulenger). 
Genotype.—Cyclura quinquecarinata Gray. 
Range.—Michoacan to the Isthmus of Tehuantepec, the Peninsula 
of Yucatan, and northern Guatemala. 
Species.—Five, all but one of which (C. palearis of Guatemala) 
occur in Mexico. 





KEY TO MEXICAN SPECIES OF ENYALIOSAURUS 


1. Whorls of enlarged caudal scales not interspersed with small scales. 

defensor (p. 77) 
Whorls of enlarged caudal scales interspersed with smaller scales_________ 2 

2. Dorsal and lateral caudal scales of nearly uniform character in each whorl. 
erythromelas (p. 77) 
Dorsal and lateral caudal scales not of nearly uniform character but median 
longitudinal series and outer lateral series differentiated______._______- 3 
3. Median row of dorsal scales extending to or almost to sacrum, noticeably 
raised, of medium height; tail twice length of body_quinquecarinata (p. 76) 
Median row of dorsal scales extending only two-thirds distance to sacrum 
and not noticeably raised; tail about 1% times body length__clarki (p. 76) 


ENYALIOSAURUS QUINQUECARINATUS (Gray) 


Cyclura quinquecarinata Gray, Zoological miscellany, 1842, p. 59. 

Enyaliosaurus quinquecarinatus, Gray, Catalogue of the specimens of lizards in 
the collections of the British Museum, 1845, p. 192. 

Cyclura (Ctenosaura) quinquecarinata, Corr, Proc. Amer. Philos. Soc., vol. 11, 
p61: 

Ctenosaura quinquecarinata, SumicuRast, Bull. Soc. Zool., vol. 10, 1880, p. 175.— 
Battey, Proc. U.S. Nat. Mus., vol. 73, art. 12, 1928, pp. 42-44, pls. 24, 25, 26. 


Type.—Brit. Mus. Nat. Hist. No. 61; collector unknown. 

Type locality—Unknown (restricted to Tehuantepec, Oaxaca, by 
Bailey, loc. cit.). 

Range.—Lowland regions of southern Oaxaca. Reported from 
Oaxaca: Tehuantepec, ‘““Hugma, Terminos,’”’ San Pedro, and Mixte- 
quilla Mountains, between Salina Cruz and Tequesixtlan. 


ENYALIOSAURUS CLARKI (Bailey) 


Enyaliosaurus quinquecarinatus, Ducts, La Naturaleza, ser. 2, vol. 2, 1897, pp. 
523-524, pl. 34 (color). 
Ctenosaura clarki Battrey, Proc. U.S. Nat. Mus., vol. 73, art. 12, 1928, pp. 44-46, 
pl. 27. 
Type-——Mus. Comp. Zool. No. 22454; Hans Gadow collector. 
Type locality —Ovopeo, Michoacan, elevation 1,000 feet. 
Range.—Known definitely only from the type locality; a record for 
Cuautla, Morelos, probably belongs here (Dugés, La Naturaleza, ser. 
2, vol. 2, 1896, p. 480). 


CHECKLIST OF REPTILES OF MEXICO CL 
ENYALIOSAURUS ERYTHROMELAS (Boulenger) 


Ctenosaura erythromelas BOULENGER, Proc. Zool. Soc. London, 1886, p. 241, pl. 
23 (color).—Bat.ey, Proc. U. 8. Nat. Mus., vol. 73, art. 12, 1928, pp. 46-48, 
pls. 28, 29.—Smirn, Occ. Pap. Univ. Michigan Mus. Zool., No. 388, 1938, p. 15. 

Cachryx erythromelas, Corr, U. 8. Nat. Mus. Bull. 32, 1887, p. 43. 

Ctenosaura (Cachryz) annectens WERNER, Jahrb. Hamburg Wiss. Anst., pt. 2, 
1911, p. 25 (locality unknown; P. Phol collector, Hamburg Museum; 
“Mexico’’). 

Type.—Brit. Mus. Nat. Hist. No. 1 (fide Bailey, loc. cit., p. 48); 
purchased alive. 

Type locality.—Unknown; here restricted to Balchacaj, Campeche. 

Range.—Known only from Balchacaj, Campeche. 


ENYALICSAURUS DEFENSOR Cope 


Cachryx defensor Corr, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 124; Proc. 
Amer. Philos. Soc., 1869, pl. 10.—Bocourt, Mission scientifique au Mexique 
. . ., Etudes sur les reptiles, livr. 3, 1874, pp. 148-149, pl. 17 bis, fig. 12, 12a. 
Ctenosaura defensor, GUNTHER, Biologia Centrali-Americana, Reptilia and Ba- 
trachia, 1890, pp. 58-59.—Baiey, Proc. U. 8. Nat. Mus., vol. 73, art. 12, 
1928, pp. 48-50, pl. 30. 
Type.—U. S. N. M. No. 12282; male, adult, and half grown, three 
cotypes; Arthur C. V. Schott collector. 
Type locality Yucatan, here restricted to Chichen Itza. 
Range.—Northern part of the Yucatan Peninsula. Reported 
only from Chichen Itz4, Yucatan. 


Genus DIPSOSAURUS Hallowell 


Dipso-saurus HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, vol. 7, 1854, p. 92. 
Genotype.—Crotaphytus dorsalis Baird and Girard. 
Range.—Sonora, Baja California, and some coastal islands of the 

Gulf. In United States: southern and central Arizona, Colorado and 


Mojave Deserts east to the Colorado River, southern Nevada, and 
Utah. 
Species.—Three, with five forms recognized. 


KEY TO MEXICAN SPECIES OF DIPSOSAURUS 


1. Two or more scale rows between nostril and rostral (62-83 percent); frequently 


with longitudinal dark lines on the body________-_ dorsalis dorsalis (p. 78) 
One scale row between nostril and rostral (75 to 80 percent); dark lateral lines 
usually tacking! 3.21 ett sent. Pope g mired oe ete rh eis get eet S 2 


2. Dorsal pattern consisting of large, round, white spots, and no longitudinal lines; 
brown reticulation on throat enclosing round or oval cream spots. 

dorsalis sonoriensis (p. 79) 

Dorsal pattern consisting of short longitudinal streaks, and few if any light 

spots, which are not dark-edged; throat either striped with dark longitudinal 

streaks or without streaks, in which case large rounded spots are present. 3 


78 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


3. Gular region without longitudinal dark streaks; large rounded spots on lower 
jaw and side of neck; central gular region suffused with dark brown; femoral 
DOTES) AVCLA CSL Oe gate oe a Lia ek ee ig csi SL eee catalinensis (p. 78) 

Gular region with longitudinal dark streaks, without definite rounded light 
spots laterally, sometimes suffused with dark centrally_____-_--------- 4 

4. Femoral pores fewer, average 18.46_-_-_--__-__--- dorsalis lucasensis (p. 78) 
Femoral pores more numerous, average 21.8__---------- carmenensis (p. 78) 


DIPSOSAURUS CARMENENSIS Van Denburgh 

Dipsosaurus carmenensis VAN DENBURGH, Occ. Pap. California Acad. Sci., No. 
10, vol. 1, 1922, pp. 81-82. 

Type.—California Acad. Sci. No. 50504; Joseph R. Slevin collector. 

Type locality —Carmen Island, Baja California. 

Range.—Carmen Island and Los Coronados Islands, Baja California. 

DIPSOSAURUS CATALINENSIS Van Denburgh 

Dipsosaurus catalinensis VAN DENBURGH, Occ. Pap. California Acad. Sci., No. 10, 
vol. 1, 1922, pp. 83-84. 

Type.—California Acad. Sci. No. 50505; Joseph R. Slevin collector. 

Type locality —Santa Catalina Island, Baja California. 

Range.—Known only from the type locality. 

DIPSOSAURUS DORSALIS LUCASENSIS Van Denburgh 

Dipsosaurus dorsalis lucasensis VAN DENBURGH, Proc. California Acad. Sci., ser. 4, 
vol. 10, 1920, p. 33; Occ. Pap. California Acad. Sci., No. 10, 1922, vol. 1, 
pp. 78-81. 

Type.—California Acad. Sci. No. 46090. 

Type locality —‘‘San José del Cabo, Baja California, Mexico.”’ 

Range.—Southern tip of Baja California, southward from Santa 
Margarita Island; Cerralbo Island. Reported from Miraflores, San 
José del Cabo, Santa Anita, Agua Caliente, Eureka, ete. 

DIPSOSAURUS DORSALIS DORSALIS (Baird and Girard) 

Crotaphytus dorsalis Barrp and Grrarp, Proc. Acad. Nat. Sci. Philadelphia, 
vol. 6, 1852, p. 126. 

Dipsosaurus dorsalis, HALLOWELL, tbid., vol. 7, 1854, p. 92. 

Dipsosaurus dorsalis dorsalis, VAN DENBURGH, Proc. California Acad. Sci., ser. 4, 
vol. 10, No. 4, 1920, pp. 33-34; Occ. Pap. California Acad. Sci., No. 10, vol. 1, 
1922, pp. 73-78, pl. 2. 

Type.—U.S.N.M. No. ?; John LeConte collector. 

Type locality.—“‘ Desert of Colorado, Cal.,”’ here restricted to Winter- 
haven (=Fort Yuma), Imperial County. 

Range.—Extreme southwestern Utah and eastern California south- 
ward into northwestern Sonora, and eastern Baja California south to 
Magdalena Island; recorded on the latter island and on those of San 
Luis, Angel de la Guarda, San Marcos, Monserrate, and San José. 
Reported from Baja California: Volcano Lake, San Felipe Bay, 13 
miles northwest of El Mejor, El Cajén Cafion, San Ignacio, 600 feet, 
etc.; Sonora: Punta Pefiasco. 


CHECKLIST OF REPTILES OF MEXICO 79 


DIPSOSAURUS DORSALIS SONORIENSIS Allen 
Dipso-saurus dorsalis sonoriensis ALLEN, Occ. Pap. Mus. Zool. Univ. Michigan, 
No. 259, 1933, pp. 4-6. 

Type.—Mus. Zool. Univ. Michigan No. 72121; John Piatt, John 
Scofield, and Morrow J. Allen collectors. 

Type locality Hermosillo, Sonora. 

Range.—Gulf slope of Sonora, except extreme northwestern part, 
and southward into northern Sinaloa. Reported from Sonora: 5 miles 
southwest of Hermosillo, Miramar, La Posa, Guaymas, Rancho Costa 
Rica, Bocoit; Sinaloa: Ahome. 


Genus SAUROMALUS Dumeéril 


Sauromalus Dum#érit, Arch. Mus. Hist. Nat. Paris, vol. 8, 1856, p. 5386.—Suaw, 
Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, pp. 269-306, map. 

Euphryne Bairp, Proc. Acad. Nat. Sci. Philadelphia, vol. 10, 1858, p. 253 (type, 
Euphryne obesus Baird). 


Genotype.—Sauromalus ater Duméril. 

Species.—Seven, one with three subspecies. 

Range.——Southern Utah and southern Nevada south through 
southern California, western Sonora and the Gulf slopes of Baja 


California. 
KEY TO SPECIES OF SAUROMALUS © 


1. One or more transverse bands dorsally, across body or rump____-------__- 4 
No transverse body, bands) present = === =--2 2-22-2222. ee eee eee 2 

2. Largest nuchal scales equally as large as or larger than frontal plates. 
hispidus (adult) (p. 80) 


Largest nuchal scales smaller than frontal plates____.__....__..__._.___---- 3 
3. Dorsal pattern of large, irregular, dark brown or black blotches on a yellowish 
PLOUNG COLON ssa aa ae ne a ee cra ee ee varius (p. 81) 


Dorsal pattern of small, dark brown or black spots on a gray ground color. 
klauberi (p. 80) 


4, Ventral scale rows usually, less; than W302... 22 3_- 2 ee 2s ee 5 
Wentraloscalesrows usually;more than W302 2225 622 ee ee ee 6 

5. Dorsal scales in a head length usually less than 20_hispidus (juvenile) (p. 80) 
Dorsal scales in a head length usually more than 20___________ slevini (p. 80) 

6. Transverse body bands with light centers and dark brown or black borders 
piv a Couble-pDandetremects see ne ere en oe se eS 9 
rancwerse bangs, Wnicolor+.5%) 205) 845 6508 8 re lee FE oes he cco A 

7. Scales around middle of upper foreleg (humeral scales) usually fewer than 
Eee en ee eS ee ee Ue ys Ri eee 8 
HormeraliscalesmoresjhanvoO sss a eee eee obesus obesus (p. 81) 


8. No reddish suffusion on dorsal and ventral areas in adult males; maximum 
adult length averaging somewhat less than in o. twmidus; average scale 
Countsylesser st). Bay. se i eee | = re obesus townsendi (p. 81) 

More or less brilliant reddish suffusion on dorsal and ventral areas in adult 
males; maximum adult length averaging more than in o. townsendi; average 


ROME COMMIS PROAtGr mila ee obesus tumidus (p. 81) 
OmVentral-scalesrows lol orsmores 404 2225 -) tee eee ee australis (p. 80) 
Ventral scale rows usually less than 151__-_---..__.._____--_--- ater (p. 80) 


60 From Shaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, p. 303. 


80 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


SAUROMALUS HISPIDUS Stejneger 


Sauromalus hispidus SresnEGER, Proc. U. S. Nat. Mus., vol. 14, 1891, p. 409.— 
Corr, Ann. Rep. U. S. Nat. Mus., 1898 (1900), p. 264, fig. 22—Van Den- 
BURGH, Occ. Pap. California Acad. Sci., No. 10, vol. 1, 1922, pp. 99-101, pls. 
5-6.—Suaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, pp. 279-280. 

Type.—U.S.N.M. No. 8563; Thomas H. Streets collector. 

Type locality. —Angel de la Guarda Island, Baja California. 

Range. —Angel de la Guarda, Smith, Pond, Nuevo Amor, Granite, 
Mejia, and South San Lorenzo Islands, Gulf of California, Baja 
California. 

SAUROMALUS KLAUBERI Shaw 


Sauromalus klaubert SHaw, Trans. San Diego Soc. Nat. Hist., vol. 9, 1941, p. 285; 
vol. 10, 1945, pp. 282-283. 
Type.—L. M. Klauber No. 6859. 
Type locality.—Santa Catalina Island, Baja California. 
Range.—The type locality. 


SAUROMALUS SLEVINI Van Denbargh 


Sauromalus slevini VAN DEeNnBURGH, Occ. Pap. California Acad. Sci., No. 10, 
vol. 1, 1922, pp. 97-99.—SuHaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 
1945, pp. 280-282. 

Type.—California Acad. Sci. No. 50503, adult male; Joseph R. 

Slevin collector. 

Type locality.—South end of Monserrate Island, Baja California. 
Range.—Monserrate, Carmen, and Coronados Islands, Baja Cali- 


fornia. 
SAUROMALUS AUSTRALIS Shaw 


Sauromalus australis SHAw, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, 
pp. 286-288. 
Type.—L. M. Klauber No. 30170; Robert S. Hoard collector. 
Type locality.—San Francisquito Bay, Baja California. 
Range.—The mainland of central and southern Baja California on 


the Gulf coast. 
SAUROMALUS ATER Duméril 


Sauromalus ater Dumérin, Arch. Mus. Hist. Nat. Paris, vol. 8, 1856, p. 536, 
pl. 23, figs. $, 3a.—Suaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, 
pp. 284-286. 
Type.—Mus. Hist. Nat. Paris; M. Jaurés collector? (“qui a été 
donné .. . par M. Jaurés, lieutenant 4 bord de la frégate la Danaide’’). 
Type locality.x—Unknown; presumably one of the islands in the 
southern part of the Gulf of California, Baja California, here restricted 
to Espiritu Santo Island. 
Range.—Espiritu Santo, Isla Partida, San Francisco, San Diego, 
Santa Cruz, and San Marcos Islands, Gulf of California, Baja Cali- 
fornia. 


CHECKLIST OF REPTILES OF MEXICO 81 


SAUROMALUS VARIUS Dickerson 


Sauromalus varius DickERsoN, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, pp. 
464-465.—Suaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, pp. 288-290. 
Type.—U.S.N.M. No. 64441; C. H. Townsend collector. 
Type locality —San Esteban Island, Gulf of California. 
Range.—The type locality. 


SAUROMALUS OBESUS TOWNSENDI Dickerson 


Sauromalus townsendi Dickrerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, 
p. 464. 
Sauromalus obesus townsendi, SHaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 
1945, pp. 290-292. 
Type.—U.S.N.M. No. 64442; C. H. Townsend collector. 
Type locality —Tiburén Island, Gulf of California. 
Range.—Tiburén Island, Sonora, and adjacent coast of Sonora at 
least as far south as Guaymas and east to the vicinity of Hermosillo. 


SAUROMALUS OBESUS TUMIDUS Shaw 


Sauromalus obesus tumidus Suaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 
1945, No. 15, pp. 269-306. 

Type.—L. M. Klauber No. 27323; L. M. Klauber collector. 

Type locality—Telegraph Pass, Gila Mountains, Yuma County, 
Ariz. 

Range.—Extreme northwestern Sonora, southwestern and south 
central Arizona. Intergrades only, with S. 0. townsendi, are recorded 
from Mexico; typical specimens undoubtedly occur in extreme north- 
western Sonora and extreme northeastern Baja California, along the 
Colorado River. 


SAUROMALUS OBESUS OBESUS (Baird) 


Euphryne obesus Batrp, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 253; United 
States and Mexican boundary survey, vol. 2, Rept., 1859, p. 6, pl. 27. 
Sauromalus obesus obesus, SHaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, 

pp. 295-301. 

Type.—U.S.N.M. No. 4172; G. H. Thomas collector (?). 

Type locality —Fort Yuma, Calif. 

Range.—Southeastern California, southern Nevada, southern Utah, 
western Arizona, and extreme northern Baja California. Although 
known from numerous localities within a few hundred feet of the 
Baja California line, none have actually been taken in that state. 


Genus HOLBROOKIA Girard 


Holbrookia Girarp, Proc. Amer. Assoc. Adv. Sci., vol. 4, 1851, p. 201. 
Cophosaurus TroscuHEL, Arch. fiir Naturg., 1850 (1852), p. 389 (type, Cophosaurus 
texana Troschel). 


82 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Genotype.—Holbrookia maculata Girard. 

Range.—Northern states of Mexico south to San Luis Potosi and 
Durango. In the United States: Texas, New Mexico, and Arizona. 

Species.—Your, with twelve forms, of which ten are known to 
occur or are to be expected in Mexico. 


KEY TO MEXICAN SPECIES OF HOLBROOKIA 


1. Tail flat, with broad, black, ventral bands; lateroventral black marks placed 
far posteriorly, continued dorsally above lateral fold, and slanting ante- 
roventrally cone) ly aaa ase a es ee texana (p. 85) 

Tail rounded; no ventral subcaudal marks save small spots in one subspecies; 
lateroventral marks placed farther anteriorly, not extending upon dorsal 
surface, and slanting posteroventrally on belly.___......------------ 2 

2. Several black spots under tail; dorsal pattern of large, well-defined spots 

equally sharp-edged on all sides; no light dots in pattern. 
maculata lacerata (p. 83) 


No black spots under tail; pattern not as described___.__....-._---.-_- 3 

3. Dorsal body scales distinctly keeled, rather small__._....__._--------- 11 
Dorsal body scales not keeled (although sometimes pointed and somewhat 
convex), larger: dat Cet Bae: <BR E 22h ARE Oe eee ee 4 

4. Tail generally shorter than the snout-vent measurement, usually not over 
51 percent of total length (the higher proportions are of males) __.--_-- 5 

Tail generally as long as or longer than snout-vent measurement, 50-58 
percent of total length (lower proportions are of females) _..._.____-_- 7 

5. Enlarged supraoculars in contact with frontals____maculata elegans (p. 84) 


Enlarged supraoculars separated from frontals by a series of small scales. 6 

6. Size greater, maximum 71 mm., average 59 mm. snout to vent; in males a 
large blue patch surrounding lateroventral black spots on belly. 

maculata dickersonae (p. 83) 

Maximum size 61 mm., average 50 mm. snout to vent; in males a smaller 

blue patch, or none, partially surrounding lateroventral black spots on 


ey eae Ma mecca etn ee maculata approximans (p. 83) 
(emHemoralaporessUsuallyalesseu marae! 2 eee meee tee eee Spey ee ny Ean eee 8 
Hemoraliporesusuall yal Zion more! ase eae a eee ee eee ee eee 10 

8. Upper labial region strongly flared, almost flat..._______- bunkeri (p. 84) 
Upper labial regioniless Haredsobtusess- eee eee eae ee eee eee 9 


9. Scales on limbs distinctly keeled; upper labials strongly imbricate. 

maculata pulchra (p. 84) 

Seales on limbs smooth; upper labials less strongly imbricate. 
maculata dickersonae (p. 83) 
10. Femoral pores about two-thirds as wide as pore scales are long in adult males, 
about one-half in adult females________-._---- maculata elegans (p. 84) 
Femoral pores about one-half as wide as the pore scales are long, in adult 
males, about one-fourth in adult females__._. maculata thermophila (p. 84) 
11. Anterior gulars a transverse row of 4 subequal scales, the inner pair not 
separated by linear or rounded scales__-_-_---- propinqua piperata (p. 85) 
INotisos.mwet) C2820. (ee Nite tine nee wilt propinqua propinqua (p. 85) 


CHECKLIST OF REPTILES OF MEXICO 83 


HOLBROOKIA MACULATA APPROXIMANS Baird 


Holbrookia approximans Bairp," Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 253. 
Holbrookia maculata approximans, STEJNEGER, North Amer. Fauna, No. 3, 1890, 
p. 109.—Scumipt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 721-723, 
pl. 60.—Smirs, Handbook of lizards, 1946, pp. 119-122, pl. 17. 
Holbrookia maculata flavilenta Corr, Proc. Acad. Nat. Sci. Philadelphia, 1883, 
p. 10 (Lake Valley, Dona Ana County, N. Mex.; Acad. Nat. Sci. Philadelphia, 
2 cotypes; E. D. Cope collector). 
Type.—Lost. 
Type locality.x—Lower Rio Grande of Tamaulipas (fide Baird, 
United States-Mexican boundary survey, Reptiles, 1859, p. 8). 
Range.—Western Texas to eastern Arizona, and from southeastern 
Utah southward through Chihuahua and central Coahuila, and 
southward in the west to Jalisco. Reported from Chihuahua: 35 and 
40 miles south of Moctezuma, between Churo and Chiricahui, Ojos 
del Diablo, Rio Santa Maria, Ramos, Casas Grandes; Coahuila: ? 
Concordia, Sierra de Santa Rosa, Buena Vista, Hda. La Rosita; San 
Imis Potosi: 5 miles northeast of Candido Navarro; Sonora: headwaters 
of San Pedro; Durango; El Oro; Jalisco: Colotlan.” 


HOLBROGCKIA MACULATA DICKERSONAE Schmidt 


Holbrookia dickersonae Scumipt, Amer. Mus. Nov., No. 22, 1921, p. 2; Bull. 
Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 724-725.—Smirn, Univ. Kansas 
Sci. Bull., vol. 22, 1935, pp. 188-189. 

Holbrookia maculata dickersonae, SmirH and Mirriteman, Trans. Kansas Acad. 
Sci., vol. 46, 1943, pp. 2438-244. 


Type.—U.S.N.M. No. 2668A; Lt. D. N. Couch collector. 

Type locality —Castafiuelas, Coahuila. 

Range.—The northern portion of the central plateau, as far south 
as Guanajuato, and adjacent areas immediately northward in southern 
Coahuila and Durango. Reported from Coahuila: Castafuelas, 
Hipélito, Jaral, Agua Nueva, Jimulco, Saltillo, Alamo de Parras, 10 
miles east of Torreédn; Durango: near Avilco, 5 miles north of Conejos, 
25 miles north of Bermejillo, Gémez Palacio; San Luis Potosi: Jestis 
Maria; Zacatecas: La Colorada, Berriozibal; Guanajuato: San Felipe. 


HGLBROOKIA MACULATA LACERATA Cope 


Holbrookia lacerata Corr, U. 8. Nat. Mus. Bull. 17, 1880, p. 15.—Scumint, Bull. 
Amer. Mus. Nat. Hist., vol. 46, 1922, p. 718. 

Holbrookia maculata lacerata, StEJNEGER, North Amer. Fauna, No. 3, 1890, 
p. 109.—Smitru, Handhook of lizards, 1946, pp. 122-124, pl. 18. 





61 The type locality for this name strongly indicates that the species involved is H. propingua, since that is 
the only one of this group whose range reasonably may include the state of Tamaulipas. Nevertheless, 
since the description better fits H. maculata, and since Baird presumably well knew H. propingua (having 
described it also), we retain the previous allocation of the name H. approzimans pending further investiga- 
tions. 

62 A number of records for ‘““Tamaulipas,’ without specific locality, are regarded erroneous either by 
misidentification or incorrect locality data. 


84 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type.—U.S.N.M. No. 10160. 

Type locality —Erath County, Tex., west of the Upper Brazos; on 
the Guadalupe River in Kendall or Comal County. Here restricted 
to Erath County, Tex. 

Range.—Central Texas plains region south to northern Coahuila. 
Recorded in Mexico only from Coahuila: Sabinas. 


HOLBROOKIA MACULATA PULCHRA Schmidt 


Holbrookia pulchra Scumipt, Amer. Mus. Nov., No. 22, 1921, p. 1. 
Holbrookia maculata pulchra, Sm1rH, Handbook of lizards, 1946, pp. 124-126, pl. 19. 


Type.—Amer. Mus. Nat. Hist. No. 14777; R. D. Camp collector. 
Type locality —Carr Canyon, Huachuca Mountains, Ariz., 5,200 feet. 
Range.—Known from the general region of the type locality; to be 
expected in adjacent Chihuahua and Sonora. 
HOLBROOKIA MACULATA THERMOPHILA Barbour 
Holbrookia thermophila Barsour, Proc. New England Zool. Club, vol. 7, 1921, 
p. 79. 
Holbrookia elegans thermophila, Smiru, Univ. Kansas Sci. Bull., vol. 22, 1935, 
pp. 194-195, pls. 27, fig. 1, pl. 28, fig. 4. 


Holbrookia maculata thermophila, Smitu, Handbook of lizards, 1946, pp. 129-132, 
pl. 21. 


Type-——Mus. Comp. Zool. No. 14281; W. W. Brown and J. E. 
Thayer collectors. 

Type locality —San José de Guaymas, Sonora. 

Range.—Sonora; possibly in northern Sinaloa. Reported from 
Sonora: 8 miles south of Nogales, 53 miles south of Nogales, 8 miles 
south of Magdalena, Canoa, near Noria, 30 miles south of Noria, 5 
miles southwest of Hermosillo, 54 miles south of Hermosillo, 10 miles 
north of Guaymas, Batamotal, Alamos, Guirocoba. ?Sinaloa: ?Bacu- 
birito. 

HOLBROOKIA MACULATA ELEGANS Bocourt 
Holbrookia Bischoffi Corr, Proc. Acad. Nat. Sci. Philadelphia, 1868, p. 310 
(nomen nudum). " 
Holbrookia elegans Bocourt, Mission scientifique au Mexique . . . , Etudes sur 
les reptiles, livr. 3, 1874, pp. 164-165, pl. 17 bis, figs. 8, 8a. 
Holbrookia elegans elegans, SmirH, Univ. Kansas Sci. Bull., vol. 22, No. 8, 1935, 
pp. 191-194. 

Type-—Mus. Hist. Nat. Paris. 

Type locality — Mazatlan, Sinaloa. 

Range.—Sinaloa. Reported from Presidio de Mazatlan, Mazatlan, 
Escuinapa, ?Bacubirito, Rosario. 


HOLBROOKIA BUNKERI Smith 


Holbrookia bunkert SmirH, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 185-188, 
pl. 28, fig. 3. 


CHECKLIST OF REPTILES OF MEXICO 85 


Type.—Univ. Kansas Mus. Nat. Hist. No. 19236; David H. Dunkle 
and Hobart M. Smith collectors. 

Type locality.—15 miles south of Judrez, Chihuahua. 

Range.——Known only from the type locality. 


HOLBROOKIA PROPINQUA PROPINQUA Baird and Girard 


Holbrookia propinqua Barrp and Grrarp, Proc. Acad. Nat. Sci. Philadelphia, 
vol. 6, 1852, p. 126.—Scumipt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, 
p. 714.—Smitu, Handbook of lizards, 1946, pp. 132-134, pl. 22. 

Holbrookia propinqua propinqua SmitH and Burcesr, Trans. Kansas Acad. Sci., 
vol. 53, 1950, p. 169. 


Type.—Unknown. 

Type locality—-Between Indianola and San Antonio, Tex., here 
restricted to 9 miles southwest of Somerset, Atascosa County. 

Range.—Central and southern Texas, southward presumably into 
Tamaulipas. No definite record for Mexico is known. 


HOLBROOKIA PROPINQUA PIPERATA Smith and Burger 


Holbrookia propinqua piperata SmirH and Buraer, Trans. Kansas Acad. Sci., 
vol. 53, 1950, pp. 167-169. 


Type.—Univ. Illinois Mus. Nat. Hist. No. 4048; R. W. Reese and 
P. W. Smith collectors. 

Type locality— Beach at Etiopa, 2 miles south of Tecolutla, Veracruz. 

Range.—Presumably southern Tamaulipas and northern Veracruz; 
recorded only from the type locality. 


HOLBROOKIA TEXANA (Troschel) 


Cophosaurus texanus TROSCHEL, Wiegmann’s Arch. fiir Naturg., Jahrg. 16, vol. 
1, 1850 (1852), p. 289, pl. 6. 

Holbrookia texana, Batrp and Grrarp, Proc. Acad. Nat. Sci. Philadelphia, vol. 
6, 1852, p. 124.—Cornr, Ann. Rep. U. S. Nat. Mus., 1898 (1900), pp. 286— 
289, fig. 30.—SmirH, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 190-191 
(very numerous localities listed). 

Holbrookia affinis Barrp and Grrarp, Proc. Acad. Nat. Sci. Philadelphia, vol. 
6, 1852 (types unknown; San Pedro, Tex.). 


Type.—Not known. 

Type locality—New Braunfels, on the Guadalupe River, Tex.; 
latitude 28° N. (=New Braunfels, Comal County, Tex.). 

Range.—Southeastern Arizona to central Texas and southward 
through the northern parts of all the border states in Mexico except 
Baja California. Reported from Tamaulipas, San Luis Potosi, 
Nuevo Leén, Coahuila, Chihuahua, Durango, and Sonora. 


Genus CALLISAURUS Blainville 


Callisaurus BLAINVILLE, Nouv. Ann. Mus. Hist. Nat. Paris, vol. 4, 1835, p. 286. 
Megadactylus FirzinarerR, Systema reptilium, 1848, p. 59 (type, Callisaurus dra- 
conoides Blainville). 


86 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Homalosaurus HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, 1852, p. 179 
(type, Homalosaurus ventralis Hallowell). 


Genotype.—Callisaurus draconoides Blainville. 

Range.—Southwestern United States, southward from Nevada to 
Baja California, Sonora, and adjacent islands. 

Species.—One species with 10 forms; only one (C. d. myurus Rich- 
ardson of Nevada) is extralimital. 


KEY TO MEXICAN SPECIES OF CALLISAURUS 


1. A fringe of small spinose scales along one side of toes somewhat as in Uma. 

draconoides crinitus (p. 87) 

No‘such ‘fringe... eo 2 as tS es De ee 2 

2. Males with two nearly vertical lateral black blotches usually followed by a 

third black spot; dorsal caudal bars never black, usually with strongly 

undulate or pointed posterior margins______ draconoides draconoides (p. 86) 

Males with two large oblique lateral black blotches usually not followed by a 

third black spot; dorsal bars black and undulate or not______.-___---- 3 

3. Dark dorsal bars on tail with strongly undulate or pointed posterior margins 
from base to beyond middle of tail; dorsal tail bars never black. 

draconoides carmenensis (p. 87) 

Dark dorsal bars on tail with nearly straight or slightly undulate posterior 

margins at least from middle to end of tail; dorsal tail bars often black__ 4 

4. Dorsal tail bars black in both sexes; usually more or less red in dorsal coloration. 

draconoides splendidus (p. 87) 

Dorsal tail bars black in males only; usually little ornoredin dorsal coloration. 5 

5. Lateroventral black bars in males ill defined, blurred or obsolete, usually united 


SVG yy eee ois ie oder ne alas, Se ee ee 6 
Lateroventral black bars in males well defined, intense, separate below__-_ 8 

6. Lamellae under fourth toe 25-28______________ draconoides bogerti (p. 89) 
Eamellac under fourth toe 80 or more...) ee ee eee ee 7 

7. Lamellae under fourth toe 30 to 33_____-___-___ draconoides brevipes (p. 88) 
Lamellae under fourth toe 35-39____________ draconoides inusitatus (p. 88) 


8. Femoral pores generally 16 or fewer; hind leg generally 92 percent of body 
length, or more; tail generally 58 percent of total length, or more; inter- 
parietal and supraorbital semicircles separated completely. 

draconoides gabbii (p. 87) 
Femoral pores generally 17 or more; hind leg generally 91 percent of body 
length, or less; tail generally 56 percent of total length, or less; interparietal 
and supraorbital semicircles usually not separated completely. 
draconoides ventralis (p. 88) 
CALLISAURUS DRACONOIDES DRACONOIDES Bilainville 

Callisaurus draconoides BLAINVILLE, Nouv. Ann. Mus. Hist. Nat. Paris, vol. 4, 
1835, p. 286, pl. 24, fig. 2, 2a—Scumipt, Bull. Amer. Mus. Nat. Hist., 
vol. 46, 1922, pp. 646-647, fig. 5, pl. 51, figs. 1-3. 

Callisaurus draconoides draconoides, Corr, Ann. Rept. U. S. Nat. Mus., 1898 
(1900), p. 272, fig. 24-—Van Densurau, Occ. Pap. California Acad. Sci., 
No. 10, 1922, pp. 142-145. 

Type.—Mus. Hist. Nat. Paris; M. Botta collector. 
Type locality—“California,”’ here restricted to Cape San Lucas, 

Baja California. 


CHECKLIST OF REPTILES OF MEXICO 87 


Range.—Cape region of Baja California; Isla Espiritu Santo. 
CALLISAURUS DRACONOIDES CARMENENSIS Dickerson 


Callisaurus carmenensis Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, 
AG5- 

Eas draconoides carmenensis, VAN DrnBuURGH, Occ. Pap. California 
Acad. Sci., No. 10 (vol. 1), 1922, pp. 145-148.—Linspa.z, Univ. California 
Publ. Zool., vol. 38, 1932, p. 358. 

Callisaurus plasticus DicKERSON, tbid., p. 466 (Agua Verde Bay, Baja California, 
Amer. Mus. Nat. Hist. No. 5349; C. H. Townsend collector). 

Type.—Amer. Mus. Nat. Hist. No. 5388, male; C. H. Townsend 
collector. 

Type locality —Carmen Island, Gulf of California, Baja California. 

Range.—The southern two-thirds of Baja California, except the 

Pacific coast in the center of that area, and the Cape region, between 

lat. 29°30’ N. and 24° N.; the islands of San José, Carmen, San Mar- 

cos, South Santa Inés, Coronados, and Santa Margarita. 


CALLISAURUS DRACONOIDES SPLENDIDUS Dickerson 


Callisaurus splendidus Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, 
p. 467.—ScumiptT, zbid., vol. 46, 1922, pp. 647-648, pl. 51, fig. 1, and text fig. 
6.—Van DeEnBorRGH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 148- 
150. 


Type—Amer. Mus. Nat. Hist. No. 5372, male; C. H. Townsend 
collector. 

Type locality—Isla Angel de la Guarda, Gulf of California, Baja 
California. 

Range.—Island of Angel de la Guarda, Baja California. 


CALLISAURUS DRACONOIDES CRINITUS Cope 


Callisaurus crinitus Corr, Amer. Nat., vol. 30, 1896, p. 1049.—Van Drnsurau, 
Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 140-142.—Scumipt, Bull. 
Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 645-646, pls. 51-52, text fig. 4. 

Callisaurus draconoides crinitus, Trevis, Copeia, 1944, pp. 7-12. 

Callisaurus rhodostictus Copx, ibid., vol. 30, 1896, pp. 1049-1050 (‘‘Lower Cali- 
fornia,” here restricted to Santo Domingo; A. W. Anthony coll.; U. 8S. Nat. 
Mus.). 


Type —U.S. N. M. No. 14895. 

Type locality—‘Lower California” [=Ballenas Bay, Baja Cali- 
fornia]. 

Range.—Along the Pacific shore of central Baja California from 
Millers Landing (lat. 28°30’ N.) to Santo Domingo (lat. 25°30’ N.). 


CALLISAURUS DRACONOIDES GABBII Cope 


Callisaurus ventralis gabbit Corn, Ann. Rep. U.S. Nat. Mus., 1898 (1900), p. 272.— 
Scumipt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 648-651, pl. 51, 
fig. 1 (part). 


88 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Callisaurus draconoides gabbii, LINSDALE, Univ. California Publ. Zool., vol. 38, 
1932, pp. 358-359; Proc. Amer. Acad. Arts and Sci., vol. 73, 1940, pp. 220—- 
221.—Trvis, Copeia, 1944, pp. 7-12, map fig. 2—Smiru, Handbook of 
lizards, 1946, pp. 138-145, pl. 24. 

Type.—Apparently none designated. 

Type locality——‘‘Northern Lower California,” here restricted to 
Gardners Laguna, Salton River. 

Range.—Southern Nevada, western Arizona, and southeastern 
California southward into extreme northwestern Sonora and north- 
eastern Baja California. Reported from several localities in Baja 
California, and from Sonora: Colorado River, Shoal Point. 


- 


CALLISAURUS DRACONOIDES VENTRALIS (Hallowell) 


Homalosaurus ventralis HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, vol. 6, 
1852, p. 179. 

Callisaurus ventralis, Barrp, United States and Mexican Boundary Survey, vol. 2, 
pt. 2, 1859, Rept., p. 8. 

Callisaurus ventralis ventralis, Camp, Univ. California Publs. Zool., vol. 12, 1916, 
pp. 508, 519.—Van Densouraa, Occ. Pap. California Acad. Sci., No. 10, 1922, 
pp. 152-163, pl. 12 (part).—Scumipr, Bull. Amer. Mus. Nat. Hist., vol. 46, 
1922, p. 651. 

Callisaurus draconoides ventralis, LINSDALE, Univ. California Publ. Zool., vol. 38, 
1932, p. 359.—Smiru, Handbook of lizards, 1946, pp. 146-147, pl. 26. 


Type.—U.S. N. M. No. 2670; S. W. Woodhouse collector. 

Type locality —New Mexico, here restricted to Tucson, Ariz. 

Range.—Southern Arizona except the western and eastern edges, 
southward into northern Sonora. Recorded in Mexico only from the 
latter state, from between Sonoyta and Puerto Pefiasco. 


CALLISAURUS DRACONOIDES INUSITATUS Dickerson 


Callisaurus inusitatus DickErson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, 
pp. 465-466.—Van Denpuren, Occ. Pap. California Acad. Sci., No. 10, 
1922, pp. 150-152. 

Callisaurus ventralis inusitatus, Scumipt, Bull. Amer. Mus. Nat. Hist., vol. 46, 
1922, p. 648, pl. 51, fig. 1, pl. 52, fig. 2. 


Type.—Amer. Mus. Nat. Hist. No. 5324, male; C. H. Townsend 
collector. 

Type locality —Tibur6én Island, Sonora. 

Range.—Tiburén Island, and adjacent Sonoran coast to Guaymas. 





CALLISAURUS DRACONOIDES BREVIPES Bogert and Dorson 


Callisaurus draconoides brevipes Bogert and Dorson, Copeia, 1942, No. 3, 
pp. 173-175.—Bocert and Ouiver, Bull. Amer. Mus. Nat. Hist., vol. 83, 
1945, pp. 314, 346-347, 401, 412, 418, 419. 

Type.—Amer. Mus. Nat. Hist. No. 63668; J. W. Hilton collector. 
Type locality —Guirocoba, 18 miles southeast of Alamos, Sonora, 
Mexico, elevation approximately 1,485 feet. 


CHECKLIST OF REPTILES OF MEXICO 89 


Range.—Extreme southern Sonora (Alamos and Guirocoba) and 
northern Sinaloa (Rio de Choix). 


CALLISAURUS DRACONOIDES BOGERTI Martin del Campo 


Callisaurus draconoides bogertti Martin del Campo, Anal. Inst. Biol. (Mexico), 
vol. 15, 1943, pp. 619-621. 
Type.—Inst. Biol. México; Victoriano Rodriguez collector. 
Type locality—Isla de los Chivos, en el puerto de Mazatlan, 
Sinaloa. 
Range.—Known only from the type locality and the vicinity of 


Mazatlan. 
Genus UMA Baird 


Uma Barrp, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 253.—He1Frerz, Copeia, 
1941, pp. 99-111, figs. 1-7. 
Genotype.—Uma notata Baird. 
Species.—Four, one of which includes two subspecies. Three forms 
occur in Mexico. 
Range.—Southeastern California and adjacent Arizona, Sonora 
and Baja California, and southwestern Coahuila. 


KEY TO MEXICAN FORMS OF UMA 


1. Two strongly enlarged auricular lobules; a black bar on rear of femur; dorsal 
pattern with 3 median rows of spots and black chevrons with reticulum out- 
Lining awhitistiidotsor blotChestese= =a == ee eee exsul (p. 89) 

Four strongly enlarged auricular lobules; no black bar on femur_-____---_-- 2 

2. Ventrolateral blotch smaller; femoral pores 20-31, usually fewer than 26. 

notata notata (p. 89) 
Ventrolateral blotch larger; femoral pores 23-32, usually more than 26. 


notata cowlesi (p. 90) 
UMA EXSUL Schmidt and Bogert 


Uma exsul Scumipt and Bocert, Amer. Mus. Nov., No. 1339, 1947, pp. 1-9, 
figs. 1-4. 

Type.—Amer. Mus. Nat. Hist. No. 67404; Karl P. Schmidt and 
C. M. Bogert collectors. 

Type locality —Dunes 12 miles north of San Pedro de Las Colonias, 
Coahuila. 

Range.—Presumably restricted to the vicinity of the type locality, 
the northern edge of the dry Laguna de Mayran. 


UMA NOTATA NOTATA Baird 


Uma notata Bairp, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 253. 
Uma notata notata, He1retTz, Copeia, 1941, pp. 101-104, figs. 1c, 2c, 3. 
Uma rufopunctata Corr, Amer. Nat., vol. 29, 1895, p. 939 (U.S.N.M. Nos. 21846-52, 
7 cotypes; ‘‘Yuma Desert,’’ Ariz.). 
Type.—U.S.N.M. No. 4124; Williamson and Heerman collectors. 
Type locality —‘‘Mojave Desert,” in error; restricted by Heifetz 


90 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


to the ‘Colorado Desert” and here restricted to the vicinity of Yuma, 
Ariz. 

Range.—Extreme southeastern Baja California, southwestern 
Arizona, and northeastern Baja California; possibly occurs in Sonora. 
Reported in Mexico only from Baja California. 


UMA NOTATA COWLESI Heifetz 


Uma notata cowlest HetretTz, Copeia, 1941, pp. 104-106, fig. 5 (distribution). 


Type.—California Acad. Sci. No. 53370; J. R. Slevin collector. 
Type locality.—Shores of Tepoca Bay, Sonora. 
Range.—The coast of northern Sonora, from Cerro Prieto to 
Tepoca Bay. 
Genus PETROSAURUS Boulenger 


Petrosaurus BOULENGER, Catalogue of the lizards in the British Museum, ed. 2, 
1885, vol. 2, p. 205.—MurtLteman, Bull. Mus. Comp. Zool., vol. 91, 1942, 
pp. 110-111. 


Genotype. —Uta thalassina Cope. 
Range.—Southern Baja California and certain adjacent islands in 
the Gulf of California. 


Species.—Two. 
KEY TO SPECIES OF PETROSAURUS 


1. Two series of scales between subocular and labials; snout longer, less truncate; 
three distinct anterior dorsal cross bars; throat yellow, orange, or pale blue 

CON Grad yf hohe a eye et Fe a Ree ch el eh oo thalassina (p. 90) 

One series of scales between subocular and labials; snout shorter and more 
truncate; four distinct anterior dark bands; throat brownish marked with 
blackishrcentralliys 2-2. ees ee a a ee a ee repens (p. 90) 


PETROSAURUS REPENS (Van Denburgh) 


Uta repens VAN DeNnBURGH, Proc. California Acad. Sci., ser. 2, vol. 5, 1895, p. 
102, pls. 7, 8, figs. a-e; Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 189- 
191.—Linspa.z, Univ. California Publ. Zool., vol. 38, 1932, pp. 360-361. 

Petrosaurus repens, MittLeEMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, p. 111. 

Type.—California Acad. Sci. No. 633; W. E. Bryant collector. 
Type locality.—Comondt, Baja California. 
Range.—The southern half of Baja California, exclusive of the 

Cape region; Isla Danzante. 


PETROSAURUS THALASSINUS (Cope) 


Uta thalassina Corn, Proc. Acad. Nat. Sci. Philadelphia, 1863, p. 104.—Van 
DensureaH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 186-189. 
Petrosaurus thalassinus, BouLENGER, Catalogue of the lizards in the British 

Museum, ed. 2, vol. 2, 1885, p. 205. 


Type.—U.S.N.M. No. 5302; John Xantus collector. 
Type locality. —Cape San Lucas, Baja California. 


CHECKLIST OF REPTILES OF MEXICO 91 


Range.—Cape region of Baja California and certain adjacent 
islands. Reported on the mainland from Cape San Lucas, La Paz, 
San José del Cabo, Miraflores, etc.; also Isla Partida, Isla Espiritu 
Santo. 


Genus STREPTOSAURUS Mittleman 
Streptosaurus MirrTLEMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, pp.111-117. 


Genotype.—Uta mearnsi Stejneger. 

Range.—Northern portion of Baja California and certain of the 
islands in the Gulf of California; in the United States: San Diego, 
Riverside, and Imperial Counties, Calif. 

Species. —T wo. 

KEY TO FORMS OF STREPTOSAURUS 


1; Throat with numerous round light spots; maximum length about 90 mm. 
mearnsi (p. 91) 
Throat mottled, without definite rounded spots; larger maximum length, 
SOO Ub ge) 4m rn eee ee eee se ee nore leas slevini ® (p. 91) 


STREPTOSAURUS MEARNSI (Siejneger) 


Uta mearnsi STEJNEGER, Proc. U. S. Nat. Mus., vol. 17, 1894, pp. 589-591.— Van 
DENnBURGH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 191-194, pl. 15. 
[Streptosaurus] mearnsi, MirTLEMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, p. 
111.—STEJNEGER and BarBour, Check list of North American amphibians 
and reptiles, ed. 5, 1943, p. 90. 
Type—U.S.N.M. No. 21882; E. A. Mearns collector. 
Type locality—‘Summit of Coast Range, United States and Mexico 
boundary line, California.” 
Range. —Extreme southern California southward through the 
northern half of Baja California. Reported from Baja Califorma: 
Ensenada, San Salado, San Matias, Parral, Matomi, Coyote Bay. 


STREPTOSAURUS SLEVINI (Van Denburgh) 


Uta slevini VAN DENBURGH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 
194-196. 

Streptosaurus slevint, MirTLEMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, p. 112. 
Type.—California Acad. Sci. No. 50506; Joseph R. Slevin collector. 
Type locality.—Mejia Island, Gulf of California. 

Range.—Mejia Island and Angel de la Guarda Island, in the Gulf of 

California. 

Genus CROTAPHYTUS Holbrook 


Crotaphytus HotBroox, North American herpetology, ed. 2, vol. 2, 1842, p. 79. 
Genotype.—Agama collaris Say. 


63 The senior author has unintentionally referred Petrosaurus repens to this genus, owing to ‘a lapsus; 
slevini was intended (Smith, Handbook of lizards, 1946, p. 175). 


861316—50——7 


92 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Range.—From Kansas south to Louisiana, west to the Pacific, as far 
north as central Oregon, and southward into northern Mexico. 
Absent from the northern Great Plains and some mountainous areas. 

Species.—F our, with five forms recognized, all but one (C. c. collaris) 
occurring in Mexico. 


KEY TO MEXICAN SPECIES OF CROTAPHYTUS 


1. Black ring or rings about neck complete or incomplete; body pattern not 

TOCICUMALCG (ee oe a ee ene 2 ee ee ee ee 2 

No black ring about neck, occasionally a black line present on shoulder; dorsal 

pattern boldly reticulated: 25-2225 ees ce ee ene reticulatus (p. 93) 

2. Collar formed of a single bar or ring; snout more elongate___insularis (p. 93) 

Collar formed of 2 rings, one or both incomplete__--....__---_---------- 3 

3. Tail more distinctly compressed; 4 or 5 scale rows between suboculars and 
labials, and slightly enlarged scales on the middorsal line on tail. 

dickersonae (p. 93) 

Tail not or scarcely discernibly compressed; minimum scale rows between sub- 

oculars and labials 2 (rarely 3); scale rows between supraorbital areas 2 (or 

MATE L yao) Bee sae Se ee ee ee See collaris baileyi (p. 92) 


CROTAPHYTUS COLLARIS BAILEYI “ Stejneger 


Crotaphytus baileyi STEJNEGER, North Amer. Fauna, No. 3, 1890, p. 103, pl. 12, 
fig. 1. 

Crotaphytus collaris baileyi, Stone, Proc. Acad. Nat. Sci. Philadelphia, 1903, p. 
30.—Van DeEnBuRGH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 
104-114; pl. 7—Smitru, Handbook of lizards, 1946, pp. 170-172, pl. 33. 

Crotaphytus collaris Burt, Oce. Pap. Mus. Zool. Univ. Michigan, No. 196, 1928, 


pp. 1-19 (part). 
Crotaphytus collaris collaris, SMirH (nec Say), Univ. Kansas Sci. Bull., vol. 22, 


1935, p. 143, pl. 24, fig. 1. 

Type.—U.S.N.M. No. 15821; C. Hart Merriam and Vernon Bailey 
collectors. 

Type locality —Painted Desert, Little Colorado River, Ariz. 

Range.—Southern Idaho and Oregon southeastward through eastern 
California and northern Baja California to San Luis Potosi. Re- 
ported in Mexico from Baja California: San Salada, Trinidad, Las 
Palmas Canyon, Comondt, La Purisima, 6 miles and 33 miles north of 
Canipole, et cetera; Chihuahua: Chihuahua, Carmen, Progreso, San 
Buenaventura; Durango: 6 miles northeast of Pedricefia; Coahuila: 
Villa Acufia, Monclova, 4 miles west of Saltillo, Cuatro Ciénegas, 
Pueblo Nuevo, Allende; Nuevo Leén: Pesqueria Grande, Santa Catar- 
ina; Sonora: Hermosillo, Sierra Tule, Los Nogales, Rio Grande 61 
miles south of Los Nogales; San Luis Potosi: 30 miles north of Mate- 
huala, Presa de Guadalupe. 

6 With sufficient series it is probable that certain populations now associated with baileyi and collaris will 


be recognized as other subspecies. For instance, certain specimens from San Luis Potosi have three scale 
series between the supraocular regions. C. collaris collaris may occur. 


CHECKLIST OF REPTILES OF MEXICO 93 
CROTAPHYTUS INSULARIS Van Denburgh and Slevin 


Crotaphytus insularis VAN DENBURGH and SLEvin, Proc. California Acad. Sci., 
ser. 4, vol. 11, 1921, p. 96.—Van Densureu, Occ. Pap. California Acad. 
Sci., No. 10, 1922, pp. 114-116. 

Type.—California Acad.Sci. No. 49151; J. R. Slevin collector. 

Type locality —Angel de la Guarda Island, 7 miles north of Pond 
Island, Gulf of California, Baja California. 

Range.—Restricted to type locality. 


CROTAPHYTUS DICKERSONAE 6 Schmidt 


Crotaphytus dickersonae Scumipt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, p. 
638, fig. 2. 
Crotaphytus collaris dickersonae, ALLEN, Occ. Pap. Mus. Zool. Univ. Michigan, 
No. 259, 1933, p. 7. 
Type—U.S.N.M. No. 64451; C. H. Townsend collector. 
Type locality —Tiburén Island, Sonora. 
Range.—Known only from the type locality. 


CROTAPHYTUS RETICULATUS Baird 


Crotaphytus reticulatus Barrp, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 253.— 
Corr, Ann. Rep. U. S. Nat. Mus., 1898 (1900), p. 254, fig. 20.—Smiru, 
Handbook of lizards, 1946, pp. 173-175, pl. 34. 


Type-—Two cotypes, U.S.N.M. No. 2692, Ringgold Barracks, 
John H. Clark collector; No. 2731, Laredo, Tex., Arthur C. B. Schott 
collector. 

Type locality.—Laredo and Ringgold Barracks, Tex., here restricted 
to Laredo. 

Range.—Extreme southern Texas, along the Rio Grande, as far west 
as Eagle Pass, and southward in the northern parts of adjacent states 
in Mexico. Reported in Mexico from Tamaulipas: Mier; Nuevo Leén: 
6 miles west of China, 4 miles west of China, 5 miles east of General 
Bravo, ‘Los Herrars”’; Coahuila: 2 miles north of Nava. 


Genus GAMBELIA Baird 


Gambelia Batrp, United States and Mexican boundary survey, vol. 2, Rept., 
1859, p. 7.—Smitru, Handbook of lizards, 1946, pp. 158-159. 
Genotype—Gambelia wislizenii Baird and Girard. 
Range.—Northern Idaho, Nevada, and Utah southward through 
southeastern California and Baja California, and into western Sonora 
and northern Coahuila. 
Species.—One, with two subspecies. Only the typical form occurs 
in Mexico. 





6s This species has been reduced to subspecifie status on the basis of specimens reported by Allen from 
Hermosillo and 40 miles north of Guaymas. Since we have not seen these specimens, and since he did not 
take into consideration some of the more important characters, we retain the specific status for this form 
pending a more considered revision. 


94 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


GAMBELIA WISLIZENII WISLIZENII (Baird and Girard) 


Crotaphytus wislizenii Barrp and Girarp, in Stansbury’s Explorations and survey 
of the valley of the Great Salt Lake of Utah, 1852, pp. 340-341, pl. 3.— 
Van DensurcH, Occ. Pap. California Acad. Sci., No. 10, vol. 1, 1922, 
pp. 116-128, pl. 8. 

Crotaphytus (Gambelia) wislizenii, Barrp, United States and Mexican Boundary 
Survey, 1859, p. 7. 

Gambelia wislizenii wislizenii, SmirH, Handbook of lizards, 1946, pp. 159-164, pl. 
30. 

Crotaphytus gambelii BArrp and Grrarp, Proc. Acad. Nat. Sci. Philadelphia, 1852, 
p. 126 (California; U.S.N.M. No. 2722; W. Gambell, collector). 

Crotaphytus fasciatus HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, 1852, p. 206 
(‘“‘Jornado del Muerte,’’ Dona Ana County, N. Mex.; U.S.N.M. No. 2736; 
L. Sitgreaves collector). 

Crotaphytus copeii YaRRow, Proc. U.S. Nat. Mus., vol. 5, 1882, p. 441 (U.S.N.M. 
No. 12663; La Paz, Baja California). 

Crotaphytus fasciatus (non Hallowell), Mocquarp, Nouv. Arch. Mus. Hist. Nat. 
Paris, ser. 5, vol. 1, 1899, p. 303, pl. 13, fig. 1 (Las Palmas, Baja California; 
Mus. Hist. Nat. Paris). 

Crotaphytus fasciolatus MocquarD, Bull. Mus. Hist. Nat. Paris, 1903, p. 209 (new 
name for C. fasciatus Mocquard). 


Type.—U.S.N.M. No. 2770; H. Mollhausen collector. 

Type locality—Near Santa Fe, N. Mex. 

Range.—Northern Idaho, Nevada, and Utah southward through 
southeastern California and all of Baja California except northwestern 
corner, western Chihuahua and northern Coahuila. Reported from 
the states of Baja California: Cedros and Magdalena Islands, numer- 
ous mainland localities; Sonora: Tiburén Island, numerous mainland 
localities; Chihuahua: 15 miles south of Juarez, Santa Maria, Pro- 
greso, Lake Santa Marfa, Chihuahua, 11 miles south of Ahumada, 2 
miles south of Moctezuma; Coahuila: Cuatro Ciénegas. 


Genus PHRYNOSOMA Wiegmann 


Tapaja (or Tapaia, p. vi) Oxen, Lehrbuch der Naturgeschichte, 1816, p. 295 
(type, Lacerta orbicularis Linnaeus by present restriction; suppressed). 
Tapayia Gray, Ann. Philos., new ser., vol. 26, 1825, p. 197 (type, Lacerta orbicu- 

laris Linnaeus, monotype; suppressed). 

Tapaya Fitz1ncEer,® Neue Classification der Reptilien, 1826, p. 17 (part only; 
type restricted to T. orbicularis Cuvier=Lacerta orbicularis Linnaeus, by 
Malcolm Smith, Fauna of British India, vol. 2, 1935, p. 211).—Bocourt, 
Mission scientifique au Mexique.. ., Etudes sur les reptiles, livr. 4, 1874, 
p. 220. 

Phrynosoma W1EGMANN, Isis von Oken, 1828, p. 367. 

Batrachosoma FirzincER, Systema reptilium, 18438, p. 79 (type, Phrynosoma coro- 
natum Blainville). 

Tropidogaster FirzIncER (nec Duméril and Bibron), loc. cit. (type, Phrynosoma 
cornutum Gray). 


68 Cuvier (Le régne animal, vol. 2, 1817, p. 35) has been credited with this name by some authorities. He 
did not, however, use the name in latinized form anywhere in the first or 1817 edition of ‘“Le Régne Animal”; 
he uses the name only as “‘les Tapayes.” 


CHECKLIST OF REPTILES OF MEXICO 95 


Anota HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, 1852, p. 182 (type, Anota 
m’callit Hallowell). 

Tapaya Grrarp (nec Fitzinger), in Wiikes, United States Exploring Expedition, 
Herpetology, 1858, p. 394 (type, Lacerta orbicularis Linnaeus). 

Doliosaurus GIRARD, op. cit. p. 407 (type, Phrynosoma platyrhinos Girard). 

Genotype.—Lacerta orbicularis Linnaeus. 

Range.—Nebraska, Kansas, Oklahoma, and Texas westward to the 
Pacific, northward to British Columbia and Montana, and southward 
through Mexico to Guatemala. 

Species —Fifteen, with about 26 forms recognized; 21 occur in 
Mexico. 

KEY TO MEXICAN SPECIES OF PHRYNOSOMA 


1. Ventral abdominal scales smooth (a few scales may be keeled on breast of 
solare and a few specimens may show faint keeling on some of abdominal 


scales, but this form will have four occipital spines) _______________-_- 6 
Ventral abdominal scales distinctly keeled (rarely indistinctly keeled in 
COTTE LU TETID re at ey ae ep ee ae ee nen SO RA 2 

2. Horns (or horny spines) present on back of skull_____________________- 3 


No horns or horny spines on back of skull, the scales that form spines in other 
species are low rounded protuberances; two flaring temporal expansions on 
back of head separated by a medial notch; lower jaw with great vertical 
development, the chin shields separated from the infralabials by 4 or 5 
BCALCMLOW Sse ONMINE = gay eee ee SO es Sores SON Be ditmarsi (p. 104) 

3. Tail at least half length of body (usually more); large species with 2 lateral 
abdominal fringes of soft spines more or less clearly developed __---- 4 
Tail shorter or but little longer than head; smaller species, with 1 series of 
lateralisoftes pines: tts sas 2 ete ha eet fell ey, haere fl, SET Bae 5 
4. Ventral scales large, 30 to 35 across widest part of belly; scales surrounding 
spines of upper lateral fringe very prominent; greater part of gular scales 
arranged in slightly serrate longitudinal series; 2 very prominent series of 
large scales widely separated on back; tail about as long as body; length, 
iA anata Poteet 2a Sage AIRE E ee S e okh te Sk asio (p. 102) 

Ventral scales smaller, 55 to 70 across widest part of abdomen; 3 temporal and 
2 occipital spines well developed; 3 brown bands across frontal area; 3 
radiating lines below eye; rarely ventral keeling is indistinct; length, 140 
SPENT Ne es cae RN ea eR ies oe cornutum (p. 99) 

5. Outer temporal part of skull prolonged backward into a very large ‘‘horn”’ 
several times size of small occipital spines; tail as long as head or slightly 
jonger-p9— 110 femoraliporess=_2 2" ss - ee ls 2e Beee a ee taurus (p. 104) 

Outer temporal region not prolonged into a large horn; temporal spines about 
the same size as occipital spines; tail shorter than head; femoral pores 
BOUL WO 0 Becca rene ere crete een ey ene braconnieri (p. 100) 

6. No ear-opening present; tympanum superficial, covered with scales (very 
rarely unscaled in some Mexican specimens of modestum and some 

TT COLULT pm omnis a puesme ee panier! 2) a, selec ienio! si hte ee EOD ar se ee epee ee eh 7 

Ear opening present, tympanum usually more or less deeply sunk__------ 10 

7. No Jateral fringe of soft spines on sides of abdomen; occipital and temporal 
spines present but rather short; chin shields bordering labial series through- 

out; greater part of tail very slender and subcylindrical, not gradually 
tapering; venter white; chin usually speckled; 84 mm__modestum (p. 101) 

At ieasth series of lateral soft spines=__-./.--_--..-.-....=.--.=.==-+- 8 


96 


10. 


11. 


12. 


13. 


14. 


15. 


16. 


BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


. Usually less than 12 femoral-preanal pores, the pore scales usually separated 


byl. or morevintervening scalesi22222 {esas 2 es eee Se eae eee 9 
Pore series 17-23, the pores surrounded by several small scales and usually 
contiguous; temporal and occipital horns very elongate; 2 series of lateral 
fringes on abdomen; an indistinctly enlarged series of longitudinal gular 
seales on each side of chin, more evident posteriorly; a dark dorsal streak; 
immaculate white on venter; tail much flattened; occasionally traces of 
abdominaltkeeling presenteso2 2224 Sos See eee m’callii (p. 99) 


. Pore series about 8 on each side; 3 enlarged temporals and 2 enlarged occipital 


spines, tending to curve laterally somewhat; small species, 72 mm. snout to 
vent; venter immaculate white___._______---- platyrhinos goodei (p. 102) 
Pore series 7-12, the temporal and occipital horn much reduced; superciliary 
spine short but heavy and prominent; a single series of scales in lateral 
fringe; venter usually punctate with dark; only rarely is the tympanum 
Visiblétin! sad whe bee beet ae) ete platyrhinos platyrhinos (p. 101) 
Four much-elongated, flattened occipital horns, contiguous with a like tem- 
poral series, all directed more or less in same general plane; breast scales 
keeled; 1 distinct lateral fringe and a small secondary fringe indicated. 
solare (p. 104) 
Never more than two strongly developed occipital horns (occasionally a very 
small median single scale may be present)_..._..------------------ 11 
Occipital horns at least twice as long as basal width; chin shields forming 
serrate series strongly visible from dorsal view of head; gulars with serrate 


series /ofvenlargedscales=- ete alae ee ee eeee eas coronatum group 17 
Occipital horns not twice as long as basal width; no serrate chin shields or 
specializedyserrate series in) gularsia_ 2. £ls- 5 Sect eued ed sees 12 


Head not strongly notched or emarginate between occipital horns, distance 
between bases of horns one-half to rarely one and one-half times basal 
width of horn; occipital horns distinctly larger than superciliary horns. 

orbiculare group 14 

Head strongly notched or emarginate between occipital horns; horns very 
short, their bases separated by a distance equal to 2 to 3% times basal width 
of a horn; occipital and superciliary horns subequal in size. 

douglassii group 13 

Tail length a little less than half snout-to-vent length but distinctly longer 

than width of head; gular scales not convex; chest scales smooth. 
douglassii hernandesi (p. 100) 

Tail very short, very slender, length less than width of head; gular scales 

convex, and chest scales keeled_-_-_-_-__- douglassii brachycercum (p. 100) 


Occipital spines nearly erect, while temporal spines approach the horizontal, 
all relatively long; top of head conspicuously concave, superciliary region 
Cle vate cer nr ey erie) Bee keen y eer Cape eeeeny eae a ee ee ever ee Ua) boucardi (p. 98) 

Occipital spines on same plane as temporals, approaching horizontal, all 
shoxrter;.¢op of head flat. <= S22 e222 eo es Se ee 15 


Temporal horns not or barely extending backward as far as posterior tip of 
occipival horns. so. 2225. (Ss ee ee Se eee 16 

Temporal horns extending backward farther than occipital horns. 
orbiculare cortezii (p. 98) 


Occipital horns extending posteriorly farther than posterior temporals; pre- 
anal area with preanal pores not confined to a single row (usually several 
irregularly placed pores or 2 longitudinal rows of 2 to 4 scales). 

orbiculare orbiculare (p. 97) 


CHECKLIST OF REPTILES OF MEXICO 97 


Occipital and posterior temporal horns extending back an equal distance (or 
very nearly so); femoral pores 14-14, confined to single row. 

orbiculare dugesii (p. 98) 

17. Fourth (or fifth) temporal horn (counting forward) sharply pointed and 


greatly exceeding fifth (or fourth) in length._.._.._.._.------------ 18 
Fourth and fifth temporal horns blunt, of nearly equal size, or fifth slightly 
larpera2 shot fee ook. tee eh eyes a ee ae Sek Le 20 

18. Nasal opening large; distance between inner edges less than 1% times maxi- 
MMUMEGIAMCLEr: Of NOStnet ee ee ee cerroense (p. 103) 
Nasal opening moderate, the distance between inner edges more than 1% 
Gime FETS OL MOSUTME sae re rere ft oe Sa ee ce a oe ager 19 

19. Postrictal spine absent or rudimentary__coronatum coronatum (p. 102) 
Postrictal, spine, present... . -...---—.-=-....- coronatum jamesi (p. 102) 


20. Frontal plates in adults smooth, convex_._-coronatum blainvillii (p. 103) 
Frontal plates in adults rough, striated or rugose, and flat or peaked. 
coronatum frontale (p. 103) 
PHRYNOSOMA ORBICULARE ORBICULARE ® (Linnaeus) 


[Tapayazin, Lacertus orbicularis HERNANDEZ, Plantas y animales de la Nueva 
Espafia, etc., 1651, C. xvi, p. 327, fig. (unnumbered).] 

Lacerta orbicularis LINNAEUS, Systema naturae, ed. 12, vol. 1, 1789, p. 1062 (part, 
based on Hernandez). 

Agama orbiculaire, Cuvier, Régne animal, vol. 2, 1817, p. 35. 

Phrynosoma orbiculare, W1EGMANN, Isis von Oken, 1828, p. 367. 

Phrynosoma orbiculare orbiculare, Smiru, Trans. Kansas Acad. Sci., vol. 37, 1934, 
p. 290, pl. 11, fig. 2, pl. 12, fig. 5. 

Phrynosoma wiegmanni Gray, The zoology of Captain Beechey’s voyage... ., 
1839, p. 96 (type unknown; type locality, ‘‘Mexico,’’ here restricted to 
Mexico City, Distrito Federal). 

Tapaya orbicularis longicaudatus Ducks, La Naturaleza, ser. 2, vol. 1, 1888, p. 117 
(Alf. Dugés Mus., Guanajuato, Guanajuato; Valley of Mexico, here re- 
stricted to México, D. F.).—Smira and Necker, Anal. Esc. Nac. Cienc. 
Biol., vol. 3, 1943, pp. 216-218, pl. 2, figs. 1, 3. 

Type.—Presumably the unnumbered figure in Hernandez, loc. cit. 

Type locality—Mexico (by inference), here restricted to México, 
be: 

Range.—The central plateau of Mexico from Chihuahua and 
Nuevo Leén southward to Michoacan, Morelos, and Puebla. Re- 
ported from Mézico: Toluca, Lerma, San Andrés, 11 miles south of 
Mexico (city), Teotihuacan, Zumpango; Puebla: Puebla, 15 kilo- 
meters northwest of San Martin, Matamoros near Tezuitlin; Distrito 
Federal: México, Tlalpam, Rio San Juan de Dios, Tlalnepantla, 
between Navitas and Chalco; Hidalgo: Guerrero near Real del 
Mineral, Actopan, El Chico Parque Nacional, 2-3 miles west of 
Tulancingo, near Pachuca; Morelos: near Tres Marias (Km. 57); 
Chihuahua: south of Chihuahua City; Samachique, 21 miles south 
of Mifiaca; Durango: Coyotes, El Salto, Ciudad; Jalisco: North of 

$1 We are unable to agree to the idea of the conspecificity of the Mexican orbiculare group and the douglassit 


group. We feel that among other important distinguishing characters the posterior indentation of the 
skull, so marked in the latter group, has been overlooked in the past. 


98 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Rio Santiago, Guadalajara, Bolafios; Guanajuato: Guanajuato (city); 

San Luis Potosi: Jesis Maria, San Pedro; Querétaro: no specific 

locality; Nuevo Leén: Hacienda Pablillo; Tlaxcala: 13 kilometers 

northeast of Tlaxcala; Michoacdn: “above 3000 feet’? (Jorullo?); 

Zacatecas: Plateado, Sierra Madre. 

PHRYNOSOMA ORBICULARE CORTEZII (Aug. Duméril and Bocourt) 

Phrynosoma orbiculare, WIEGMANN, Herpetologia Mexicana, 1834, p. 53, pl. 8, 
figenle 

Tapaya orbicularis var. A, Bocourt, ibid. (infra), livr. 4, 1874, pp. 223-224, pl. 11, 
fig. 2. 

Tapaya Cortezii Auc. Dumirit and Bocovurt,® Mission scientifique au 
Mexique . . ., Etudes sur les reptiles, livr. 1, 1870, pl. 9, fig. 2, 2a-g. 

Phrynosoma orbiculare cortezii, BONLENGER, Catalogue of the lizards in the British 
Museum, ed. 2, vol. 2, 1885, p. 242.—Smirn, Trans. Kansas Acad. Sci., vol. 
37, 1934, pp. 291-292, pl. 11, fig. 1, and pl. 12, fig. 6. 

Type.—Mus. Hist. Nat. Paris; Lucien Biart collector. 

Type locality—‘Hacienda del Jasmin,’ between ‘Orisaba and 
Cordoba,” Veracruz. 

Range.—Eastern half of Puebla and adjoining parts of Veracruz. 
Reported from Veracruz: 15 miles east of San Marcos, 5 kilometers 
east of Las Vigas, Perote, Jalapa, Orizaba; Puebla: San Diego south 
of Tehuacan, Atzitzintla. 

PHRYNOSOMA ORBICULARE DUGESII (Aug. Duméril and Bocourt) 

Tapaya Dugesiit ® Ava. Dumférit and Bocourt, Mission scientifique au Mex- 
ique . . ., Etudes sur les reptiles, livr. 1, 1870, pl. 11, fig. 3, 3a-g. 

Phrynosoma orbiculare (var.) dugestt, BoULENGER, Catalogue of the lizards in the 
British Museum, ed. 2, vol. 2, 1885, p. 248. 

Tapaya orbicularis var. B, Bocourt, ibid., livr. 4, 1874, pp. 224-225, pl. 17, 
fig. 3, 3a—g. 

Type.—Mus. Hist. Nat. Paris, two cotypes; Alfredo Dugés collector. 

Type locality —Colima. 

Range.—Known only from Colima. 

PHRYNOSOMA BOUCARDI (Aug. Duméril and Bocourt) 

Tapaya Boucardii Auc. Dumfrit and Bocourt, Mission scientifique au Mex- 
ique . . ., Etudes sur les reptiles, livr. 1, 1870, pl. 11, fig. 4, 4a-g.—Bocourt, 
tbid., livr. 4, 1874, pp. 225-226. 

Phrynosoma boucardii, BouLENGER, Catalogue of the lizards in the British 
Museum, ed. 2, vol. 2, 1885, p. 248.—Smiru, Journ. Washington Acad. Sci., 
vol. 39, 1949, p. 38. 


Type.—Mus. Hist. Nat. Paris, two males, two females; A. Boucard 
collector. 
Type locality. ‘Plateau of Mexico,” here restricted to Zimap4n, 
Hidalgo. 
68 The name cortezii must be accredited to both authors of livr.1. The name when it appears is Tapaya 
cortezii (nobis), and when quoted by Bocourt both names are used. 


69 In 1870 the name was used as a full species, when associated with the figure. The name is dropped in 
the livr. 4 of the same work where it is regarded as a variety B of Tapaya orbicularis. 


CHECKLIST OF REPTILES OF MEXICO 99 


Range.—Central plateau region. Reported from Guanajuato: 
Leén; Hidalgo: Zimapan. 
PHRYNOSOMA CORNUTUM (Harlan) 


Agama cornuta HARLAN, Journ. Acad. Nat. Sci. Philadelphia, vol. 4, 1825, p. 299, 
1. 20. 

gas cornuta, CuviER, Régne animal, vol. 2, 1829, p. 37. 

Phrynosoma cornutum, Gray, in Griffith, Cuvier’s ‘Animal Kingdom” (Appen- 
dix), 1831, p. 9.—Smiru, Handbook of lizards, 1946, pp. 290-293, pl. 74. 

Phrynosoma bufonium W1EGMANN, Isis von Oken, vol. 21, 1828, pp. 367-369 (type 
locality uncertain (see Herpetologia Mexicana, 1834, p. 54), here restricted 
to Los Nogales, Sonora; collector unknown; Zool. Mus. Berlin). 

P{hrynosoma] harlanii W1EGMANN, Herpetologia Mexicana, 1834, p. 54 (substitute 
name for Agama cornuta Harlan). 

Phrynosoma planiceps HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, 1852, 
p. 178 (western Texas near the Rio Grande, here restricted to El Paso). 
Phrynosoma cornutum planiceps, BOoULENGER, Catalogue of the lizards in the 

British Museum, vol. 2, 1885, p. 246. 

Type.—Unknown. 

Type locality—Great Plains east of the Rocky Mountains, here 
restricted to Fort Riley, Geary County, Kans. 

Range.—Kansas to Texas, west to Arizona, south to Durango and 
San Luis Potosi. Reported from Tamaulipas: Alta Mira, Mier, Soto 
la Marina, Charco Escondido, Matamoros, Tampico, Marmolejo; 
Nuevo Leén: Monterrey, Pesqueria Grande, 20 miles south of Nuevo 
Laredo, San Juan, 1 mile south of Nuevo Laredo; San Luis Potosi: 
Charco Largo; Coahuila: Castafiuelos, Alamo de Parras, Monclova, 
San Pedro, Patos, Rio Nazas, El Zacate, Buena Vista, Mtizquiz, et 
cetera; Durango: 7 miles south of La Loma, 25 miles north of Berme- 
jillo; Chihuahua: Batopilas, 5 miles north of Moctezuma, Progreso, 
Chihuahua (city), Huajuquilla (Jiménez), Sauz, Guzman, et cetera; 
Sonora: Los Nogales. 

PHRYNOSOMA M’CALLII Hallowell 


Anota m’callii HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, 1852, p. 182.} 
Doliosaurus m’calli, GiraRD, Wilkes United States Exploring Expedition, Herpe- 
tology, 1858, p. 408. 
Phrynosoma m’calli, STEJNEGER, North Amer. Fauna, No. 7, 1893, p. 190.—Van 
Densurau, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 428-430, pl. 
37. 
Anota maccalli,” Bryant, Univ. California Publ. Zool., vol. 9, No. 1, 1911, pp. 5, 
54, pl. 7. 
Type.—Unknown. 
Type locality—Great Desert of the Colorado, between Vallecita 
and Camp Yuma, about 100 miles east of San Diego. 
Range.—Northwestern Sonora and northeastern Baja California; 
in United States: Southeastern California and southwestern Arizona. 


7 The senior author has expressed the opinion elsewhere that if the characters on which the genus Anofe is 
based should prove constant this name should replace Phrynosoma for this species. 


100 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Reported from Sonora: East bank of Colorado River, 5 miles south of 
United States border. Baja California: ‘“Yuhu Basin north of 
Petrified Forest.” 


PHRYNOSOMA DOUGLASSIT HERNANDESI (Girard) 


Tapaya hernandesi Girard, Wilkes United States Exploring Expedition, Herpe- 
tology, 1858, p. 395. 

Tapaya hernandezii, Barrp, United States and Mexican Boundary Survey, vol. 2, 
Rept., 1859, p. 8. 

Phrynosoma hernandesi, StEsNEGER, North Amer. Fauna, No. 3, 1890, p. 112. 

Phrynosoma douglassit hernandesi, Corr, Ann. Rep. U.S. Nat. Mus., 1898 (1900), 
p. 413, fig. 70.—Van DeEnsBurau, Occas. Pap. California Acad. Sci., No. 10, 
vol. 1, 1922, pp. 382-386, pl. 31. 

Phrynosoma orbiculare hernandesit, KuauBER, Bull. Zool. Soc. San Diego, vol. 4, 
1939, pp. 91-93. 

Phrynosoma douglassii Corr, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 302. 


Type.—U.S.N.M. No. 197, a cotype. 

Type locality —New Mexico and ‘Sonora,’ restricted to Santa Fe, 
N. Mex. (Cope, loc. cit., 1900). 

Range.—Northern Sonora, Chihuahua; in United States: Colorado, 
southern New Mexico, southeastern and central Arizona, south- 
western Texas. Reported from Sonora: ?‘Mexican Boundary line’; 
Chihuahua: Ramos. 


PHRYNOSOMA DOUGLASSII BRACHYCERCUM Smith 


Phrynosoma douglassit brachycercum Smitu, Proc. U. 8. Nat. Mus., vol. 92, 1942, 
pp. 361-363. 


Type.—U.S.N.M. No. 23993; Edward Palmer collector. 
Type locality —‘ Durango.” 
Range.—Known only from the type locality. 


PHRYNOSOMA BRACONNIERI Duméril and Bocourt 


Phrynosoma Braconniert Auc. Dumérit and Bocourt, Mission scientifique au 
Mexique . . ., Etudes sur les reptiles, livr. 1, 1870, pl. 12, fig. 7, 7a-—g.— 
Bocourt, livr. 4, 1874, pp. 233-234.—BouLencEr, Catalogue of lizards in 
the British Museum, vol. 2, 1885, p. 248.—Smiru, Trans. Kansas Acad. Sci., 
vol. 37, 1934, pp. 287-289. 


Type.—Mus. Hist. Nat. Paris, adult; collected by Aide-Major 
Jacob; several specimens collected by Alfredo Dugés and A. Boucard. 

Type locality—Oaxaca. 

Range.—Extreme southern edge of the central Mexican plateau, in 
semiarid portions of Puebla and Oaxaca (a specimen has been taken 
within a few hundred meters of the border of Veracruz, and very 
probably the species occurs in the mountains in the western part of 
that state). Reported from Pucbla: Tehuacd4n, 14 miles northeast of 
Tehuacén, near Cacoloapam (Km. 226); Oaxaca: No specific locality, 
unless Oaxaca (city) is meant. 


CHECKLIST OF REPTILES OF MEXICO 101 
PHRYNOSOMA MODESTUM Girard 


Phrynosoma modestum GrrarpD, in Stansbury, Exploration and survey of the 
valley of the Great Salt Lake of Utah, 1852, pp. 361-362 and 365, pl. 6, figs. 
4-8.—Van DensureH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 
430-433, pl. 38. 

Doliosaurus modestus, GiraARD, Wilkes United States Exploring Expedition, 
Herpetology, 1858, p. 409. 

Anota modesta, Corr, Amer. Nat., vol. 30, 1896, p. 1014. 


Type.—U.S.N.M. (?), several cotypes; General Churchill and ‘the 
party under Col. J. D. Graham”’ collectors. 

Type locality.—The Rio Grande west of San Antonio, Tex. (Churchill), 
and from between San Antonio and El Paso (Graham); here restricted 
to Las Cruces, N. Mex. 

Range.—Western Texas to Arizona, southward to San Luis Potosf. 
Reported from Nuevo Ledn: Santa Catarina, Pesquerfa Grande. 
Chihuahua: 11 miles south of Ahumada, Rio Santa Marfa, Guzman, 
near Janos, Juarez, Chihuahua (city); Coahuila: 1.5 miles west of 
Saltillo, Mtsquiz, Jaral, Castafiuelas, Palau, Alamo de Parras, San 
Pedro, Agua Nueva, Monclova, Buena Vista, Rio Nazas, Cuatro 
Ciénegas, et cetera; Sonora: Nariz Temporal, Los Nogales, Sierra de 
la Nariz; Durango: between Lerdo and La Goma, Pedricefia, 5 kilo- 
meters west of Torreén, 25 miles north of Bermejillo, 7 miles south of 
La Loma; Zacatecas: 3 miles west of La Colorada; San Luis Potosi: 
near Charco Largo (130 kilometers north of San Luis Potosi city), 
Hacienda La Parada. 


PHRYNOSOMA PLATYRHINOS PLATYRHINOS Girard 


Phrynosoma platyrhinos Girarp, Stansbury’s Explorations and survey of the 
valley of the Great Salt Lake of Utah, 1852, pp. 361, 363-364, pl. 7, figs. 1-5.— 
Van Densoureu, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 421- 
426, pl. 36. 

Doliosaurus platyrhinos, GrraRD, in Wilkes, United States Exploring Expedition, 
Herpetology, 1858, p. 409. 

Anota platyrhina, Corr, Ann. Rep. U.S. Nat. Mus., 1898 (1900), p. 448, fig. 81. 

Phrynosoma platyrhinos platyrhinos, KLAUBER, Bull. Zool. Soc. San Diego, vol. 14, 
1939, p. 13. 

Anota calidiarum, Corr, Amer. Nat., vol. 30, No. 358, 1896, p. 333 (Death Valley, 
Calif.). 

Phrynosoma calidiarum, Dirmars, The reptile book, 1907, p. 157, pl. 46, figs. 10, 
14, pl. 50, fig. 2. 


Type.-—U.S.N.M. No. 189?; H. Stansbury collector. 

Type locality —Great Salt Lake, Utah. 

Range.—Northeastern Baja California; widespread in Nevada, 
Idaho, Utah, Washington, California, and Arizona. Reported in Baja 
California: 20 miles north of San Felipe, San Felipe Bay, Pozo, San 
Agustin, Seven Wells on the Salton River. 


102 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


PHRYNOSOMA PLATYRHINOS GOODEI Stejneger 


Phrynosoma goodei StEINEGER, North Amer. Fauna, No. 7, 1898, p. 191 (foot- 
note), pl. 2, figs. 8a—3c. 

Anota goodei, Corg, Ann. Rep. U. 8S. Nat. Mus., 1898 (1900), p. 442, fig. 80. 

Phrynosoma platyrhinos goodet, KLAUBER, Copeia, 1935, pp. 178-179. 


Type.—U.S.N.M. No. 8567a; T. H. Streets collector. 

Type locality.—‘‘Coast deserts of the state of Sonora, Mexico,”’ here 
restricted to Puerto Libertad. 

Range.—Known only from the gulf coast of Sonora. Reported 
from Puerto Libertad, 8 miles northwest of Alamo Muerto, west 


coast of Sonora. 
PHRYNOSOMA ASIO Cope 


Phrynosoma asio Corr, Proc. Acad. Nat. Sci. Philadelphia, vol. 16, 1864, pp. 
178-179.— BouLEencErR, Catalogue of lizards in the British Museum, vol. 2, 
1885, p. 244. ; 

Batrachosoma asio, Bocourt, Mission scientifique au Mexique. . . , Etudes 
sur les reptiles, livr. 4, 1874, pl. 17, fig. 9, 9a-c. 

Phrynosoma spinimentum Peters, Monatsb. Akad. Wiss. Berlin, 1873, pp. 742- 
743 (Tehuantepec; Mus. Berlin No. 7219). 

Type.—U.S.N.M. (?); John Xantus collector. 

Type locality Colima, Colima (by present restriction). 

Range.—F¥rom Colima through coastal Michoac4n, Guerrero, Oaxaca 
to Chiapas, and in the Balsas Basin. In Central America, recorded 
from Guatemala. Reported from Oazaca: San Pedro, Tequesixtlin, 

San Gerénimo, Ranchero Poso Rio, vicinity of Tehuantepec, vicinity 

of Salina Cruz; Guerrero: Mexcala (on Balsas River); Michoacdn: 

La Salada, Cofradia, San Salvador, Apatzingan; Colima: Tecomaén, 

Colima (city); Chiapas: no definite locality. 


PHRYNOSCMA CORONATUM CORONATUM (Blainville) 


Agama (Phrynosoma) coronata BLAINVILLE, Nouv. Ann. Mus. Hist. Nat. Paris, 
vol. 4, 1835, p. 284, pl. 25, figs. 1-1ce. 

Phrynosoma coronatum coronatum, LiNspALE, Univ. California Publ. Zool., vol. 
38, 1932, p. 368.—KuauBER, Copeia, 1936, pp. 103-110, fig. 1—Tervis, 
Copeia, 1944, pp. 13-15, fig. 2, map. 

Type.—Mus. Hist. Nat. Paris; P. E. Botta collector. 
Type locality. —‘‘California,” here restricted to Cape San Lucas, 

Baja California. 

Range. —Southern part of Baja California, north to about lat. 26° N. 


PHRYNOSOMA CORONATUM JAMESI Schmidt 


Phrynosoma jamest Scumipt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 
668-669, pls. 55, 56. 
Phrynosoma coronatum jamesi, KLAUBER, Copeia, 1936, p. 110.—Trvis, Copeia, 
1944, pp. 138-15, map fig. 2. 
Type.—U.S.N.M. No. 64450; H. Townsend collector. 


Type locality.—Shore of San Bartolomé Bay, Baja California. 


CHECKLIST OF REPTILES OF MEXICO 103 


Range.—Central Baja California, between lat. 26° N. and lat. 
28°30’ N. 
PHRYNOSOMA CORONATUM FRONTALE Van Denburgh 


Phrynosoma frontalis VAN DenBURGH, Proc. California Acad. Sci., ser. 2, vol. 4, 
1894, pp. 296-298. 

Phrynosoma blainvillit frontale, VAN DeNBURGH, Occ. Pap. California Acad. Sci., 
No. 5, 1897, p. 95; Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 
395-401, pl. $3.—Kuavuger, Copeia, 1936, pp. 103-110. 

Phrynosoma coronatum frontale, LinspaLe, Univ. California Publ. Zool., vol. 38, 
1932, p. 368.—Tervis, Copeia, 1944, pp. 13-15, fig. 2 (map). 

Phrynosoma schmidti Barsour, Proc. New England Zool. Club, vol. 7, 1921, 
p. 1138 (Mus. Comp. Zool. No, 15142; W. W. Brown collector; Cedros Is- 
land). 

Phrynosoma nelsoni Scumipt, Bull. American Mus. Nat. Hist., vol. 46, 1922, 
p. 666 (Amer. Mus. Nat. Hist. No. 37585; E. W. Nelson and E. A. Goldman 
collectors; San Quintin, Baja California). 


Type.—Stanford Univ. Mus. No. 93, adult male; C. H. Gilbert 
and W. W. Price collectors. 

Type locality —Bear Valley, San Benito County, Calif. 

Range.—Discontinuous; central California east of the Sierra Nevada 
south to but not including extreme southwestern California; north- 
western coast of Baja California except a small area adjacent to the 
United States border; Cedros Island. 


PHRYNOSOMA CORONATUM BLAINVILLII Gray 


Phrynosoma blainvilliti Gray, The zoology of Captain Beechey’s voyage... , 
Reptiles, 1839, p. 96, pl. 29, fig. 1. 

Phrynosoma blainvillii blainvilliit, GRINNELL and Camp, Univ. California Publ. 
Zool., vol. 17, 1917, p. 164.—Van Dernsurea, Occ. Pap. California Acad. 
Sci., No. 10, 1922, pp. 388-395, pl. 32.—Kuavuser, Copeia, 1936, pp. 103-110, 
fig. 2. 

Phrynosoma ochoterenai TreRRon, Anal. Inst. Biol., vol. 3, 1932, p. 109 (type 
formerly in Mus. Hist. Nat., Mexico, now lost; J. M. Gallegos collector; 
Tecate, Baja California). 


Type.—Brit. Mus. Nat. Hist., young specimen; de Blainville 
collector. 
Type locality “California,” here restricted to San Diego. 
Range.—Southern California and extreme northeastern Baja 
California. 
PHRYNOSOMA CERROENSE Stejneger 


Phrynosoma cerroense STEJNEGER, North Amer. Fauna, No. 7, 1893, p. 187 (in 
footnote).—Van DrnspuraH, Occ. Pap. California Acad. Sci., No. 10, 1922, 
pp. 401-403.—Ktuavser, Copeia, 1936, p. 108. 
Type-—U.S.N.M. No. 11977; L. Belding collector. 
Type locality——Cerros” Island, Pacific coast of Baja California 
[=Cedros Island]. 


Range.—Known only from the type locality. 


104 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


PHRYNOSOMA TAURUS Dugés 


Phrynosoma taurus Ducés, MS. 1868, fide Duméril and Bocourt loc. cit., p. 234; 
and La Naturaleza, 1869, p. 143 (nomen nudum).—Avua. Dumférit and 
Bocourt, Mission scientifique au Mexique ... , Etudes sur les reptiles, 
livr. 1, 1870, pl. 12, figs. 8, 8a-g.—Duats, La Naturaleza, 1873, pp. 302-305, 
figs. 1-4 (this description bears the date Nov. 23, 1870, but was not pub- 
lished until 1873; there were two specimens, one from ‘‘Cozcatlain,”’ the other 
from Estado de Puebla).—Bocourt, Mission scientifique au Mexique ... , 
Etudes sur les reptiles, livr. 4, 1874, pp. 234-235.—BovuLencer, Catalogue 
of lizards in the British Museum, vol. 2, 1885, p. 249.—SmirxH and NEcKER, 
Anal. Ese. Nac. Cienc. Biol., vol. 3, 1948, pp. 208-210, pl. 3, figs. 1-2. 


Type."—Mus. Hist. Nat. Paris, 9 cotypes?; M. Boucard collector. 

Type locality.—‘‘Matamoras Iztcar,” Puebla. 

Range.—Puebla and Guerrero. Reported from Puebla: Zapotitlan, 
San Diego near Tehuac4an, Cozcatlan (Dr. Bedel collector), Izticar 
de Matamoros; Guerrero: 5 miles north of Chilpancingo, Amula 


8,000 feet. 
PHRYNOSOMA SOLARE Gray 


Phrynosoma solaris Gray, Catalogue of the specimens of lizards in the collection 
of the British Museum, 1845, p. 229. 

Phrynosoma solare, VAN DrengureH, Proc. California Acad. Sci., ser. 2, vol. 5, 
1895, p. 115; Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 406-409, pl. 34. 

Phrynosoma regale GirRARD, in Wilkes, United States Exploring Expedition, 1858, 
p. 406 (U.S. N. M. No. 161; Sierra de la Nariz, near Zuiii, Sonora). 


Type.—Brit. Mus. Nat. Hist. (not listed in Boulenger, Catalogue of 
lizards in the British Museum, vol. 2, unless it is spec. a, o [Cali- 
fornia], listed under Phrynosoma regale, p. 245); ? de Blainville col- 
lector. 

Type locality. —‘‘California,’”’ here restricted to Yuma, Ariz. 

Range.—Southern Arizona southward into northeastern Baja 
California, western Sonora and into Sinaloa. Reported from Sonora: 
Santa Ana, Nogales, Hermosillo, 15 to 20 miles south and 45 miles 
southwest of Hermosillo, Tiburén Island, Empalme, 10 miles north- 
west of Guaymas, Alamos, Guirocoba; Baja California: Las Animas 
Bay; Sinaloa: Ahome, Sierra de Choix. 


PHRYNOSOMA DITMARSI Stejneger 


Phrynosoma ditmarsit StEsJNEGER, Proc. U. S. Nat. Mus., vol. 29, 1906, p. 565.— 
Dirmars, The reptile book, 1907, p. 154, pl. 46, figs. 3, 7, pl. 48, fig. 1.— 
Smith, Handbook of lizards, 1946, pp. 297-299, pl. 97. 


Type.—U.S.N.M. No. 36022; Mr. Eustace collector. 


71 Whether the specimens mentioned above are really the types may be open to question. A description 
in manuscript was made by Dugés from a specimen in Mexico and presumably sent from Mexico to August 
Duméril in Paris. The Paris Museum had nine specimens of the same species. Drawings were made of 
one, and the drawings published under the name Phrynosoma taurus Dugés. Since the specimen figured 
is the first description or indication, we believe that the original of the drawing becomes the type and this, 
we believe, is a specimen in the Museum d’Histoire Naturelle de Paris. Thus the type specimen was 
presumably never seen by the author of the name. The specimen described by Dugés is in the Museum 
“Alfredo Dugés,’’ Colegio del Estado de Guanajuato. 


CHECKLIST OF REPTILES OF MEXICO 105 


Type locality.—State of Sonora, not far from the boundary of 
Arizona. 
Range.—Known only from the type locality. 


Genus SCELOPORUS Wiegmann 


Sceloporus W1IEGMANN, Isis von Oken, 1828, p. 369. 

Tropidolepis CuviER, Régne animal . , . , ed. 1, vol. 2, 1817, p. 38 (type, Agama 
undulata Daudin). 

Lysoptychus Corr, Proc. U. S. Nat. Mus., vol. 11, 1888, pp. 397-398 (type, L. 
lateralis Cope=Sceloporus couchii Baird). 


Genotype.—Sceloporus torquatus Wiegmann. 

Species.—In all, 104 forms are recognized at present, consisting of 
54 species representing 15 groups. Eighty-nine forms of 52 species 
are known from Mexico. 

Range. —Southern Canada on the west and southern New York on 
the east south throughout North and Central America to western 


Panama. 
KEY TO MEXICAN GROUPS OF SCELOPORUS 


i Postremoral’ dermal pocket present=* 2225225 > 5 fe oe See ee 2 
Eostiemoral dermal pocket absent. 225-2 222 20 ee Se ee eee 4 
2. Postrostral scales absent; nasals and internasals in contact with rostral. 
maculosus (p. 133) 
Postrostral scales present; internasals and nasals separated from rostral__ 3 
3. Tail strongly compressed in males, rounded and light pink in females; femoral 
pores 24’ or more on'each sides 22-2 - ee pyrocephalus (p. 138) 
Tail rounded in both sexes, not pink in females; femoral pores 20 or less. 
variabilis (p. 129) 
2 soterar poudy scales not mbricates= 2: 2>=222 222s * Seow es setae ee 5 
PaperainoOdy Seales uIMDriCAtOns-— fon! Meas ne Se ae ee Sete nee 6 
5. Preanal scales keeled in females; males with poorly developed postanals; no 
distinct belly patches in males; tail over twice length of body. 
utiformis (p. 135) 
Preanal scales smooth in females; males with well-developed postanals; 
lateral belly patches distinct in males; tail less than twice length of body. 
merriami (p. 132) 
6. No postrostrals; nasals and internasals in contact with rostral____._______- 7 
Postrostrals present; nasal and internasals separated from rostral_--_-__-_-_- 8 
7. Dorsal scales 50 or more; femoral pores 17 or more; preanals smooth in 
RTE eee i ak ee BR Se es SE Se a scalaris (p. 136) 
Dorsal scales 46 or less; femoral pores 16 or less; preanals keeled in females. 
siniferus (p. 133) 
8. Ventral scales, at least laterally, pointed, not notched; preanal scales keeled 
in females; no lateral belly markings in males; femoral pore series widely 
separated medially; two postrostrals__..._---.--------- siniferus (p. 133) 
Ventral scales notched, or, if pointed, femoral pore series closely approxi- 
mated medially (separated by four scales or less); preanal scales smooth 


ATUL TNA ee ae eee to ree Ne Uae ee eee cc et ee ee ore eee et rae ee ee 9 
9. Males uniform white below; anterior section of frontal longitudinally divided; 
femoral pore series widely separated medially____-_- chrysostictus (p. 133) 


Males with lateral belly patches distinct or, if not, anterior section of frontal 
CTL Cee ee eee Senet ne eae See See een RR em Sebel on Cee ey te 10 


106 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


10. 


11. 


12. 


13. 


14. 


15. 


16: 


17 


18. 


195 


20. 


21. 


22. 


23. 


Femoral pore series closely approximated medially; two postrostrals; lateral 
scale rows parallel or nearly so; scales on posterior surface of thigh granular; 
SMA SPCClOS. aa Se eee ee scalaris (p. 136) 

Femoral pore series widely separated medially, or, if closely approximated, 
scales on posterior surface of thigh not granular; lateral scale rows strongly 


divergent; typically four postrostrals, sometimes three, rarely two__-_ 11 
Scales .on posterior, surface of thigh granular_—_—---—--_-- 12 
Seales on posterior surface of thigh not granular____-____-__-_-_-------_- 15 
Dorsal scales highly irregular in size.._.__----.-------- grammicus (p. 119) 
OTS SCA Sy SU oe CA NL AN PD STZ, Ce ee ag ee 13 


Lateral nuchal scales much smaller than and well differentiated from dorsal 
nuchal seales; scales of lateral row of dorsal nuchals enlarged, strongly 
keeledvand mucronate: 224 has a ee ee ee grammicus (p. 119) 

Lateral nuchal seales not well differentiated from dorsal nuchal scales____ 14 

Throat light blue, barred or mottled with white; lateral belly patches confluent 
medially; preanal scales large, about seven in a row from a line between 


FEIN OLA GO OLEsSCEIES mb Oyen LU Ss eemesea ene ee eres eee grammicus (p. 119) 
Throat not as described or, if so, lateral belly patches not confluent medially 
and preanal scales smaller. = 2229252 besser eee eee 15 


Seales on posterior surface of thigh granular, femoral pore series widely 
separated medially, the folds defining the preanal area passing between the 
femoral pore series; throat mottled with blue and white, never with a 
distinct, blue spot medially or on either side posteriorly_graciosus (p. 118) 

Seales on posterior surface of thigh not granular or, if so, femoral pore 
series more closely approximated medially, the folds defining preanal area 
not or barely passing between femoral pore series___-.---....--_--- 16 

Dorsal, ventral, and lateral scales subequal in size; small species, 

megalepidurus (p. 121) 

Dorsal, ventral, and lateral scales distinctly differing in size; large or small 


SPOCless awa Ss bee ee ol cee ee pe eS ee Be eee 17 
A distinct, dark, light-bordered nuchal collar____.-__-_- torquatus (p. 121) 
No distinct, dark, light-bordered nuchal collar_-_.-....._..-.-_____--- 18 
Gular region mottled, not barred nor with a median or two lateral blue spots 

DOSLeTIONy Seana a feo eee! oe se eens oe torquatus (p. 121) 
Gularresioninotrmptileds=: 2 As 825 eka 8.99 i ee 19 
Gular region barred in males and lacking a median posterior dark blue or 

plac kas pot ie == Sea ss eae ee ee ee spinosus (p. 110) 
Gular region not barred; or, if barred, with a median posterior dark blue or 

TROIS PO Gee a ce Be Se Le SD 20 


Gular region barred in males, with a median posterior dark blue or black spot. 
undulatus (p. 117) 

Guilar-regionnot barred2 225 = 22e ss wee ee en eee 2 
Supraoculars large, separated from superciliaries by no more than one com- 
plete and one incomplete row of small scales; and one or more of the posterior 
supraoculars in contact with median head scales__.__._._._._._------- 28 
Supraoculars large or small; if any of the posterior supraoculars are in con- 
tact with median head scales, the supraoculars are separated from super- 
ciliaries by three or more complete or incomplete rows of small scales 22 


Males lacking lateral belly patches____.___._..---------- undulatus (p. 117) 
Males with lateral sbelly, patehess yo ee arse are ee ee eee 23 
A pair of dark blue or black spots on posterior part of gular region. 


undulatus (p. 117) 
No pair of dark spots on posterior part of gular region_-_______-_----- 24 


CHECKLIST OF REPTILES OF MEXICO 107 


24. Three or more rows of small scales between supraoculars and superciliaries; 
or, if two, the scales of inner row considerably larger than those of outer 


TO Wetter sewn sae a ee ee eile a eae eee eee ee 25 

One complete and one incomplete row of small, subequal scales between 
supravculars and’ superciliariess! {1200 §s05h) west SA ak eee 27 

25. One canthal, or, if two, the first forced above canthal ridge_formosus (p. 107) 
Two canthals, the first not forced above canthal ridge_______________-_ 26 

26. A broad, black nuchal collar complete about neck dorsally, or anterior section 
of frontallongitudinally divideda=== 2-22-22. 5-22 55 55— formosus (p. 107) 
INeigheraUnie= se ae mes tee aes ee a eee ee © undulatus (p. 117) 
Smee renter TIC PCR DTCSCM boa. oe eS Se ee ee ee formosus (p. 107) 
BrOnvaenrid ges) vbSenite sees ee ees ee See Se en yale eae spinosus (p. 110) 

28. Adult males a uniform bright green above_______________ formosus (p. 107) 
SING Gh Ss ese ee SS. sen ep eh re es Ne spinosus (p. 110) 


FORMOSUS GROUP 


Species.—Six, with a total of 11 forms. Extralimital are lunaei, m. 
malachiticus, and m. smaragdinus.” 
Range.—Michoacan and Veracruz south to western Panama. 


KEY TO MEXICAN SPECIES OF THE FORMOSUS GROUP OF SCELOPORUS 


1. Median frontonasal separated from lateral frontonasals________________- 2 
Median frontonasal in contact with lateral frontonasals________________-_ 4 
2. Dorsals about 31; internasals large, keeled, 3 from rostral to median fron- 
LOTS Fa merece | LenS ene get Sr ema BE prezygus 7 (p. 109) 
Dorsals 37 to 45; internasals smaller, not keeled, rugose or not__-___-____-_ 3 

3. A complete nuchal collar, sometimes narrowly interrupted medially. 
malachiticus salvini (p. 108) 
Nuchal collar incomplete________-_--- malachiticus taeniocnemis (p. 107) 
4, Anterior section of frontal usually longitudinally divided; canthals sharply 
ridged; no nuchal collar; ventrals a fourth, laterals not more than two-thirds, 
SIZGIOL GOrsals. inj an WF nie Fag eet) ey ee Ve AY: asper (p. 109) 
Anterior section of frontal rarely longitudinally divided; canthals rounded; 
a nuchal collar or not; ventrals a half, laterals three-fourths size of dorsals_ 5 
5. A broad, nearly or quite complete, nuchal collar____........_.__..---.---- 6 
Collar, u' present, resiricved to sides of Meck. 2.22722. 2°25 oes <2 ic 
6. Femoral pores 12-16; one or more supraoculars generally in contact with 
median head/seales_ 2-2. 2us2f24_ 55522. malachiticus acanthinus (p. 108) 
Femoral pores 16-21; supraoculars not in contact with median head scales. 
stejnegeri (p. 109) 
7. Males without yellow on throat; dorsal scale rows black edged; dorsal head 


scales with a light blue center_________-___- formosus scitulus (p. 108) 
Males with yellow or orange on throat; dorsal surface uniform blue; head 
scalesimotaicht spoued asso — === ee formosus formosus (p. 108) 


SCELOPORUS MALACHITICUS TAENIOCNEMIS Cope 


Sceloporus taeniocnemis Copz, Proc. Amer. Philos. Soc., vol. 22, 1885, p. 399. 
Sceloporus malachiticus taeniocnemis, SmitH, Proc. U. 8. Nat. Mus., vol. 92, 1942, 
p. 356; Journ. Washington Acad. Sci., vol. 39, 1949, pp. 39, 40. 


72 See Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, p. 356, for a key to these forms. 
72a Material examined since this key was originally prepared places this species in the torquatus group. 
861316—50——_8 


108 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type.—U.S.N.M. No. 24768. 

Type locality Guatemala, here restricted to Cob4n. 

Range.—Highlands of Chiapas and northern Guatemala in Alta 
Verapaz. Known in Mexico only from Chiapas: Pinobete, Mount 
Ovando, Teopisca. 


SCELOPORUS MALACHITICUS SALVINI Giinther 


Sceloporus salvint GUNTHER, Biologia Centrali-Americana, Reptilia and Batra- 
chia, 1890, p. 68.—Smirn, Proc. Biol. Soc. Washington, vol. 49, 1936, p. 95; 
Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 55-58, pl. 3. 

Sceloporus malachiticus salvinii, Smiru, Proc. U. 8S. Nat. Mus., vol. 92, 1942, p. 
356; Journ. Washington Acad. Sci., vol. 39, 1949, p. 39. 


Type.—Brit. Mus. Nat. Hist. cotypes: Jalapa, Hoege collector; 
Guatemala, Salvin collector. Lectotype specimen 0 in Boulenger, 
Proc. Zool. Soc. London, 1897, p. 503. 

Type locality—Jalapa and Guatemala.” Restricted (Smith, 
1939, p. 40) to Jalapa, Veracruz. 

Range.—Central western Veracruz to the Isthmus of Tehuantepec. 
Recorded only from the states of Veracruz and Oaxaca. 


SCELOPORUS MALACHITICUS ACANTHINUS Bocourt 


Sceloporus acanthinus Bocourt, Ann. Sci. Nat. Zool., ser. 5, vol. 17, No. 6, 1873, 
p. [24] (acathhinus); No. 10, 1873, p. 1; Mission scientifique au Mexique 
. . . , Etudes sur les reptiles, livr. 3, 1874, pp. 180-181, pl. 18, figs. 10, 10a, 
10b, pl. 19, figs. 4, 4a.—Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 
1939, pp. 74-78, pls. 6, 7. 

Sceloporus malachiticus acanthinus, SMirn, Journ. Washington Acad. Sci., vol. 39, 
1949, pp. 39-40. 

Sceloporus guentheri SresNEGER, Proc. Biol. Soc. Washington, vol. 31, 1918, p. 92 
(type, Brit. Mus. Nat. Hist.; ‘‘Mexico,’”’ here restricted to La Esperanza, 
Chiapas). 

Type.—Mus. Hist. Nat. Paris, male and female cotypes; Commis- 
sion Scientifique collector. 
Type locality —San Agustin, near Volcan de Atitlin, Guatemala, 

610 meters. 

Range.—Pacific coast foothills, Chiapas to El] Salvador. Known 
in Mexico only from the vicinity of Escuintla, Chiapas. 


SCELOPORUS FORMOSUS FORMOSUS Wiegmann 


Sceloporus formosus WIEGMANN, Herpetologia Mexicana, 1834, p. 50, pl. 7. 

Tropidolepis formosus, DumfrRiL and Bisron, Erpétologie générale, vol. 4, 1837, 
pp. 303-304. 

Sceloporus formosus formosus, WETTSTEIN, Sitzb. Akad. Wiss. Wien, math. nat. 
Kl., vol. 148, Abt. 1, 1934, p. 25.—Smiru, Publ. Field Mus. Nat. Hist., zool. 
ser., vol. 26, 1939, pp. 34-41 (part). 

Sceloporus viviparus Copz, Proc. Amer. Philos Soc., vol. 22, 1885, p. 398 (Mirador, 
Veracruz; type, U.S.N.M. No. 25073). 


CHECKLIST OF REPTILES OF MEXICO 109 


Type.—? Zool. Mus. Berlin; F. Deppe collector. 

Type locality Mexico, here restricted to Acultzingo, Veracruz. 

Range.—Central Veracruz south in mountainous regions to the 
Isthmus of Tehuantepec. Recorded from various localities in the 
states of Veracruz, Puebla, and Oaxaca. 


SCELOPORUS FORMOSUS SCITULUS Smith 


Sceloporus formosus scitulus SmiruH, Proc. U. 8. Nat. Mus., vol. 92, 1942, pp. 
352-354, 
Type-—EHT-HMS No. 26962; R. C. Taylor and E. H. Taylor 
collectors. 
Type locality—Omilteme, Guerrero. 
Range.—Sierra Madre del Sur Mountains in Guerrero. 


SCELOPORUS ASPER Boulenger 


Sceloporus asper BOULENGER, Proc. Zool. Soc. London, 1897, pp. 476, 497, 498, 
pl. 33 (colored).—Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 51-55. 
Sceloporus obscurus VAN DENBURGH, Proc. Acad. Nat. Sci. Philadelphia, 1897 
(1898), p. 462 (Tepic, Nayarit; type, California Acad. Sci. No. 3213). 
Type.—Brit. Mus. Nat. Hist. 
Type locality—lLa Cumbre de los Arrastrados, Jalisco. 
Range.—Mountainous regions of western Mexico, from Nayarit to 
Guerrero. Recorded from Nayarit, Jalisco, Michoacan, and Guer- 


rero.”? 
SCELOPORUS STEJNEGERI Smith 


Sceloporus stejnegert SmitTuH, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 350-352. 


Type —U.S.N.M. No. 112635; E. H. Taylor, R. C. Taylor, and 
H. M. Smith collectors. 

Type locality.—Tierra Colorada, Guerrero. 

Range.—Guerrero, on southern low slopes of the Sierra Madre del 


Sur. 
SCELOPORUS PREZYGUS Smith “4 


Sceloporus prezygus SmitTu, Proc. U. 8. Nat. Mus., vol. 92, 1942, pp. 354-355. 


Type —U.S.N.M. No. 46881; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality—Conjab, 5,300 feet (between San Bartolomé and 
Comitaén), Chiapas. 

Range.—Known only from the type locality in Chiapas. 


78 The record from this state (Chilapa) (Ahl, Zool. Anz., vol. 106, 1934, p. 184) is open to doubt. The iden- 
tification needs verification. 

™ Group allocation of this species was not certain until after this account was in type. It belongs in the 
torquatus group. 


110 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 
SPINOSUS GROUP 


Species—Nine, with a total of 21 forms. 
Range.—Guatemala northward to southern Nevada, southwestern 
Colorado, southern Oklahoma, and central Texas. 


KEY TO THE SPINOSUS GROUP OF SCELOPORUS 


iz Hemorall poresion; both thighs) totaly/“ormonel ss sae ee ee 3 
Kemoral pores’ on: both. thighs total.Gior less". 122k bas See ae 2 

2. Ventral surfaces immaculate in adults of both sexes. 
horridus albiventris (p. 116) 
Sides of belly blue, frequently dark-bordered, in males; gular region barred 
in males, sometimes in females_____________ horridus oligoporus (p. 116) 
3. Supraoculars completely in contact with superciliaries, or with one to four 
very small scales between; posterior superciliary greatly enlarged; venter 
immaculate im cboOth sexes -- =. aa edwardtaylori (p. 112) 
Supraoculars separated from superciliaries by at least one row of small scales; 
posterior superciliary not enlarged; venter with distinct markings in 


a 2] Ss BOS Pah Re eee Sees re ee reste NS Ay AR ee Oe ee Sree ee 4 

4. Femoral pores on both thighs total 12 or less; 4 supraoculars, the fourth in 
contact with median head scales______------- horridus horridus (p. 116) 
Femoral pores on both thighs total 13 or more; or, if 12, all supraoculars 
Sseparatedtirom medianvhesd scalegs =. 42-5 ee eee eee 5 

5. BHemoral spores; (one: side)) liv, ors mores. = oe ee eee eae ee 6 
Hemoraljporesa(Onelside) alGorlessie 2 a ee ee ee eee een 10 

6. A black shoulder patch present, with a light posterior border___________ 10 


No black shoulder patch; or, if present, without a light posterior border__ 7 

7. Snout black; a broad, black bar across head at middle of supraocular region; 
chest and middle of belly orange in males; gular region in females white, 
with irregular, black marks; first canthal not in contact with lorilabials; 
vientralisealeés yd 4 hae sgh ts eer ere ne a 8 
Snout not black, similar in color to rest of head; no black bar across head; 
chest and middle of belly not orange; gular region in both sexes bluish, with 
light lines following scale rows at anterior part of throat; first canthal in 
contact with lorilabials; ventral scales 38—42_.magister lineatulus (p. 115) 

8. Femoral pores usually (84 percent) 20 or less on each side; total pore counts 
usually. (SOspercent)i40ror less2e) ee ee ee 9 
Femoral pores usually (78 percent) 21 or more on each side; total pore counts 
usually (83 percent) 41 or more; lateral belly patches usually not confluent 


mediallyeinumaless. ales eee melanorhinus melanorhinus (p. 112) 

9. Lateral nuchal pocket greatly reduced, with very few if any granules, and very 
little if any bare skin, shallow___________ melanorhinus stuarti (p. 113) 
Lateral nuchal pocket with numerous granules, considerable bare skin, and 
considerableidepthess see a= ae melanorhinus calligaster (p. 113) 

10. One or more supraoculars in contact with median head scales_________- 14 
None of supraoculars in contact with median head scales_____________-_ 11 

11. Gularcregionubarred= 2 22 locien eres are ayer tes tek pee Fe ole a 12 
Gular region mot barred: 5 45.6 ij oe ea SS Rae Se eee 13 


12. Dorsal scales usually more than 30; femoral pores usually more than 9 (one 
side); supraoculars usually 5 to 7___spinosus caeruleopunctatus (p. 116) 
Dorsal scales usually 30 or less; femoral pores usually 9 or less (one side); 
Supraccularsmisuallys 4a = == = eee ee ee spinosus spinosus (p. 116) 


13. 


14. 


15. 


16. 


ive 


18. 


ro: 


20. 


21. 


22. 


23. 


24, 


25. 


CHECKLIST OF REPTILES OF MEXICO 11] 


Belly patches in males continuous across chest; prefrontals never in contact; no 
dorsolateral light lines in males; posterior surface of thigh mottled or with 
a, broad Jightalineidark=-bordereds {25226 45 a2 SB ieee eae ee ee 14 
Belly patches small, confined to sides of belly in males; prefrontals usually in 
contact; dorsolateral light lines present in males; posterior surface of thigh 
nearly immaculate or with a narrow, short black line near insertion of hind 


Ne pak meh ce Mee byes al eee ucnien wey iiy dl into biped Be eit a olivaceus (p. 117) 
Gulsr region. barred. - 15 -e4! write) ce 088 aU hie od pepe tl ag be oy pga acy 9s 15 
Gilariresion«nobtbarred =o: 32! tela inte Doom ly ween Ath lyf 16 
Femoral pores 13 or more (one side) _____________-_-- orcutti licki (p. 114) 
Femoral pores 12 or less (one side).______--_--- spinosus spinosus (p. 116) 
Hemoralipores: (onejside) Wl 2ior lesseaee sone eee: Werte sae Biel 2 17 
Hemoralspores)(one;side)» 13, or-mores2.—- - Sin Seance ~ 21 


Outer row of labiomental scales in contact with mental. 
magister magister (p. 114) 


Outer row of labiomental scales separated from mental_______________- 18 
First canthal in contact with lorilabials___.________ orcutti orcutti (p. 114) 
hirsiacanthaliseparated inom lortlabialssesss 92 = eee eae ee 19 


Lateral scales about one-half size of dorsal scales; median frontonasal not in 
contact with frontal; lateral belly patches in males not passing over 
GHeS besee ee Fk es ys tern a Pes FT eR pees AN eee td eet 21 

Lateral scales considerably more than one-half size of dorsal scales; median 
frontonasal usually in contact with frontal; lateral belly patches in males 
DASSIT PAOV CLs CHES Geer oe os Bere eee er eee ta ee ee 20 


Dorsal markings absent or very dimly visible; a broad, light line on posterior 
SUTPACCLOM hI NS yok een ee lundelli gaigeae (p. 112) 
Dorsal markings present and distinct at least in females, consisting of narrow, 

dark cross bars; posterior surface of thigh mottled. 
lundelli lundelli (p. 112) 


Remoralspores (one side), .0) oF lesse 2 2 3-2 Jobe 5 eee oe eee Le 22 
Bemoral pores! (one side) *LINOrimores 3222. 45254. ee ge ee 23 
First canthal rarely in contact with lorilabials; gular region never barred; 

outer row of labiomentals rarely in contact with mental. 
clarkii boulengeri (p. 113) 


First canthal usually in contact with lorilabials; gular region barred, or outer 
row of labiomentals sometimes in contact with mental (not in o. orcut- 
ta) 5 ee ee Ot EE eee SE 8 ete ee Nye at Beh care BE 23 


First canthal separated from labiomentals; femoral pores 16 or less; lower fore- 
legs distinctly barred; gular region never barred___clarkii clarkii (p. 113) 
First canthal in contact with lorilabials, lower forelegs not barred dis- 
ITC Uy Seer a ra ee a Ss Hed SA RENE, TF Pe ee. oe A ee 24 
Outer row of labiomentals in contact with mental; gular region not barred; 
Shoulder patch GIshimet ome ok ue WE ee oe Fe eee 26 
Outer row of labiomental scales separated from mental; gular region barred or 
SHOUIer PALGIUNGIShNCh= ee coe ek ee ere ee a ee ee 25 
Dorsal scales 32 to 40; femoral pores 13-19; usually one or more scales in con- 
tact with both subocular and supralabials; a distinct, light-bordered black 
shoulder patch; dorsolateral light lines present, indistinct posteriorly, the 
medial edge sometimes indistinet_____._._________- orcutti licki (p. 114) 
Dorsal scales 28 to 36; femoral pores 10-15; usually 2 complete rows of lorila- 
bials below subocular; shoulder patch indistinct; no dorsolateral light 
pan yo Se orcutti orcutti (p. 114) 


112 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


26. Supraoculars usually 5; femoral pores (one side) usually 15 or less. 
magister magister (p. 114) 
Supraoculars usually 6 or 7; femoral pores usually 16 or more______-___- 27 
27. No dark lines in adult males on sides of lateral scale rows; femoral pores (one 
side) usually less than 19; lateral belly patches confluent medially in adult 
Malesiwet wees fart) ats ee ee Le, magister rufidorsum (p. 115) 
Dark lines on sides of lateral scale rows present in adult males; lateral belly 
patches not confluent medially in adult males___-___-___-__-__----- 28 
28. A distinct, narrow dorsal stripe about 144 scale rows wide; females and young 
with 2 rows of dark spots, one on each side of middorsal stripe. 
magister monserratensis (p. 115) 
A broad, light-colored dorsal area about 6 scale rows wide; dark spots on back 
absentronundistinctaes== =e) eee magister zosteromus (p. 115) 


SCELOPORUS LUNDELLI LUNDELLI Smith 


Sceloporus lundelli lundelli SmitH, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 
1939, pp. 66-71, pl. 4. 

Type.—Univ. Michigan Mus. Zool. No. 80674; C. L. Lundell 
collector. 

Type locality.—Cohune Ridge (20 miles southeast of Benque Viejo), 
British Honduras. 

Range.—The southern portion of the Yucatan Peninsula. Recorded 
in Mexico only from Campeche. 


SCELOPORUS LUNDELLI GAIGEAE Smith 


Sceloporus lundelli gaigeae Smrru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 
1939, pp. 71-74, pl. 5. 
Type.—Chicago Nat. Hist. Mus. No. 31524; H. M. Smith collector. 
Type locality.—Meérida, Yucatan. 
Range.—Northern Yucat4n; recorded only from Yucat4n. 


SCELOPORUS EDWARDTAYLORI Smith 


Sceloporus edwardtaylori Smirn, Herpetologica, vol. 1, 1936, pp. 6-8; Publ. Field 
Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 78-82, fig. 9. 
Type.—EHT-HMS No. 8331 (EHT field No. 4221); E. H. Taylor 
and H. M. Smith collectors. 
Type locality.—Ixtepec (= San Gerénimo), Oaxaca. 
Range.—Central and southern Oaxaca. 


SCELOPORUS MELANORHINUS MELANORHINUS Bocourt 


Sceloporus melanorhinus Bocourt, Ann. Sci. Nat., Zool., ser. 6, vol. 3, No. 12, 
1876, pp. 2-4.—Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 82-87 (part). 

Sceloporus melanorhinus melanorhinus, SmirH, Proc. U. 8. Nat. Mus., vol. 92, 
1942, p. 360. 


Type —Mus. Hist. Nat. Paris, three females, one male, cotypes; 
F. Sumichrast collector. 


CHECKLIST OF REPTILES OF MEXICO £13 


Type locality—Isthmus of Tehuantepec, Oaxaca, here restricted 
to Tehuantepec (City). 
Range.—Pacific slopes of Oaxaca. 
SCELOPORUS MELANORHINUS CALLIGASTER Smith 
Sceloporus melanorhinus, Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 
1939, pp. 82-87 (part), pls. 9, 10. 
Sceloporus melanorhinus calligaster SmitruH, Proc. U. S. Nat. Mus., vol. 92, 1942, 
pp. 360-361. 
Type.—U.S.N.M. No. 112201; H. M. Smith collector. 
Type locality—Acapulco, Guerrero. 
Range.—Pacific slopes below about 3,500 feet, from Nayarit to 
Guerrero. Recorded from Nayarit, Jalisco, Colima, Michoac4n, and 


Guerrero. 
SCELOPORUS MELANORHINUS STUARTI Smith 


Sceloporus melanorhinus calligaster, SmirH, Rev. Soc. Mex. Hist. Nat., vol. 7, 
1947, p. 66. 

Sceloporus melanorhinus stuartt SmitH, Nat. Hist. Misc., No. 20, 1948, pp. 1-3. 
Type.—Univ. Michigan Mus. Zool. No. 96759; L. C. Stuart collector. 
Type locality—Finea Canibal, 3,000 feet, Huehuetenango, Guate- 

mala. 

Range.—The upper valley of the Rio Grijalva in Chiapas and Guate- 

mala. Known in Mexico only from Chiapas: Piedra Parada, 12 

miles north of Ocozucoautla. 


SCELOPORUS CLARKII CLARKII Baird and Girard 


Sceloporus clarkiti Bairp and Grrarp, Proc. Acad. Nat. Sci. Philadelphia, vol. 6, 
1852, p. 127 (part).—SrrnsJnecER, North Amer. Fauna, No. 7, 1893, pp. 
178-181, figs. la-c, pl. 1—Smirx, Handbook of lizards of the United States, 
1945, pp. 206-208, pl. 45. 

Sceloporus clarkii clarkiit, Copr, U.S. Nat. Mus. Bull. 1, 1875, pp. 49, 92 (part).— 
SmitH, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 118-128. 


Type-—U.S.N.M. No. 2940, three cotypes, of which two belong to 
S. m. magister, and one, with a white tag on its leg, is designated 
lectotype of S. clarkii; John H. Clark collector. 

Type locality —‘‘Province of Sonora’? (~southern Arizona), here 
restricted to Santa Rita Mountains. 

Range.—Southern and central Arizona (except the western part of 
the State), southwestern New Mexico, and all of Sonora except the 
extreme western and southern parts. Recorded only from the states 
of Sonora and Chihuahua, and the islands of San Pedro Nolasco and 
Tiburén. 


SCELOPORUS CLARKII BOULENGERI Stejneger 
Sceloporus boulengeri STEJNEGER, North Amer. Fauna, No. 7, 1893, p. 180, pl. 1; 
figs. 5a-c. 
Sceloporus clarkit boulengeri, Burt, Trans. Amer. Micr. Soc., vol. 54, 1935, 171-172 
(part).—Smitu, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 
128-1383, pl. 15. 


114 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type-—U.S.N.M. No. 14079, three cotypes; A. Forrer collector. 

Type locality —Presidio (about 50 miles south from Mazatlan), 
Sinaloa. 

Range.—Pacific slopes, southern Sonora south to northern Jalisco. 
Recorded from Sonora, Sinaloa, Nayarit, and Jalisco. 


SCELOPORUS ORCUTTI ORCUTTI Stejneger 


Sceloporus orcutti StesnEcER, North Amer. Fauna, No. 7, 1893, p. 181, pl. 1, 
figs. 4a-c. 

Sceloporus orcutti orcuttt, SmirH, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 
1939, pp. 133-140, fig. 12. 

Sceloporus diguett Mocquarp, Nouv. Arch. Mus. Hist. Nat. Paris, ser. 4, vol. 1, 
1899, pp. 311-318, pl. 12, fig. 2-2b (Mus. Hist. Nat. Paris; Santa Rosalia, 
Baja California). 

Type.-—U.S.N.M. No. 16330; C. R. Orcutt collector. 

Type locality—Milquatay Valley, San Diego County, Calif. 

Range.—California, from southern San Bernardino county south- 
ward into Baja California to the Sierra de la Gigante, and on some 
adjacent islands. 


SCELOPORUS ORCUTTI LICKI Van Denburgh 


Sceloporus licki VAN DENBURGH, Proc. California Acad. Sci., ser. 2, vol. 5, 1895, 
pp. 79, 110-114, pl. 10. 

Sceloporus orcutti licki, Smirn, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 
1939, pp. 140-145, pl. 4. 


Type.—Stanford Univ. Mus. No. 2987a, lectotype (one of two 
paratypes; type in California Acad. Sci. destroyed in 1906); Gustav 
Eisen, and Vaslet collectors. 

Type locality —Sierra San Lazaro, Baja California. 

Range.—Cape region of Baja California, including Isla Espiritu 
Santo and Isla Partida. 


SCELOPORUS MAGISTER MAGISTER Hallowell 


Sceloporus magister HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, vol. 7, 1854, 
p. 938.—StTEJNEGER, North Amer. Fauna, No. 7, 1893, pp. 160, 178-188, 
pl. 1, figs. 2, a-c. 

Sceloporus magister magister, LINSDALE, Univ. California Publ. Zool., vol. 38, 
1932, p. 365.—Smitu, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 145-161, pl. 15. 


Type.—U.S.N.M. No. 2967; A. L. Heermann collector. 

Type locality —Fort Yuma, Ariz. 

Range.—Extreme western Texas to southern California (excluding 
western slopes), north to southern Nevada and southwestern Colorado, 
south to northern Durango and southern Sonora. Reported from 
Chihuahua, Coahuila, Durango, Sonora, and Baja California. 


CHECKLIST OF REPTILES OF MEXICO 115 
SCELOPORUS MAGISTER RUFIDORSUM Yarrow 


Sceloporus rufidorsum Yarrow, Proc. U.S. Nat. Mus., vol. 5, 1882, pp. 442-443 
(part). 

Sceloporus magister rufidorsum, LINSDALE, Univ. California Publ. Zool., vol. 38, 
1932, p. 366.—Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 161-165. 


Type.—U.S.N.M. No. 11981; L. Belding collector. 

Type locality —‘San Quentin Bay,” Baja California. In Yarrow, 
U.S. Nat. Mus. Bull. 24, 1882, p. 64, the locality is listed ‘‘La Paz, 
California.” 

Range.—Northern Baja California, exclusive of the northeastern 
section, southward to include the Vizcaino Desert and Cedros Island; 
Coronados Islands. 


SCELOPORUS MAGISTER LINEATULUS Dickerson 


Sceloporus lineatulus Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, pp. 
467-468. 

Sceloporus magister lineatulus, Smita, Publ. Field Mus. Nat. Hist., zool. ser., vol. 
26, 1939, pp. 168-170, fig. 13. 


Type—U.S.N.M. No. 64263; C. H. Townsend collector. 


Type locality—Santa Catalina Island, Gulf of California. 
Range.—Known only from Santa Catalina Island. 


SCELOPORUS MAGISTER MONSERRATENSIS Van Denburgh and Slevin 


Sceloporus monserratensis VAN DENBURGH and SteEvin, Proc. California Acad. 
Sci., ser. 4, vol. 11, 1921, p. 396. 

Sceloporus magister monserratensis, SMirH, Publ. Field Mus. Nat. Hist., zool. ser., 
vol. 26, 1989, pp. 165-168. 

Sceloporus magister rufidorsum, LInspALE, Univ. California Publ. Zool., vol. 38, 
1932, p. 366 (part). 


Type.—California Acad. Sci. No. 50509; Joseph R. Slevin collector. 

Type locality —Monserrate Island, Gulf of California. 

Range.—Southern edge of the Vizcaino Desert in Baja California 
southward to the southern end of the Sierra de la Gigante; islands 
adjacent to the coast in the gulf except Santa Catalina. 


SCELOPORUS MAGISTER ZOSTEROMUS Cope 


Sceloporus zosteromus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1863, p. 105.— 
STEJNEGER, North Amer. Fauna, No. 7, 1893, pp. 178, 181, pl. 1, fig. 3, a-e. 

Sceloporus magister zosteromus, LINSDALE, Univ. California Publ. Zool., vol. 38, 
1932, p. 366.—SmitH, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 170-172. 


Type.—U.S.N.M. Nos. 5298 (23 specimens), 69472-69488; John 
Xantus collector. 

Type locality —Cape San Lucas, Baja California. 

Range.—Cape region of Baja California. 


116 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 
SCELOPORUS HORRIDUS HORRIDUS Wiegmann 


Sceloporus horridus WirGMaANnn, Herpetologia Mexicana, 1834, p. 50. 
Sceloporus horridus horridus, Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 
1939, pp. 98-106, pl. 11. 
Type.—Zool. Mus. Berlin; F. Deppe collector. 
Type locality —Mexico, here restricted to Cuernavaca, Morelos. 
Range.—Known in southern Morelos, Guerrero, and Oaxaca in the 
Balsas Basin, and southern Puebla. 


SCELOPORUS HORRIDUS OLIGOPORUS Cope 


Sceloporus oligoporus Corn, Proc. Acad. Nat. Sci. Philadelphia, 1864, pp. 177-178. 
Sceloporus horridus oligoporus, TAyLor, Univ. Kansas Sci. Bull., vol. 24, 1938, 
p. 520.—Smirn, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 
106-108. 
Type.—U.S.N.M. Nos. 31386-31393, cotypes. 
Type locality —Colima, Colima. 
Range.-—Known from western Guerrero, central and southern 
Michoacan, Colima, and central Jalisco to Durango. 


SCELOPORUS HORRIDUS ALBIVENTRIS Smith 


Sceloporus horridus albiventris Smiru, Publ. Field Mus. Nat. Hist., zool. ser., 
vol. 26, 1939, pp. 108-110, pl. 12. 
Type-—EHT-HMS No. 8519; E. H. Taylor collector. 
Type locality —Tepic, Nayarit. 
Range.—Known from coastal regions from northern Jalisco through 
Nayarit to Sinaloa. 


SCELOPORUS SPINOSUS SPINOSUS Wiegmann 


Sceloporus spinosus WIEGMANN, Isis von Oken, vol. 21, 1828, p. 370; Herpetologia 
Mexicana, 1834, p. 50, pl. 7, fig. 3. 

Sceloporus spinosus spinosus, Martin peut Campo, Anal. Inst. Biol. Mexico, 
vol. 8, 1937, p. 262.—Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 
1939, pp. 87-94, pl. 11. 

Type.—? Zool. Mus. Berlin; F. Deppe collector. 
Type locality —Mexico, here restricted to Puebla, Puebla. 
Range.—Durango and western Tamaulipas to northern Jalisco, 

Michoacaén, Hidalgo, and Puebla on the Mexican Plateau. Recorded 

from Durango, Zacatecas, San Luis Potosi, Tamaulipas, Jalisco, 

Aguascalientes, Guanajuato, Hidalgo, Veracruz, México, Michoacan, 

Puebla, and Distrito Federal. 


SCELOPORUS SPINOSUS CAERULEOPUNCTATUS Smith 


Sceloporus spinosus caeruleopunctatus Smita, Univ. Kansas Sci. Bull., vol. 245 
1936 (1988), pp. 469-473; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26; 
1939, pp. 94-98, fig. 10. 


Type.—EKHT-HMS No. 8467; H. M. Smith collector. 


CHECKLIST OF REPTILES OF MEXICO U7 


Type locality —Slopes of Cerro de San Luis, about 15 miles north of 
Oaxaca, Oaxaca. 
Range.—Known only from the highlands of Oaxaca. 


SCELOPORUS OLIVACEUS Smith 


Sceloporus spinosus floridanus, STEJNEGER, North Amer. Fauna, No. 7, 1893, 
p. 181 (nec Baird). 
Sceloporus olivaceus SmiruH, Trans. Kansas Acad. Sci., vol. 37, 1934, pp. 263, 
277-279; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 110-118, 
pls. 13-14; Univ. Kansas Sci. Bull., vol. 31, 1946, pp. 103-106. 
Type-—EHT-HMS No. 2508; E. H. Taylor and John S. Wright 
collectors. 
Type locality —Arroyo Los Olmos, 3 miles southeast of Rio Grande 
City, Starr County, Tex. 
Range.—Extreme south-central Oklahoma, southward through 
central Texas to southern Tamaulipas, central Nuevo Leén, south- 
eastern Coahuila, and San Luis Potosi. Recorded from each state 


named. 
UNDULATUS GROUP 


Species.—Four, with a total of 13 forms. Extralimital are 9 forms 
all restricted to the United States and Mexico. 

Range.—British Columbia and southern New York south to 
northern Baja California and central Zacatecas. 


KEY TO MEXICAN SPECIES OF THE UNDULATUS GROUP OF SCELOPORUS 


1. Supraoculars large, entire; femoral pore series separated by 9 or more scales; 


gular patches small, widely separated____..__.__--_-------- cautus (p. 117) 
Supraoculars smaller, divided; femoral pore series separated by 8 or fewer 
scales, or, gular patches large, often covering entire throat__-_--------- 2 


2. Femoral pore series separated by 9 scales or more; scales on posterior surface 
of thigh abruptly differentiated from dorsal scales of same member, the 
median posterior scales not distinctly larger than adjacent lateral posterior 
SC alesse S11 Sr ee ee eA eee ee occidentalis biseriatus (p. 118) 

Femoral pore series separated by 8 scales or less; scales on posterior surface of 
thigh gradually merging with larger dorsals of same member, at least the 
median posterior scales distinctly larger than the adjacent lateral posterior 
Remon sees =. Foss Meee. A ae b= bg 7 ae Pe I ae 3 

3. Males without lateral belly patches; dorsolateral and lateral light stripes very 
plearivsdetined =. « 1,252 Boe Ao halk: See undulatus virgatus (p. 118) 

Males with lateral belly patches; dorsolateral and lateral light stripes poorly 
Gelined 23st ee ee ee ee undulatus consobrinus (p. 118) 


SCELOPORUS CAUTUS Smith 


Sceloporus cautus Smiru, Occ. Pap. Mus. Zool. Univ. Michigan, No. 387, 1938, 
pp. 2-7; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 173-174. 


Type—EHT-HMS No. 13027; E. H. Taylor and H. M. Smith 
collectors. 


118 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type locality —Thirty miles north of El Salado, San Luis Potosf, 
in the state of Coahuila, Mexico. 
Range.—Southern Coahuila, northern San Luis Potosi, and eastern 
Zacatecas. 
SCELOPORUS OCCIDENTALIS BISERIATUS Hallowell 


Sceloporus bi-seriatus HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, vol. 7; 
1854, pp. 93-94. 

Sceloporus occidentalis bi-seriatus, Camp, Univ. California Publ. Zool., vol. 17, 
1916, p. 65. 

Type.—Acad. Nat. Sci. Philadelphia Nos. 8474-6, cotypes. 

Type locality—Borders of El Paso Creek and Tejon Valley,” 
Calif., here restricted to El Paso Creek. 

Range.—Southern Idaho, southwest through Nevada and western 
Utah into California and the San Pedro MArtir range in Baja Cali- 
fornia. 

SCELOPORUS UNDULATUS CONSOBRINUS Baird and Girard 


Sceloporus consobrinus Barrp and GiraArD, in Marcy and McClellan, Exploration 
of the Red River of Louisiana, . . ., 1853, pp. 224-226, 237, pl. 10, figs. 
5-12. 

Sceloporus undulatus consobrinus, Corn, Ann. Rep. U. S. Nat. Mus., 1898 (1900), 
p. 337, fig. 60.—Smirn, Publ. Field Mus. Nat. Hist., zool ser., vol. 26, 1939, 
pp. 175-176. 

Type.—Destroyed. 

Type locality —Beckham County, Okla., near the confluence of the 
north fork of the Red River and Suydam Creek. 

Range.—Texas except eastern fourth, southwestern Oklahoma, ex- 
treme western and southern fourth of New Mexico, southeastern 
Arizona in vicinity of San Pedro River, south in Mexico to central 
Zacatecas, central Chihuahua on the west and central Tamaulipas 
on the east. Recorded only from Chihuahua, Coahuila, Durango, 
Nuevo Leén, and Zacatecas. 


SCELOPORUS UNDULATUS VIRGATUS Smith 


Sceloporus undulatus virgatus Smirx, Occ. Pap. Mus. Zool. Univ. Michigan, No. 
387, 1938, pp. 11-14; Publ. Field Mus. Nat. Hist., zool ser., vol. 26, 1939, 
p. 176. 
Type——Univ. Mich. Mus. Zool. No. 81912; Berry Campbell 
collector. 
Type locality—Above Santa Maria mine, El Tigre Mountains, 
Sonora. 
Range.—Southeastern Arizona and adjacent Mexico, at high ele- 
vations. Recorded in Mexico only from Sonora and Chihuahua. 


GRACIOSUS GROUP 


Species.—One, with three subspecies, only one of which occurs in 
Mexico. 


CHECKLIST OF REPTILES OF MEXICO 119 


Range.—Montana and Washington south to extreme northwestern 
Baja California. 


SCELOPORUS GRACIOSUS VANDENBURGIANUS Cope 


Sceloporus vandenburgianus Corr, Amer. Nat., vol. 30, 1896, pp. 834-835; Ann. 
Rep. U. S. Nat. Mus. 1898 (1900), p. 390, fig. 64. 

Sceloporus graciosus vandenburgianus, Camp, Univ. California Publ. Zool., vol. 
17, 1916, p. 67. 


Type.-—U.S.N.M. No. 21931; E. A. Mearns collector. 


Type locality—Summit of the coast range, San Diego County, 
Calif. 


Range.—Mountains south from Ventura County, Calif., into 
northwestern Baja California. 


GRAMMICUS GROUP 


Species.—Two, one of which includes three subspecies. 
Range.—Extreme southern Texas and Chihuahua south to Oaxaca. 


KEY TO SPECIES OF THE GRAMMICUS GROUP OF SCELOPORUS 


1. Dorsal scales unequal; a series of enlarged scales on each side of middorsal 

line, separated from each other by small, flat scales.__heterolepis (p. 121) 

Dorsalescales;moresor less) Unif OnMMny SZC = a= ea a = = ee eee 2 

2. Scales on sides of neck not abruptly differentiated from dorsal nuchal scales; 
no enlarged series of scales on sides of neck; dorsal scales 48-66. 

grammicus grammicus (p. 119) 

Seales on sides of neck abruptly differentiated from dorsal nuchal scales; two 

series of enlarged scales on sides of neck posterior to ear; dorsal scales 

FO ee ents pay eee ad Ey ae Lee ape Ie EN ee 3 

3. Dorsal scales usually less than 70 (52-74)___grammicus disparilis (p. 120) 

Dorsal scales usually 70 or more (68-93) _grammicus microlepidotus (p. 120) 


SCELOPORUS GRAMMICUS GRAMMICUS Wiegmann 


Sceloporus grammicus WIEGMANN, Isis von Oken, vol. 21, 1828, p.370.—Smiru, 
Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 179-182, fig. 16. 
Sceloporus grammicus grammicus, SMirH and Laure, Trans. Kansas Acad. Sci., 
vol. 48, 1945, pp. 332-333. 

Sceloporus pleurostictus W1EGMANN, Isis von Oken, vol. 21, 1828, p. 370 (Zool. 
Mus. Berlin; F. Deppe collector; Mexico, here restricted to Chilpancingo, 
Guerrero). 


Sceloporus rubriventris GinrHer, Biologia Centrali-Americana, Reptilia and 
Batrachia, 1890, p. 72, pl. 32, fig. C (Omilteme, Guerrero; Brit. Mus. Nat. 
Hist.). 

Type.—Zool. Mus. Berlin; F. Deppe collector. 

Type locality —Mexico, here restricted to Chilpancingo, Guerrero. 

Range.—Southern Oaxaca (San Pedro Quiechapa, Ozolotepec) and 
the Sierra Madre del Sur in Guerrero (Omilteme, Chilpancingo, 

Tamazulapan, Iguala). 


120 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 
SCELOPORUS GRAMMICUS MICROLEPIDOTUS Wiegmann 


Sceloporus grammicus var. alpha W1EGMANN, Isis von Oken, vol. 21, 1828, p. 370. 

Sceloporus microlepidotus W1EGMANN, Herpetologia Mexicana, 1834, p. 51.— 
Bocourt, Mission scientifique au Mexique ..., Etudes sur les reptiles, 
livr. 3, 1874, pp. 194-195, pl. 18 bis, fig. 13, a—d. 

Sceloporus microlepidotus microlepidotus, SmirH, Publ. Field Mus. Nat. Hist., 
zool. ser., vol. 26, 1939, pp. 184-191 (part), fig. 17, 18. 

Sceloporus grammicus microlepidotus, SmMirH and Laure, Trans. Kansas Acad. 
Scei., vol. 48, 1945, pp. 332-333. 

Sceloporus dispar Barrp and Girarp, Proc. Acad. Nat. Sci. Philadelphia, vol. 6, 
1852, p. 127 (Veracruz, here restricted to Cruz Blanca; type lost). 

Sceloporus heterurus Corr, Proc. Acad. Nat. Sci. Philadelphia 1866 (1867), pp. 
322-323 (Mirador, Veracruz; type now lost). 

Sceloporus microlepidopterus HERRERA, La Naturaleza, ser. 2, vol. 1, 1890, p. 331 
(lapsus calamt). 

Sceloporus microlepis BOULENGER, Proc. Zool. Soc. London, 1894, p. 731 (lapsus 
calami). 


Type.—Zool. Mus. Berlin; F. Deppe, collector. 

Type locality Mexico, here restricted to México, D. F. 

Range.—Southern part of the Mexican plateau from Jalisco, 
northern Guanajuato, and northern Hidalgo south to Oaxaca. Re- 
corded from the states of Colima, Distrito Federal, Guanajuato, 
Hidalgo, Jalisco, Oaxaca (Reyes), Puebla, Veracruz, México, 
Morelos, Tlaxcala, and Michoacan. 


SCELOPORUS GRAMMICUS DISPARILIS Stejneger 


Sceloporus disparilis STEINEGER, Proc. Biol. Soc. Washington, vol. 29, 1916, pp 
227-230. 

Sceloporus microlepidotus disparilis, DuNN, Proc. Acad. Nat. Sci. Philadelphia, 
vol. 88, 1936, p. 472.—Smitu, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 
1939, pp. 191-197, fig. 18, pl. 16. 

Sceloporus grammicus disparilis, SmirH and Laurs, Trans. Kansas Acad. Sci., 
vol. 48, 1945, pp. 332-333. 

Sceloporus microlepidotus microlepidotus, Dunn, Proc. Acad. Nat. Sci. Phila- 
delphia, vol. 88, 1936, pp. 472-473, 474. 

Sceloporus pilsbryt Dunn, Proc. Acad. Nat. Sci. Philadelphia, vol. 88, 1936, pp. 
473-474 (Alvarez, San Luis Potosf; type, Acad. Nat. Sci. Philadelphia No. 
20085). 


Type—U.S.N.M. No. 33041; William Lloyd collector. 

Type locality —Lomita Ranch, 6 miles north of Hidalgo, Tex. 

Range.—Northern Hidalgo, Guanajuato, and southern Zacatecas 
northward to central Chihuahua and northern Coahuila and the 
southern tip of Texas. Recorded from Chihuahua, Coahuila, 
Durango, Guanajuato, Hidalgo, Nuevo Leén, San Luis Potosi, 
Tamaulipas, and Zacatecas. 


15 Intergrades of S. g. grammicus and S. g. microlepidotus are from Cerro San Felipe and Cerro San Juan» 
north of Oaxaca City. 


CHECKLIST OF REPTILES OF MEXICO 12h 


SCELOPORUS HETEROLEPIS Boulenger 

Sceloporus heterolepis BOULENGER, Proc. Zool. Soc. London, 1894, pp. 724, 731, 
pl. 48, fig. 4—Smitu, Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 
197-199, pl. 16. 

Type.—Brit. Mus. Nat. Hist., part of cotypes; one cotype, Mus. 
Comp. Zool. No. 32346. 

Type locality —La Cumbre de los Arrastrados (to which the type 
locality is here restricted), Real Alto,“‘Riocho (=Rancho?)” La Ber- 
beria, Sierra de Bolafios. 

Range——Known only from western Jalisco (type localities and 


Cerro Tequila.) 
MEGALEPIDURUS GROUP 


Species.—Two. 
Range.—Western central Veracruz and eastern Puebla. 


KEY TO SPECIES OF THE MEGALEPIDURUS GROUP OF SCELOPORUS 


1. Dorsal scales 44 to 56; scales between femoral pore series 2 to 5; basal sub- 
caudals keeled in females; males with distinct blue areas on sides of belly. 
pictus (p. 121) 
Dorsal scales 54 to 62; scales between series of femoral pores 4 to 8; basal sub- 
caudals smooth in females; males immaculate below. 
megalepidurus (p. 121) 
SCELOPORUS MEGALEPIDURUS Smith 
Sceloporus megalepidurus Smitu, Trans. Kansas Acad. Sci., vol. 37, 1934, pp. 
272-277, pl. 9, figs. 7, 8, pl. 10, fig. 13; Publ. Field Mus. Nat. Hist., zool. ser., 
vol. 26, 1939, pp. 204-209, fig. 20. 
Type-—EHT-HMS No. 7543 (EHT field No. 2908); E. H. Taylor 
and H. M. Smith collectors. 
Type locality — Near Totalco, Veracruz. 
Range.—Known only from northern Puebla, western Veracruz, and 


Tlaxcala (Apizaco). 
SCELOPORUS PICTUS Smith 


Sceloporus pictus Surry, Amer. Mus. Nov., No. 892, 1936, pp. 1-4; Publ. Field 
Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 200-204, pl. 17, two figs. 
Type-—Amer. Mus. Nat. Hist. No. 18744; Paul D. R. Rithling, 
collector. 
Type locality —Santa Catarina, Puebla. 
Range.—Central Puebla and central western Veracruz. 
TORQUATUS * GROUP 


Species.—Twenty-one forms, representing 10 species, are recognized. 
Range.—Arizona and Texas south to Guatemala. 


76 This group was in 1938 termed the poinsettii group, inasmuch as Sceloporus poinsettii Baird and Girard 
(1854) was the oldest name in the group after exclusion of S. torguatus Wiegmann, 1828. The latter name 
was considered unavailable because it was once (Wagler, 1830) placed in the genus Tropidurus, in 
which the combination Tropidurus torquatus (Wied, 1820) had already been made (Wied, 1824). Secondary 
homonyms, such as this, have not, however, been regarded in recent years as permanently suppressed; 
they are suppressed only as long as they actually remain homonyms. Accordingly, we revert to the 
name torquatus for this group. 


io2 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


10. 


11. 


12. 


13. 


KEY TO SPECIES OF THE TORQUATUS GROUP OF SCELOPORUS “63 


. Supraoculars in a single series, with no scale divided_____.________----_ 2 
Supraoculars in 2 series; or, if in one series, with one or more scales divided. 9 

; Femoral’ pores'S—14; dorsal scales: 25-3522 55-2. 222 oS ee ee ee 3 
Femoral] pores more than 14; or dorsal scales more than 35_______-___--- 5 

. Dorsal nuchal collar broadly interrupted medially by a space about 5 scales 
WAdess throats mo tt lec ase =e eee torquatus binocularis (p. 126) 
Dorsal nuchal collar broad, complete; throat not mottled______________ 4 

. Femoral pore counts on both sides usually total more than 21; dorsal scales 
USUalhy sole Oren OTe ss see ee a ee eee serrifer plioporus (p. 124) 
Femoral pore counts on both sides usually total less than 22; dorsal scales 
SeMeraliya less UL ey Mey) eee ee eee ere a serrifer serrifer (p. 123) 
Dorsal'scales Shor less) 2S Ae ee ee ee ee es see eee 6 
Dorsal scales * more wham ae es ye eee ee ee ee a 


. Nuchal collar divided on each side of neck, the area between lighter in color; 


dorsal color light, with dark and light spots irregularly placed; size large 
(maximum snout-vent measurement 129 mm.); dorsal scales more strongly 
keeled and mucronate__________--_--- torquatus melanogaster (p. 126) 
Nuchal collar broad, complete; dorsal color dark, without light spots; maxi- 
mum snout-vent measurement 98 mm.; dorsal scales more weakly keeled 
DIA INU OTT eh Ge eee eae ee torquatus torquatus (p. 125) 


. Lateral scales about one-half as large as median dorsals_lineolateralis (p. 126) 


Lateral scales as large as or larger than dorsals____._____._.____---_------- 8 


. Dorsal scales 40 or more; nuchal collar narrow, with light borders broken; a 


light line on side of head and another on side of neck; each dorsal scale 
usually with a light median spot___.___.___-__--- jarrovii jarrovii (p. 128) 
Dorsal scales usually less than 40; nuchal collar broad, with unbroken light 

borders; no light lines on sides of head and neck; no spots on dorsal scales. 
bulleri (p. 125) 


. Lateral scales with the terminal point arising well within the free posterior 


marpins dorsaliscales 4:1 oramorel ss 2e- =  e  e ee 10 
Lateral scales with the terminal point arising at or very near the free posterior 
margin; head scales not microscopically rugose; no oblique dark blue lines 
ON ConOatse<s eo ae ee oe eee Le ee ee eee eee en 11 
Dorsal scales 47 to 54; throat with very distinct, oblique, dark blue lines; 
head scales not microscopically rugose; oblique dark bands on sides of body 
distinct; maximum snout-vent measurement 79 mm. 
dugesii intermedius (p. 127) 
Dorsal scales 41 to 50; throat without or with very faint oblique lines; head 
scales microscopically rugose; oblique dark bands on sides of body indistinct 
or absent; maximum snout-vent measurement 87.5 mm; femoral pores not 


OVET GS cee ee 2 aie ye Senne Meyer ee see ea er ee dugesii dugesii (p. 127) 
DorsallscalesiS5vor mores 22 ee ee ornatus ornatus (p. 127) 
Morsaliscalestessithan 55 atwel be Aye he eyeie n Re ee Oe eee 12 
Dorsaliscalest4 toro 22 542 ee ee ee ornatus caeruleus (p. 127) 
IDorsalscalestless thami.4: (fete ace eo eee irene pet cree na er eee 13 


Tail with very distinct, broad, alternating dark and light bands, most distinct 
toward tip, where they are complete; supraoculars in 2 complete rows; 
head scales very irregular; light borders of nuchal collar broad; a broad light 
band across neck behind occiput; preocular usually entire; inner row of 
labiomental scales generally terminating at a point posterior to suture be- 
tween second and third infralabials____.______________-_ poinsettii (p. 125) 


7a For one other species of this group, see footnote 72a. 


14. 


15. 


16. 


17. 


18. 


19. 


20. 


21. 


CHECKLIST OF REPTILES OF MEXICO 123 


Tail without distinct alternating dark and light bands of nearly equal width; 


bands about tail not complete toward tip; dorsal scales 40 or less___-_-__ 14 
Dorsal seaies ot OF less 2e— ee ae ee eee 15 
Ore AliSeRtens So. Ol INOlG sae oe ie ee SiS oe ee ee ee 16 


A middorsal series of very large, dark blotches, sometimes fused bandlike; 
all except males with a pair of parallel, closely placed dark lines down 
middle of throat; collar 4 scales Jong or less. 

mucronatus mucronatus (p. 124) 

No middorsal series of dark blotches; parallel throat lines absent; collar 

involving 4% scale lengths or more___mucronatus omiltemanus (p. 124) 


Each dorsal scale row, in adult males, with a longitudinal light line; dorsals 
SUSCONO Orta 2 fees ree eee Rie aN Tee mucronatus aureolus (p. 124) 
Scale rows net with continuous Imes. {559 «A BB oe a 1624 


Nuchal black collar 4 or 5 scales wide, bordered anteriorly and posteriorly 
by a light band 1 or 1% scales wide; each border interrupted medially by a 
light scale; snout-vent length often more than 100 mm., reaching 143 
FOO OT a a Ne Tt ye re cyanogenys (p. 125) 

Not so; maximum snout-vent length 100 mm________________-_______ 18 


Lateral body scales distinctly decreasing in size laterally, at a point halfway 
between axilla and groin, distinctly smaller than dorsal scales______ 19 
Lateral body scales not decreasing in size laterally at least up to a point half- 
way between axilla and groin, where they are still subequal to, or even a 
littledareer than; middorsal ‘seales-—_ 2-5 225 eon = nee 20 
Dorsal scales on lower foreleg about half size of those on upper foreleg; dark 
transverse streaks in lateral belly patches; nuchal collar covering six scales 
medially;“or more. 220 ee es ae ie ee ne jarrovii sugillatus (p. 128) 
Dorsal scales on lower foreleg but little if any smaller than those on upper 
foreleg; no dark streaks in lateral belly patches; nuchal collar less than four 
scales vongeimiddorsally= 2-2 eee jarrovii immucronatus (p. 128) 


Supraoculars essentially in 1 row; if an outer row is evident, it is composed of 
scales much smaller than those of inner row, and usually number no more 


Chant woe se a eee eee 2 ee eee se jarrovii jarrovii (p. 128) 
Supraoculars in 2 rows, those of outer row a little smaller than those of inner, 
MsUslly NUM pering o.Ol MOLES <2 eo a8 ee eh he a ge 21 


Adult males black above and below, with orange areas and spots on sides of 
head, belly, and tail; only throat, underside of tail, and posterior surface of 
hind leg not black; females somewhat similar, very dark above, the collar 
poorly defined, young with poorly defined, narrow, light borders on neck 
collar; dorsal scales average 37.5__.____________- jarrovii oberon (p. 129) 

Adult males light brown above, with very broad, very well defined nuchal 
collar; a median area on belly white except in very largest males; sides of 
abdomen blue, black-edged; females and young with more distinct light 
borders on nuchal collar; dorsal scales average 40.6_jarrovii minor (p, 128) 


SCELOPORUS SERRIFER SERRIFER Cope 


Sceloporus serrifer Corn, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 124.— 


Smiru, Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), pp. 558-564, figs. 5, 6, 
pl. 47, fig. 2. 


Sceloporus serrifer serrifer, SMiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 


1939, p. 212, fig. 23. 
861316—50——_9 


124 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type—U.S.N.M. No. 34868; Arthur Schott collector. 

Type locality— Yucatan, here restricted to Mérida. 

Range.—Northern part of the Yucatan Peninsula. Recorded only 
from the state of Yucatén; intergrades with S. s. plioporus are re- 
corded from Balchacaj, Campeche. 


SCELOPORUS SERRIFER PLIOPORUS Smith 


Sceloporus serrifer plioporus SmitH, Publ. Field Mus. Nat. Hist., zool. ser., vol. 
26, 1939, pp. 212-214, pl. 18. 

Type.—Chicago Nat. Hist. Mus. No. 32004; E. H. Taylor collector. 

Type locality —Four miles east of Encero, Veracruz. 

Range.—Gulf coast of Mexico from southern Tamaulipas southward 
through the basal part of the Yucatén Peninsula to Petén, Guatemala. 
Recorded from the states of Tamaulipas, San Luis Potosi, Veracruz, 
and Tabasco (Tenosique); intergrades with S. s. serrifer are recorded 
from Balchacaj, Campeche. 


SCELOPORUS MUCRONATUS MUCRONATUS Cope 


Sceloporus torquatus mucronatus Corr, Proc. Amer. Philos. Soc., vol. 22, 1885, p. 
402. 

Sceloporus mucronatus mucronatus, SMitH, Univ. Kansas Sci. Bull., vol. 24, 1936 
(1938), pp. 583-591, figs. 10, 11, pl. 49, fig. 2; Publ. Field Mus. Nat. Hist. 
zool. ser., vol. 26, 1939, pp. 218-220, figs. 26, 27. 


Type.—U.S.N.M. Nos. 25074-9; No. 25077 designated as lectotype 
(Smith, op. cit., 1938, p. 583). 
Type locality Mirador, Veracruz. 
Range.—Parts of Hidalgo, Veracruz, Puebla, and México, at high 
elevations. 
SCELOPORUS MUCRONATUS AUREOLUS Smith 


Sceloporus mucronatus omiltemanus, SMitH, Univ. Kansas Sci. Bull., vol. 24, 1936 
(1938), pp. 591-598 (part), figs. 11, 12, pl. 50. 

Sceloporus mucronatus aureolus Smiru, Proc. U. 8. Nat. Mus., vol. 92, 1942, pp. 
356-357. 


Type.-—U.S.N.M. No. 112232; H. M. Smith collector. 


Type locality —Two miles west of Acultzingo, Veracruz. 
Range.—Western central Veracruz and southern Puebla. 


SCELOPORUS MUCRONATUS OMILTEMANUS Giinther 


Sceloporus omiltemanus GintHeER, Biologia Centrali-Americana, Reptilia and 
Batrachia, 1890, p. 66, pl. 32, fig. A. 

Sceloporus t.[orquatus] omiltemanus, Taytor, Proc. Biol. Soc. Washington, vol. 
44, 1931, p. 129. 

Sceloporus mucronatus omiltemanus, SmMirH, Univ. Kansas Sci. Bull., vol. 24, 1936 
(1938), pp. 591-598. 


Type.—Brit. Mus. Nat. Hist.; H. H. Smith collector. 


CHECKLIST OF REPTILES OF MEXICO 125 


Type locality —Omilteme, Guerrero, 8,000 feet elevation. 
Range.—Central highlands of Oaxaca and the Sierra Madre del Sur 
in Guerrero.” 
SCELOPORUS POINSETTII Baird and Girard 


Sceloporus poinsettit Barrp and Grrarp, Proc. Acad. Nat. Sci. Philadelphia, vol. 
6, 1854, pp. 126-127.—Smiru, Univ. Kansas Sci. Bull., vol. 24, 1936 (1988), 
pp. 606-617, pl. 51, fig. 1, text figs. 14, 15. 
Type-—U.S.N.M. No. 2948 cotypes, 2 specimens, and No. 2952, 2 
specimens, “‘Sonora’’; J. H. Clark collector. 
Type locality —‘Rio San Pedro of the Rio Grande del Norte, and 
the province of Sonora,” here restricted to the former locality. 
Range.—Southern New Mexico east to central Texas, south through 
western Nuevo Leén and southern Coahuila to central Durango. 
Recorded in Mexico from Chihuahua, Coahuila, Durango, and Nuevo 
Leén. 


SCELOPORUS CYANOGENYS Cope 


Sceloporus torquatus cyanogenys Corr, Proe. Amer. Philos. Soc., vol. 22, 1885, 
p. 402. 

Sceloporus cyanogenys, SmirH, Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), 
pp. 599-606, pl. 51, fig. 2, text figs. 13-14. 

Sceloporus torquatus mucronatus, BoULENGER, Catalogue of the lizards in the 
British Museum, 1885, vol. 1, p. 220 (part). 

Sceloporus torquatus poinsettii, Burt, Copeia, 1932, No. 3, p. 158 (part), 


Type.—U.S.N.M. Nos. 31373-31377, cotypes; Acad. Nat. Sci. Phila- 
delphia, 11304-11305, cotypes. 

Type locality —Monterrey, Nuevo Leén. 

Range.—Southern Texas from Devils River and Starr County to 
central Tamaulipas and Nuevo Leén. Recorded in Mexico only from 
the states of Tamaulipas and Nuevo Leén. 


SCELOPORUS BULLERI Boulenger 


Sceloporus bulleri BouLENGER, Proc. Zool. Soc. London, 1894, pp. 729-730, pl. 
48, fig. 3.—Smirn, Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), pp. 579-583, 
figs. 8, 9, pl. 49, fig. 1. 


Type.—Brit. Mus. Nat. Hist. 


Type locality —La Cumbre de los Arrastrados, Jalisco. 
Range.—Western and southern parts of Jalisco. 


SCELOPORUS TORQUATUS TORQUATUS Wiegmann 7 


Sceloporus torquatus WIEGMANN, Isis von Oken, 1828, p. 369. 

Sceloporus torquatus torquatus, Corr, Proc. Amer. Philos. Soc., vol. 22, 1885, 
pp. 402, 403.—Smiru, Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), pp. 
564-571, figs. 7, 8, pl. 48, fig. 1. 


7 A record for Chiapas (Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, p. 221) is in error; 
the specimen referred to is an S. prezygus. 
78 See footnote 76 for use of this name, 


126 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Agama torquata PEALE and GREEN, Journ. Acad. Nat. Sci. Philadelphia, vol. 6, 
1830, pp. 231-234 (type Acad. Nat. Sci. Philadelphia; Mexico, here restricted 
to México, D. F.). 

Sceloporus ferrariperezi Corr, Proc. Amer. Philos. Soc., vol. 22, 1885, p. 400 
(cotypes, U.S.N.M. Nos. 9874, 9876, 9878, 9880, 9895, and No. 9897 which 
is an S. spinosus spinosus; Guanajuato, Guanajuato, by present restriction). 

Sceloporus ferrariperezt ferrariperezt, SmirH, Publ. Field Mus. Nat. Hist., zool. 
ser., vol. 26, 1939, pp. 214-216, fig. 24. 


Type.—Zool. Mus. Berl.; F. Deppe collector. 

Type locality.—Mexico, here restricted to México, D. F. 

Range.—Central Mexico, including Hidalgo, west-central Veracruz, 
México, Distrito Federal, northern Puebla, eastern Morelos, southern 
Guanajuato, and northern Michoacén. Recorded only from the 
states cited. 


SCELOPORUS TORQUATUS BINOCULARIS Dunn 


Sceloporus binocularis DunN, Proc. Acad. Nat. Sci. Philadelphia, vol. 88, 1936, 
pp. 474-475. 

Sceloporus ferrariperezi binocularis, SmitH, Publ. Field Mus. Nat. Hist., zool. 
ser., vol. 26, 1939, pp. 216-218, pl. 18. 


Type.—Acad. Nat. Sci. Philadelphia No. 20032; Henry A. Pilsbry, 
Francis W. Pennell, and Cyril H. Harvey collectors. 

Type locality.—Trail from Pablillo to Alamar, Nuevo Le6n. 

Range.—Known only from the general region of the type locality. 


SCELOPORUS TORQUATUS MELANOGASTER Cope 


Sceloporus melanogaster Corr, Proc. Amer. Philos. Soc., vol. 22, 1885, pp. 400-401. 

Sceloporus torquatus melanogaster, BOULENGER, Catalogue of the lizards in the 
British Museum, vol. 2, 1885, p. 220.—Smitu, Univ. Kansas Sci. Bull., vol. 
24, 1936 (1938), pp. 571-579, pl. 48, fig. 2, text figs 1, 8. 

Sceloporus ferrartperezt melanogaster, SmitH, Publ. Field Mus. Nat. Hist., zool. 
ser., vol. 26, 1939, p. 216. 


Type.—U.S.N.M. No. 9877; Alfredo Dugés collector. 

Type locality.—‘‘Probably from Guanajuato’’=Noria (Michoacan), 
or TupAtaro, near Cueramaro, Guanajuato fide Dugés (La Naturaleza, 
ser. 2, vol. 1, 1887, pp. 114-115), here restricted to TupAtaro. 

Range.—Northern Jalisco, west through most of Guanajuato, north 
through central and southern San Luis Potosi and Zacatecas. Re- 
corded from Aguascalientes, Guanajuato, Jalisco, Michoacan, San 
Luis Potosi, and Zacatecas. 


SCELOPORUS LINEOLATERALIS Smith 


Sceloporus lineolateralis Smiru, Proc. Biol. Soc. Washington, vol. 49, 1936, pp. 
92-96; Univ. Kansas Sci. Bull., vol. 24, 1986 (1938), pp. 617-624, pl. 52, 
figs. 16, 17; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 225-226, 
fig. 31. 


CHECKLIST OF REPTILES OF MEXICO 127 


Type.—EHT-HMS, No. 4323; E. H. Taylor and H. M. Smith 
collectors. 

Type locality.—Six miles northeast of Pedricefia, Durango. 

Range.—Durango. 


SCELOPORUS ORNATUS ORNATUS Baird 


Sceloporus ornatus Barrp, Proc. Acad. Nat. Sci. Philadelphia, 1858 (1859), p. 254. 
Sceloporus ornatus ornatus, SmitH, Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), 
pp. 647-651, figs. 21, 22, pl. 54, fig. 1. 


Type.—U.S.N.M. No. 2845; Darius Nash Couch collector. 
Type locality —Patos, Coahuila. 
Range.—Southeastern Coahuila. 


SCELOPORUS ORNATUS CAERULEUS Smith 


Sceloporus ornatus caeruleus SmitH, Copeia, 19386 (1937), No. 4, pp. 227-230; 
Univ. Kansas Sci. Bull., vol. 24, 1936 (1938) pp. 652-657, figs. 22-23, pl. 54, 
fig. 2. 


Type-—Univ. Kansas Mus. Nat. Hist. (DHD-HMS field No. 
350); D. H. Dunkle and H. M. Smith, collectors (lost?). 

Type locality—Five miles south of San Pedro, Coahuila. 

Range.—Known only from the type locality. 


SCELOPORUS DUGESII DUGESII Bocourt 


Sceloporus dugesii Bocourt, Ann. Sci. Nat., Zool., ser. 5, vol. 17, No. 10, 1873, 
p. 2; Mission scientifique au Mexique . . ., Etudes sur les reptiles, livr. 3, 
1874, pp. 188-190, pl. 18, figs. 7-7b. 

Sceloporus dugesii dugesiit, SmitH, Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), 
pp. 657-663, figs. 22, 24, pl. 55, fig. 1. 


Type.—Mus. Hist. Nat. Paris; Alfredo Dugés collector. 

Type locality —Colima. 

Range.—Along the coastal ranges of western Mexico from southern 
Nayarit to Colima. Recorded from the states of Jalisco, Colima, 


and Nayarit. 
SCELOPORUS DUGESI INTERMEDIUS (Dugés) 


Trlopidolepis] intermedius Ducts, La Naturaleza, vol. 1, 1869, p.143 (nomen 
nudum). 

Sceloporus intermedius Ducts, La Naturaleza, vol. 4, 1877, pp. 29-34, pl. 1, 
figs. 21-32. 

Sceloporus dugesii intermedius, SmMirH, Univ. Kansas Sci. Bull., vol. 24, 1936 
(1938), pp. 663-670.—SmitH and Necker, Anal. Esc. Nac. Cien. Biol., 
vol. 3, 1948, pp. 212-214, pl. 2, figs. 2, 3. 

Scleloporus] Westphalii Ducks, La Naturaleza, vol. 4, 1877, p. 30 (substitute 
name for S. intermedius, which Dugés retained for his description of the 
species in 1877, although he says “‘Querfa haberle dado el nombre de Sec. 
Westphalii’’). 


Type.—Mus. Alf. Dugés; two cotypes; collector unknown. 


128 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type locality—La Noria, near Zamora, hacienda of D. Epifanio 
Jiménez (in Michoacan). 
Range.—Known from Guanajuato and northern and central 


Michoacan. 
SCELOPORUS JARROVII JARROVII Cope 


Sceloporus jarrovii Corn, in United States Geological and Geographical Surveys 
West of the 100th Meridian, vol. 5, 1875, p. 569. 

Sceloporus jarrovii jarrovit, Smiru, Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), 
pp. 624-631, figs. 17, 18, pl. 47, fig. 1. 


Type-—U.S.N.M. Nos. 8494, two specimens, and 8495, cotypes. 

Type locality—Southern Arizona. 

Range.—Central Arizona east to western New Mexico, and south 
through Chihuahua and western Sonora to extreme western Zacatecas 
and extreme northern Nayarit. Recorded in Mexico from Chihuahua, 
Durango, and Sonora. 


SCELOPORUS JARROVII SUGILLATUS Smith 


Sceloporus jarrovit sugillatus Smrru, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 
357-359. 


Type.—U.S.N.M. No. 112100; H. M. Smith collector. 
Type locality.—Lake No. 4, Zempoala, México. 10,000 feet 
elevation. 
Range.—High elevations in Ajusco range. Known only from the 
type locality. 
SCELOPORUS JARROVII IMMUCRONATUS Smith 


Sceloporus jarrovit immucronatus SmirH, Copeia, 1936 (1937), pp. 223-227; Univ. 
Kansas Sci. Bull., vol. 24, 1936 (1938), pp. 640-646, figs. 17, 20, pl. 53, fig. 1. 


Tupe.—EHT-HMS No. 9358A (EHT-HMS field No. 500); E. H. 
Taylor and H. M. Smith collectors. 

Type locality.—Ten miles north of El Pinalito, Hidalgo. 

Range.—Western Querétaro through Hidalgo, eastern San Luis 
Potosi, to south-central Veracruz. Recorded from each state cited. 


SCELOPORUS JARROVII MINOR Cope 


Sceloporus torquatus minor Corn, Proc. Amer. Philos. Soc., vol. 22, 1885, p. 402. 
Sceloporus jarroviit minor, SmituH, Univ. Kansas Sci. Bull., vol. 24, 1936 (1988), 
pp. 631-639, figs. 17-19, pl. 53, fig. 2. 


Type.—U.S.N.M. Nos. 26166-26167. 

Type localt-y.—Zacatecas, here restricted to Valparaiso Mountains. 

Range.—Northern Quéretero and Guanajuato through México, 
western Zacatecas, and most of San Luis Potosi, to western Nuevo 
Leén. Recorded from each state cited. 


CHECKLIST OF REPTILES OF MEXICO 129 


SCELOPORUS JARROVII OBERON Smith and Brown 


Sceloporus jarrovii oberon Smith and Brown, Publ. Field Mus. Nat. Hist., zool. 
ser., vol. 24, 1941, pp. 253-257, fig. 24. 


Type.—U.S.N.M. No. 105823; H. M. Smith and Rozella Smith, 
collectors. 

Type locality.—Arteaga, Coahuila. 

Range.—Southeastern Coahuila. Recorded only from the type 
locality, from Diamond Pass, Mount Zapalinamé, near Saltillo, and 
from Sierra Guadelupe. 

VARIABILIS GROUP 


Species.—Nine forms, representing five species, are recognized in 
this group. One form, S. variabilis olloporus Smith, is extralimital, 
occurring from Guatemala to Costa Rica. 

Range.—-Southern central Texas south to Costa Rica, entirely on 
Atlantic slopes west of the Isthmus of Tehuantepec. 


KEY TO SPECIES OF THE VARIABILIS GROUP OF SCELOPORUS 


1. Series of femoral pores separated medially by no more than 6 scales________ 2 
Series of femoral pores separated medially by 10 or more scales_________-_- 3 
2. Dorsal scales 69 to 76; scales around body 70 to 81; dorsal scale rows at nape 
SCOR D Te anes eee men ee ee eRe Snr Lee nen es eet parvus scutulatus (p. 131) 
Dorsal scales 58 to 69; scales around body 61 to 69; dorsal scale rows at nape 
LUBY SO) TES hn NO parvus parvus (p. 131) 
3. Ventral interfemoral scales separated from ventral thigh scales by a group of 
small scales one-third or one-fourth size of adjacent scales; a rudimentary 
gular fold; lateral scales much less than half size of ventral scales; dorsal 
SCAICSIOO TONS Ba ee BO ee a edd SAS 2 Ee couchii (p. 132) 
Ventral interfemoral scales more or less continuous with ventral thigh scales; 
no rudimentary gular fold; lateral scales more than one-half size of ventral 
scales dorsalsiusualliy lessythaniG9o= === see == ae eee ee ee 4 
4. Dorsal scales 36 to 47; subnasal usually absent; shank and posterior surface of 
lower foreleg distinctly banded; spots on back distinct in both sexes; preocular 
usually divided; frontoparietals usually in contact medially_teapensis (p. 130) 
Domaecaless 4cOFMOreso 2 Saas hee ee ees ee ee a ee eee 5 
5. Males and females immaculate below; femoral pores usually 9 or less on each 
side (occasionally more in females); dorsal scales 48 to 55; frontoparietals 
usually separated by an azygous scale; preocular usually entire; subnasal 
MATELY APTOS CM baa aa ar ee eae es ee ee he Se eS 6 
Males with red, blue-bordered areas on sides of abdomen; subnasal usually 
present; frontoparietals usually in contact medially; preocular usually 
GUVA Ged Sore ee ee ee ee eS Se a 
6. Postrostrals usually 2, never 4; scales around body usually 53 to 58. 

cozumelae (mainland) (p. 130) 

Postrostrals usually 4, rarely 2 or 3; scales around body usually 59 to 64. 
cozumelae (typical) (p. 130) 
(peorsalpealesshOvormore-cty sts bys ta. tae Yk eee hey, Tope, | 8 
Dorsal scales usually less than 59; dorsolateral light lines 1 and 2 half scale 
rows wide posteriorly; maximum snout-vent measurement about 74mm_ 9 


130 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


8. Dorsolateral light stripes very distinct, 2 and 2 half scale rows wide posteriorly; 
females with sides of belly marked as in males, but less distinctly; maxi- 
mum snout-vent measurement 71 mm________~_ variabilis smithi (p. 131) 

Dorsolateral light stripes not so distinct, 1 and 2 half scale rows wide posteriorly; 
spots between dorsolateral light stripes very distinct; females with sides of 
belly immaculate; maximum snout-vent measurement 53 mm. 

variabilis marmoratus (p. 131) 

9. Femoral’ pores) 2.or mores.) eens fs be Le variabilis variabilis (p. 130) 

Femoral spores: Visor, less 22. 6 se ee variabilis olloporus (p. 129) 


SCELOPORUS COZUMELAE Jones 


Sceloporus cozumelae JONES, Occ. Pap. Mus. Zool. Univ. Michigan, No. 186, 1927, 
pp. 1-4.—Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 
246-251. 


Type.—U.S.N.M. No. 13904; “U.S. Fish Commission” collector. 

Type locality —Cozumel Island, Quintana Roo. 

Range.—Northern half of Yucat&én Peninsula and adjacent Islands. 
Recorded from Cozumel and Mujeres Islands and the states of 
Yucatan and Quintana Roo. 


SCELOPORUS TEAPENSIS Giinther 


Sceloporus teapensis GUNTHER, Biologia Centrali-Americana, Reptilia and Ba- 
trachia, 1890, pp. 75-76.—Smitru, Publ. Field Mus. Nat. Hist., zool. ser., 
vol. 26, 1939, pp. 256-262, pl. 21. 


Type.—Brit. Mus. Nat. Hist., six cotypes; H. H. Smith collector. 

Type locality —Teapa, Tabasco. 

Range.—Restricted to Atlantic slopes from southern Veracruz and 
northeastern Oaxaca, eastward through Chiapas, Tabasco, and Cam- 
peche, and through the Petén region of Guatemala to British Hon- 
duras, south to Coban, Alta Verapaz. Recorded from each state 
cited in Mexico. 


SCELOPORUS VARIABILIS VARIABILIS Wiegmann 


Sceloporus variabilis W1rmaGMANN, Herpetologia Mexicana, 1834, p. 51. 

Sceloporus variabilis variabilis, Smrtu, Proc. Biol. Soc. Washington, vol. 47, 1934, 
pp. 127-129; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 262-272, 
fig. 45, pl. 22. 


Type.—? Zool. Mus. Berlin; F. Deppe collector. 

Type locality.—Mexico, here restricted to Veracruz, Veracruz. 

Range.—Southern Tamaulipas along the Atlantic coast to south- 
central Veracruz (Rio Blanco), inland to eastern Querétaro and 
Puebla; through eastern Oaxaca and western Guatemala, reaching 
the coast only in Chiapas. Recorded from each state mentioned, 
and from San Luis Potosi, Guanajuato, and Hidalgo. 


CHECKLIST OF REPTILES OF MEXICO 131 


SCELOPORUS VARIABILIS SMITHI Hartweg and Oliver 


Sceloporus variabilis smithi HartTwea and Outver, Occ. Pap. Mus. Zool., Univ. 
Michigan, No. 356, 1937, pp. 1-5.—Smiru, Field Mus. Nat. Hist., zool. ser., 
vol. 26, 1939, pp. 278-282, fig. 45, pl. 23. 
Type.—Univ. Mich. Mus. Zool. No. 81777; N. Hartweg and J. A. 
Oliver collectors. 
Type locality —Guengola Mountain, 6 kilometers northwest of the 
city of Tehuantepec, Oaxaca. 
Range.—Eastern Oaxaca. 


SCELOPORUS VARIABILIS MARMORATUS Hallowell 


Sceloporus marmoratus HaLtuoweEuL, Proc. Acad. Nat. Sci. Philadelphia, vol. 6, 
1852, p. 178. 

Sceloporus variabilis marmoratus, SmitH, Proc. Biol. Soc. Washington, vol. 47, 
1934, pp. 125-128; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 
272-278. 

Sceloporus delicatissimus HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, vol. 
6, 1852, p. 178 (San Antonio, Tex.; type, U.S.N.M. No. 16020). 

Type.—Presumably lost. 

Type locality —San Antonio, Tex. 

Range.—Atlantic lowlands from San Antonio, Tex., south to 
southern Tamaulipas, west to Coahuila. Recorded in Mexico only 
from the states of Coahuila, Nuevo Leén, and Tamaulipas. 

SCELOPORUS PARVUS PARVUS Smith 

Sceloporus parvus Smiru, Trans. Kansas Acad. Sci., vol. 37, 1934, pp. 263-267, 
figs. 1, 3, 10. 

Sceloporus parvus parvus, SmiruH, Occ. Pap. Mus. Zool. Univ. Michigan, No. 358, 
1937, pp. 3-4; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 251- 
255, fig. 44. 

Type-—EHT-HMS No. 7120 (EHT field No. 279); E. H. Taylor 
and H. M. Smith collectors. 

Type locality.— Hills 5 miles west of Sabinas Hidalgo, Nuevo Leén. 

Range.—Northern Nuevo Leén to southern San Luis Potosi, west 
of the Sierra Oriental. Recorded only from the two states cited, and 
from Coahuila (Arteaga). 


SCELOPORUS PARVUS SCUTULATUS Smith 


Sceloporus parvus scutulatus SmiruH, Occ. Pap. Mus. Zool. Univ. Michigan., No. 
358, 1937, pp. 4-6; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 255-256. 
Type —EHT-HMS No. 7129; E. H. Taylor and H. M. Smith 
collectors. 
Type locality.—Thirty miles north of Matehuala, San Luis Potosi. 
Range.—Northwestern San Luis Potosi south along the Sierra 
Oriental to southern Hidalgo. Recorded only from the two states 
cited. 


132 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


SCELOPORUS COUCHII Baird 


Sceloporus couchtt Batrp, Proc. Acad. Nat. Sci. Philadelphia, 1858 (1859), p. 254.— 
Smiru, Publ. Field Mus, Nat. Hist., zool. ser., vol. 26, 1939, pp. 240-246, 
pl. 19, fig. 43. 

Lysoptychus lateralis Corr, Proc. U. 8. Nat. Mus., vol. 11, 1888, pp. 397-398, 
pl. 26, fig. 1 (San Diego, Duval Countv, Tex.; U. S. N. M. No. 14741). 

Sceloporus lateralis, BOULENGER, Proc. Zool. Soc. London, 1890, p. 78. 


Type —U.S.N.M. No. 2739 (9 specimens), cotypes; Lt. Darius 
Nash Couch collector. 

Type locality —Santa Catarina, Nuevo Leén (specimens collected 
by Lt. Couch were also at hand from “‘ Pesqueria Grande,’’ Mexico). 

Range.—Central and northern Nuevo Leén and eastern Coahuila. 
In the United States, reported from southern Texas.” 


MERRIAMI GROUP 


Species—One, with two subspecies. 
Range.—Southwestern Texas and adjacent Coahuila. 


KEY TO SPECIES OF THE MERRIAMI GROUP OF SCELOPORUS 


1. Anterior section of frontal usually divided; frontoparietals usually divided; 
outer row of labiomental scales rarely terminating with the first scale wedged 
between first infralabial and first postmental; head scales rugose; sub- 
caudal surface distinctly banded; gular bars extensive, confluent medially. 

merriami annulatus (p. 133) 

Anterior section of frontal rarely divided; frontoparietals rarely divided; outer 
row of labiomentals terminating with the first scale wedged between first 
infralabial and first postmental; head scales smooth; subcaudal surface 
nearly or quite immaculate, not barred; gular bars short, usually separate 
medialliyo fae Ase eet OR er ee ee merriami merriami (p. 1382) 


SCELOPORUS MERRIAMI MERRIAMI Stejneger 


Sceloporus merriami SresneGER, Proc. Biol. Soc. Washington, vol. 17, 1904, pp. 
17-20. 

Sceloporus merriami merriami, SmiruH, Proc. Biol. Soc. Washington, vol. 50, 1937, 
pp. 83-86. 


Type.—U.S.N.M. No. 33039; W. Lloyd collector. 

Type locality.—Kast Painted Cave, near mouth of the Pecos River, 
Tex. 

Range.—The Rio Grande and its immediate tributaries from western 
Brewster County to southeastern Val Verde County, Tex. The species 
occurs on the bluffs along the Mexican side of the river, but no specific 
records for either Coahuila or Chihuahua (in both of which states it 
should occur) are known. 


" Except possibly for the type specimen of Lysoptychus lateralis, no specimen of S. couchii has ever, to our 
knowledge, been taken in Texas. We consider it doubtful that the specimen came from there. 


CHECKLIST OF REPTILES OF MEXICO 133 


SCELOPORUS MERRIAMI ANNULATUS Smith 


Sceloporus merriami annulatus Smiru, Proc. Biol. Soc. Washington, vol. 50, 1937, 
pp. 83-86; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 289-290, 
pl. f- 
Type.—EHT-HMS No. A787; E. H. Taylor and J. S. Wright 
collectors. 
Type locality.—East slope of the Chisos Mountains, Brewster 
County, Tex. 
Range.—Southern and central Brewster County, Tex., and eastern 
Coahuila (Cuatro Ciénegas, Jaral). 


MACULOSUS GROUP 
Species.—One. 
Range.—Eastern Durango. 


SCELOPORUS MACULOSUS Smith 


Sceloporus maculosus Smirn, Trans. Kansas Acad. Sci., vol. 37, 1934, pp. 267-269, 
pl. 8, figs. 2, 4, 5, pl. 10, fig. 11; Publ. Field Mus. Nat. Hist., zool. ser., vol. 
26, 1939, pp. 291-294, fig. 47. 
Type.—EHT-HMS No. 7638; E. H. Taylor and H. M. Smith 
collectors. 
Type locality.—Fourteen miles northeast of Pedricefia, Durango. 
Range.—Known only from east-central Durango. 


CHRYSOSTICTUS GROUP 
Species. —One. 
Range.—Yucatan Peninsula. 


SCELOPORUS CHRYSOSTICTUS Cope 


Sceloporus chrysostictus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1866 (1867), 
p. 125.—Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 
295-300, pl. 24. 

Type.—U.S.N.M. No. 24865, six cotypes. 

Type locality. —Yucatan, here restricted to Chichen ItzA. 

Range.—The Yucatan Peninsula, and the area at its eastern base. 
Recorded in Mexico from Yucatan, Campeche, and Quintana Roo. 


SINIFERUS GROUP 


Species. —Four, one of which includes two subspecies. 
Range.—Pacific slopes from Guerrero to Nicaragua. 


KEY TO SPECIES OF THE SINIFERUS GROUP OF SCELOPORUS 


1. Nasals and anterior internasals broadly in contact with rostral; no postrostrals; 
postanals enlarged in males; ventral scales notched___ochoterenai (p. 135) 
Nasals and anterior internasals separated from rostral by two or more postros- 
iralssvenoral scalesenot MOUCDEO = eee ee eee se ea 2 


134 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


2. One CRNUDAl = a2 ee cine ee See ee ee cage enn ee 3 
HU oh eR UE Ua a 2 a a oir pe tn ee eh 4 
3. Femoral pores 11-12; dorsal scales 38-44____________- carinatus (p. 135) 
Femoral pores 3-6; dorsal scales 28-37___._._.________-- squamosus (p. 135) 


4. Femoral pores 3-11; postanals not or slightly enlarged in males. 
siniferus siniferus (p. 134) 
Femoral pores 12-14; postanals distinctly enlarged in males. 
siniferus cupreus (p. 134) 


SCELOPORUS SINIFERUS SINIFERUS Cope 


Sceloporus siniferus Corn, Proc. Amer. Philos. Soc., vol. 11, 1869, pp. 159-160, 
161.—Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 313- 
319 (part), pl. 25, 

Sceloporus humeralis Bocourt, Ann. Sci. Nat. zool., ser. 5, vol. 17, No. 10, 1873, 
p. 2 (Mus. Hist. Nat. Paris; Oaxaca, here restricted to the city of Tehuante- 


pec). 
Type.-—U.S.N.M. Nos. 30453-30471, cotypes. 
Type locality——FPacific side of the Isthmus of Tehuantepec, here 
restricted to Tehuantepec (city), Oaxaca. 
Range.—Pacific slopes from western Guerrero to extreme western 
Guatemala, and inland to Morelos. Recorded in Mexico from 
Guerrero, Morelos, Oaxaca, and Chiapas. 


SCELOPORUS SINIFERUS CUPREUS Bocourt ° 


Sceloporus cupreus Bocourt, Ann. Sci. Nat., Zool., ser. 5, vol. 19, No. 4, 1873, p. 3; 
Mission scientifique au Mexique . . ., Etudes sur les reptiles, livr. 4, 1874, 
pp. 210-212, pl. 18 bis, fig. 2, 2a-b.—Smiru, Occ. Pap. Mus. Zool. Univ. 
Michigan, No. 358, 1937, pp. 6-9; Publ. Field Mus. Nat. Hist., zool. ser., 
vol. 26, 1939, pp. 305-308. 

Sceloporus cochranae Smitu, Proc. Biol. Soc. Washington, vol. 49, 1936, pp. 87- 
89, pl. 2 (Mount Zempoaltepec, Oaxaca; U.S.N.M. No. 47605). 

Sceloporus siniferus, Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
p. 317 (part). 


Type.—Mus. Hist. Nat. Paris; Adolphe Boucard collector. 





% It is with some doubt that the name cupreus is applied to the highland form of siniferus. This was 
discussed in some detail previously (Publ Field Mus. Nat. Hist., zool, ser., vol. 29, 1939, p. 317). Speci- 
mens then available showed a higher femoral pore count for the types of cupreus and of cochranae (the latter 
from Mount Zempoaltepec, Oaxaca), for which the name cupreus was used, than for the few specimens 
referred to siniferus then available from the vicinity of Oaxaca City. However, new material from the 
latter region (including EHT-HMS 19195-19202, 19479-80) reveals a variation from 38 to 50 (ay. 44.2) in 19 
counts of dorsal scales, and from 6 to 15 (av. 9.4) in 40 counts of femoral pores. 

From these data it is apparent that the highland population of siniferus is recognizably different from the 
typical subspecies and deserves a name. Handicapped by lack of adequate material from Mount Zem- 
poaltepec and its environs, we find it difficult to determine whether the population from that area is the same 
as that from the Oaxaca region, since the single specimen available has enlarged postanals and broad dor- 
solateral light stripes. Typically siniferus (including Oaxaca specimens) lacks enlarged postanals; none 
has been observed with them well developed. Likewise, distinct and broad dorsolateral stripes do not 
occur in male siniferus, although they are present in females. None of the other characters previously thought 
to be peculiar to the Mount Zempoaltepec specimen are actually so, as shown by data on the Oaxaca series. 
Moreover, there is some slight variation in the size of the basal caudal scales and in the width and distinct- 
ness of the dorsolateral light stripes in males. In view of this variability in the only two characters defining 
the Mount Zempoaltepec specimen (cupreus, sensu stricto), we believe it best under the circumstances to 
extend the concept of this name to include the Oaxaca specimens; it should then, of course, be treated as a 
subspecies, if the present interpretation of the name is correct. 


CHECKLIST OF REPTILES OF MEXICO 135 


Type locality.—Oaxaca, here restricted to Mount Zempoaltepec. 
Range.—Highlands of central Oaxaca. 


SCELOPORUS SQUAMOSUS Bocourt 


Sceloporus squamosus Bocourt, Mission scientifique au Mexique .. ., Etudes 
sur les reptiles, livr. 4, 1874, pp. 212-214, pl. 18 bis, figs. 7, 7a—c, pl. 19, fig. 
3.—SmirtH, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 319-324, 
fig. 52, pl. 24. 

Sceloporus fuluus Bocourt, tbid., pp. 214-215, pl. 18 bis, figs. 8, 8a-c (La Unién, 
El Salvador; Commission Scientifique collector; Mus. Hist. Nat. Paris). 

Type-—Mus. Hist. Nat. Paris; collected by (?) Commission Scien- 
tifique du Mexique. 
Type locality —‘ Environs de Guatemala et de l’Antigua”’ [~Volc4n 

Antigua], 1,500 meters. 

Range.—Pacific slopes from eastern Chiapas to Costa Rica. Re- 
corded only from Chiapas in Mexico. 


SCELOPORUS CARINATUS Smith 


Sceloporus carinatus Smitu, Proc. Biol. Soc. Washington, vol. 49, 1936, pp. 89- 
91, pl. 2, figs. 2, 3; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 303-305. 
Type —EHT-HMS No. 15205 (EHT-HMS field No. 4866); E. H. 
Taylor and H. M. Smith collectors. 
Type locality — Near Tuxtla Gutiérrez, Chiapas. 
Range.—Plateau of Chiapas. 


SCELOPORUS OCHOTERENAI Smith 


Sceloporus ochoterenae Smitu, Trans. Kansas Acad. Sci., vol. 37, 1934, pp. 269- 
272, pl. 9, figs. 6, 9, pl. 10, fig. 12; Publ. Field Mus. Nat. Hist., zool. ser., 
vol. 26, 1939, pp. 308-313, text fig. 51. 

Type-—EHT-HMS No. 7158 (EHT field No. 1075); E. H. Taylor 
and H. M. Smith collectors. 

Type locality—Two miles north of Mazatlan, Guerrero (12 miles 
south of Chilpancingo, Guerrero). 

Range.—Morelos and Guerrero. 


UTIFORMIS GROUP 
Species.—One. 
Range.—Western Mexico, Sinaloa to Guerrero. 


SCELOPORUS UTIFORMIS Cope 


Sceloporus utiformis Corn, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 177.— 
Smitu, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 325-330, 
fig. 53, pl. 26. 
Type.—U.S.N.M. No. 42089; John Xantus collector. 
Type locality —Near Colima, Colima. 


136 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Range.—Pacific slopes from southern Sinaloa to western Guerrero, 
inland about 125 miles in the southern part of its range. Recorded 
only from Sinaloa, Nayarit, Jalisco, Colima, Michoacan, and Guerrero. 


SCALARIS GROUP 


Species.—Seven forms are recognized at present, representing four 


species. 
Range.—High elevations from southeastern Arizona and central 
Nuevo Leén southward to the edge of the plateau in Oaxaca. 


KEY TO SPECIES OF THE SCALARIS GROUP OF SCELOPORUS 


1. Nasals and internasals in contact with rostral; no postrostrals_jalapae (p. 136) 


Nasals and internasals separated from rostral by two postrostrals________ 2 

2. Dorsal scales 50 or more; lateral scales in slightly, although distinctly, oblique 
TOWA sere = So ee Oe ee ee ee en goldmani (p. 137) 
Dorsal scales less than 50; lateral scales in parallel rows__-______________- 3 

3. .Onercanthalass 2.22224 S6 ee a eee eee ee oe ee eae 4 
Two canthals, the first occasionally forced above canthal ridge by contact of 
BECOME Uc surat ka all en ra Cl ess Ua kT Sa a ee 6 


4, Males with much black in ventral coloration; females suffused with black 
below; gular region never barred; black shoulder spot with a light blue spot, 
if present, on its anterior edge; tail with a continuous dark median dorsal 
stripe; tibia/head proportion usually less than 0.95; maximum snout-vent 
measurement 09 MMo 22 ss osee sae mece aaa eee aeneus aeneus (p. 137) 

Black, if present on ventral surface, confined to bars in gular region, and a few 
dark, transverse bars on sides of abdomen; black shoulder spot with the light 
blue'spotan itssmiddlet.. 2222. “Sees se sae ee eee Ee Se 5 

5. Tibia/head proportion usually less than 0.90; scales of second pair of post- 
mentals separated medially; dorsal scales usually more than 40; maximum 
snout-vent measurement 61 mm___--_---_------- scalaris slevini (p. 138) 

Tibia/head proportion usually more than 0.90; scales of second pair of post- 
mentals usually in contact medially; dorsal scales usually less than 40; 
maximum snout-vent measurement 65 mm_-_scalaris unicanthalis (p. 138) 

6. Males with much black in ventral coloration; black shoulder spot with the 
light blue spot, if present, on its anterior edge; tail with a continuous dark 
median dorsal stripe; tibia/head proportion usually less than 0.90; maximum 
snout-vent measurement 56 mm__-_-____----- aeneus bicanthalis (p. 137) 

Black, if present on the ventral surface, confined to bars in gular region, and a 
few dark transverse bars on sides of abdomen; black shoulder spot with the 
light blue spot in its middle; tail with dark chevron-shaped bars; tibia/head 
proportion usually more than 0.90; maximum snout-vent measurement 
GOLAN oe = one ore ere crc eee ee eee es er scalaris scalaris (p. 137) 


SCELOPORUS JALAPAE Giintker 


Sceloporus jalapae GtnrueEr, Biologia Centrali-Americana, Reptilia and Batra- 
chia, 1890, p. 74.—Smrru, Proc. Biol. Soc. Washington, vol. 47, 1934, pp. 
121-125, fig. 1; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 
333-338, figs, 55, 56, pl. 27. 


Type.—Brit. Mus. Nat. Hist.; Mr. Hége collector. 
Type locality. Jalapa, Veracruz. 


CHECKLIST OF REPTILES OF MEXICO 17 


Range.—Central Veracruz south through eastern Puebla to central 
Oaxaca. Recorded only from the states cited. 


SCELOPORUS AENEUS AENEUS Wiegmann 


Sceloporus aeneus WIEGMANN, Isis von Oken, vol. 21, 1828, p. 370. 

Sceloporus aeneus aeneus, SmitH, Occ. Pap. Mus. Zool. Univ. Michigan, No. 361, 
1937, p. 6; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 353-356, 
text fig. 57. 

Type.—Zool. Mus. Berlin; F. Deppe collector. 

Type locality—Mexico, by inference, here restricted to Tres 
Cumbres. 

Range——Western Puebla, to central western Michoacin, and 
Jalisco; north on the plateau to northern Guanajuato. Recorded 
from the states cited and from Distrito Federal, México, Morelos, 
and questionably, Jalisco. 


SCELOPORUS AENEUS BICANTHALIS Smith 


Sceloporus aeneus bicanthalis SmirH, Occ. Pap. Mus. Zool. Univ. Michigan, No. 
361, 1937, pp. 6-8; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 356-361, fig. 57, pl. 20. 
Type —EHT-HMS No. 7939; E. H. Taylor, collector. 
Type locality —Cofre de Perote, near Cruz Blanca, Veracruz. 
Range.—From northern Hidalgo along the eastern escarpment of 
the plateau to central Oaxaca, including parts of Puebla, México, 
and Veracruz. Recorded only from the states cited. 


SCELOPORUS GOLDMANI Smith 


Sceloporus goldmant Smit, Occ. Pap. Mus. Zool. Univ. Michigan, No. 361, 1937, 
p. 5; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 350-353. 


Type.—Univ. Michigan Mus. Zool. No. 80896; C. L. Lundell 
collector. 

Type locality —Charcas, San Luis Potosi. 

Range.—Southern Coahuila southward in the central part of the 
plateau of Mexico to central San Luis Potosi. Recorded only from 
the states cited. 


SCELOPORUS SCALARIS SCALARIS Wiegmann 


Sceloporus scalaris WIEGMANN, Isis von Oken, vel. 21, 1828, p. 370. 

Sceloporus scalaris scalaris, SmirH, Occ. Pap. Mus. Zool. Univ. Michigan, No. 361, 
1937, pp. 2-3; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 338- 
343, pl. 20. 


Type.—Zool. Mus. Berlin; F. Deppe collector. 

Type locality.—Mexico, here restricted to México, D. F. 

Range.—Central Durango, southward over the central and southern 
parts of the Mexican Plateau except (apparently) Guerrero. Re- 


138 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


corded from Distrito Federal and the states of Durango, Guanajuato, 
Hidalgo, Jalisco, México, Michoac4n, Puebla, and Zacatecas. 


SCELOPORUS SCALARIS UNICANTHALIS Smith 


Sceloporus scalaris unicanthalis Smrra, Occ. Pap. Mus. Zool. Univ. Michigan, No- 
361, 1937, pp. 4-5; Publ. Field Mus. Nat. Hist. zool. ser., vol. 26, 1939, 
pp. 349-350. 
Type.—EHT-HMS No. 7699; H. M. Smith collector. 
Type locality — Magdalena, Jalisco. 
Range.—Known only from the southern border of the Mexican 
plateau in central Jalisco, from the eastern border of Lake Chapala 
westward to Nayarit. Recorded only from the state of Jalisco. 


SCELOPORUS SCALARIS SLEVINI Smith 


Sceloporus scalaris slevini Smita, Occ. Pap. Mus. Zool. Univ. Michigan, No. 361, 
1937, pp. 3-4; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 
343-349. 

Type.—Calif. Acad. Sci. No. 48013; J. R. Slevin collector. 
Type locality.—Miller Peak, Huachuca Mountains, Cochise County, 

Ariz. 

Range.—Southern Arizona south to northern Durango, and east to 
to Nuevo Leén in Mexico. Recorded from the states cited and from 

Sonora, Chihuahua, and Coahuila. 


PYROCEPHALUS GROUP 
Species.—Three. 
Range.—Pacific slopes from southwestern Chihuahua to north- 
western Oaxaca. 


KEY TO SPECIES OF THE PYROCEPHALUS GROUP OF SCELOPORUS 


1, Scales on posterior surface of thigh granular; postfemoral dermal pocket 


POLESC TG eee i a rs A ES Sect ay eee rece gadoviae (p. 138) 
Scales on posterior surface of thigh larger, imbricate; no postfemoral dermal 
pocketswcves ole S Leet Pe EE ee Shee Soe Se 2 


2. Dorsal scales larger, 36-41; females not red-headed; males with unbroken 
lateral belly patches; dark spot on interparietal not enclosing or touching 
light pinealispotics) 7200. oo Bek 2 ees ee ee ee nelsoni (p. 139) 

Dorsal scales smaller, 41-50; females red-headed, conspicuously barred below 
on throat; males (and usually females) with a series of broad dark bars on 
each side of belly; a dark spot surrounding light pineal spot. 

pyrocephalus (p. 139) 


SCELOPORUS GADOVIAE Boulenger 


Sceloporus gadoviae BOULENGER, Proc. Zool. Soc. London, 1905, vol. 2, pp. 246= 
247, pl. 7, fig. 1.—Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 373-377, fig. 53, pl. 31. 
Type.—Brit. Mus. Nat. Hist.; Hans Gadow collector. 
Type locality. —Mezquititlan, north of Chilpancingo, Guerrero. 


CHECKLIST OF REPTILES OF MEXICO 139 


Range. —Southern Michoacan through Guerrero, Morelos, southern 
Puebla to northern and western Oaxaca. Recorded only from the 


states cited. 
SCELOPORUS PYROCEPHALUS Cope 


Sceloporus pyrocephalus Corn, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 177.— 
Smitu, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 368-373, 
pls. 29, 30. 

Sceloporus pyrrhocephalus Corr, Proc. Amer. Philos. Soc., vol. 22, 1885, pp. 
394-397 (emendation). 


Type.—U.S.N.M. No. 31495; John Xantus collector. 

Type locality—Near Colima, Colima. 

Range.—Central Jalisco, south and east through Colima and 
Michoacin to Guerrero. Recorded only from states cited. 


SCELOPORUS NELSONI Cochran 


Sceloporus nelsoni CocHraAN, Journ. Washington Acad. Sci., vol. 13, 1923, pp. 
185-186.—Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 
364-368, pl. 28. 

Type.-—U.S.N.M. No. 47676; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality—Plomosas, Sinaloa. 

Range—Pacific slopes from southwestern Chihuahua to northwestern 
Jalisco. Recorded from Chihuahua, Sonora, Sinaloa, Nayarit, and 
Jalisco. 

Genus SATOR Dickerson 


Sator DickERSON, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, pp. 468-469. 


Genotype.—Sator grandaevus Dickerson. 
Range.—Santa Cruz and Cerralvo Islands in the Gulf of California. 


Species.—Two. 
KEY TO SPECIES OF SATOR 


1. A poorly defined anterior gular fold, without granules; a posterior (true) gular 
fold immediately in front of arm insertions, but not continuous across throat; 
median lateral scales very small, often granular, abruptly differentiated from 
dorsalsyand) ventrals: 22-2 223222554225) obese Se grandaevus (p. 140) 

A well defined anterior gular fold, with granules; no evidence whatever, even 
at sides in front of arm insertions, of a posterior gular fold; median lateral 
scales larger, gradually merging with dorsals and ventrals_angustus (p. 139) 


SATOR ANGUSTUS Dickerson 


Sator angustus DickERsoN, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, p. 469.— 
Scumipt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 665-666, fig. 10.— 
Van DENBURGH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 256-258. 

Type.—Amer. Mus. Nat. Hist. No. 5712; C. H. Townsend collector. 
Type locality—Santa Cruz Island, Gulf of California, Baja Cali- 
fornia. 


Range.—Restricted to the type locality. 
861316—50-——10 


140 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 
SATOR GRANDAEVUS Dickerson 


Sator grandaevus Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, pp. 
469-470.—Scumipt7, 2zbid., vol. 46, 1922, p. 665, figs. 8, 9 —Van DEeNBURGH, 
Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 259-261. 


Type.—Amer. Mus. Nat. Hist. No. 5491; C. H. Townsend collector. 
Type locality.—Cerralvo Island, Gulf of California, Baja California. 
Range.—Restricted to the type locality. 


Genus UROSAURUS Hallowell 


Urosaurus HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, vol. 7, 1854, p. 92.— 
MirrLeMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, pp. 103-181, pls. 1-16. 

Phymatolepis Dumirit, Arch. Mus. Hist. Nat. Paris, vol. 8, 1856, p. 548 (type, 
Phymatolepis bicarinatus Duméril). 


Genotype.—Urosaurus graciosus Hallowell. 

Range.—Southern Texas west to California, entering Utah and 
Colorado, southward through Baja California and many adjacent 
islands, and through mainland Mexico from the western Texas border 
south to the plateau and along the western slopes to Chiapas. 

Species.—Ten species and 21 forms are recognized; all 10 species 
and 19 forms are known to occur or are to be expected in Mexico. 


KEY TO MEXICAN % SPECIES OF UROSAURUS 


1. Enlarged anterodorsal femoral scales smooth___-_--- auriculatus (p. 146) 

Enlarged anterodorsal femoral scales strongly keeled____._----.--------- 2 

2. Enlarged dorsals in a single broad band, uninterrupted by an intervening 

series of vsmealleniscales. 24. a eS ee ae ee eee ee 3 

Enlarged dorsals separated into two or more parallel series by the presence of 

aavertebral series of smalleriscaleshaes eee eee ee ee ae di 

3. Tail two or more times length of head and body combined_graciosus (p. 144) 

Tail less than twice length of head and body combined_-_-------------- 4 

4. Dermal folds, when present, not heavily crested with tubercles; blue abdominal 

patches only in males; enlarged dorsals comparatively small___--_----- 5 

Dermal folds present, always crested with tubercles of fairly large size; 

abdominal blue patches sometimes in females as well as males____--_---- 6 

5. Enlarged dorsals larger, from 17 to 24 in length of head from tip of snout to 

posterior edge of interparietal; gular region in males deep yellow or orange. 

nigricaudus (p. 145) 

Enlarged dorsals smaller, 32 to 36 in length of head from tip of snout to 
posterior edge of interparietal; gular region in males usually blue. 

microscutatus (p. 145) 

6. Four to seven rows of enlarged dorsal scales; abdomen of both sexes with a 

blue wash and/or blue patches; dorsolateral folds not converging in sacral 

BO QUOT bre tAPh Hor ANU fia (A aaa hal TR ep ae cae cease Ei ae gadovi (p. 145) 

About three rows of enlarged dorsal scales; only males with a blue abdomen; 

dorsolateral folds converging in the sacral region to form prominent ridges. 

irregularis (p. 146) 

(= chhreeordewer rows of lateral tuberclesee= = == == soe ae ee eee eee 12 

Hour.onmoremows ollaterallitulbencl estes. = se a= ee eee ee 8 


81 Adapted from Mittleman, op. cit., pp. 127-133. 


10. 


te 


12. 


13. 


14, 


15. 


16. 


CHECKLIST OF REPTILES OF MEXICO 141 


. Enlarged dorsals commencing caudad of a line joining the anterior points of 


insertion of the forelimbs; dorsals weakly keeled, rounded posteriorly, 
prominently pavemented; general habitus not at all rugose_unicus (p. 147) 


Enlarged dorsals commencing craniad of a line joining the anterior points of 
insertion of the forelimbs, or else equal with such a line; dorsals prominently 
keeled, usually mucronate or spinose, imbricate; ventrals imbricate; general 
SppeaLance Wu OSes S225 LSa= hoes Soo ss So eee on eee eee eee 9 


. Form rugose; enlarged dorsals strongly carinate and prominently mucronate; 


tubercles of lateral and dorsolateral folds well developed; ventrals mucro-_ 
nate; gular surface generally stippled, with a light median area; blue ab-_ 
dominal patches of males quite extensive__bicarinatus bicarinatus (p. 146) 


General appearance somewhat less rugose; enlarged dorsals not so often 
mucronate; tubercles smaller, sometimes absent; ventrals less mucronate, 
occasionally rounded; gular surfaces evenly stippled; abdominal blue of 
males sometimes restricted to small sternal patches__--------------- 10 


Ventrals rounded; dorsolateral and lateral tubercles very poorly developed; en- 
larged dorsals commencing on the nape_-_-_--- bicarinatus nelsoni (p. 147) 


Ventrals submucronate to mucronate; dorsolateral and lateral tubercles well 
developed; enlarged dorsals commencing on the shoulders just craniad of a 
line joining the anterior points of insertion of the forelimbs_________- il 


Ventrals mucronate, prominently carinated laterally; gular scales with a 
tendency toward pavementation, especially anteriorly; gular surfaces evenly 
stippled; abdominal blue of males restricted to small sternal patches. 

bicarinatus anonymorphus (p. 146) 

Ventrals submucronate (occasionally rounded), only faintly keeled on the 
lateral portions of the belly, or else not at all; gular scales imbricate; gular 
region with an even blue wash, and only barely flecked if at all; abdominal 


blue of males evenly distributed_-_-_--_- bicarinatus tuberculatus (p. 147) 
HMolareed dorsals commencing om Wape=—- —-- 2. 2 = =e sae eee 13 
Enlarged dorsals commencing on shoulders or caudad of them_---_----- 14 


Enlarged dorsals strongly keeled; scales of primary and secondary serieS 
almost equal in size; postfemoral dermal pocket absent or rudimentary; no 
prominent lateral pattern of dark whorls__--------- clarionensis (p. 142) 

Enlarged dorsals not so rugose, scales of primary series prominently larger 
than those of the secondary series; postfemoral dermal pocket regularly 
present; a distinct lateral pattern of dark whorls_ornatus schottii (p. 144) 


Tail two or more times length of head and body combined_graciosus (p. 144) 
Tail less than twice length of head and body combined__-------------- 15 


Enlarged dorsals extending onto basal portion of tail for a distance equal to 
length of femur, or more; entire gular region, including sublabials, a uniform 
bright) le bt 2 bet os ees Pee oe ete ornatus caeruleus (p. 143) 

Enlarged dorsals extending onto basal portion of tail for a distance equal to less 
than length of femur; entire gular region including sublabials never com- 
TON Ge ly, oO ae 16 

Enlarged dorsals often irregularly arranged; lateral tubercles not affecting 
diagonal arrangements; average size less than 45 mm. from snout to 
SU EST a a 17 

Enlarged dorsals regularly arranged in parallel series on either side of the 
vertebrals; tubercles in parallel diagonal series; average size greater than 45 
MIMeirOMESHOUL tOMWeNts 2.2) —— = o—- ae ene eee eae eee 18 


142 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


17. Scales of primary series not twice as Jarge as those of the secondary series; 
largest of the dorsals inferior in size to enlarged femorals and tibials; ventral 
interhumeral and interfemoral areas immaculate, or but slightly stip- 
Pledue oe 2 Se ea See eee eee re ornatus schmidti (p. 143) 

Seales of primary series almost twice as large as those of secondary series; 
largest of dorsals equal to, or larger than, enlarged femoral and tibial scales; 
ventral interhumeral and interfemoral areas heavily maculated. 

ornatus ornatus (p. 142) 

18. Largest of dorsals equal to, or larger than enlarged femorals; vertebrals extend- 
ing onto basal portion of tail for a distance equal to half, or slightly more, of 
length of femur; entire gular region in males, except sublabials, an intense 
blue; head length/head width ratio averaging 81 percent. 

ornatus chiricahuae (p. 143) 

Largest of dorsals inferior in size to enlarged femorals; vertebrals extending 
only onto rump, or but slightly farther; no uniform intense blue color present 
in male gular region; head length/head width ratio averaging 75 percent or 
LOSS! Asa oe aCe SR A ee ee ee Se i ee 19 

19. Enlarged dorsals separated into 2 parallel series by width of vertebral series, 
which is greater in width than broadest of enlarged dorsals; prefrontals and 
frontonasals usually 3 each; general coloration pallid, light tan above, 
whitish below, males with bright-blue abdominal patches; average head 
length/width ratio 75.4 percent; average length, snout to vent, 55.1 mm. 

ornatus symmetricus (p. 144) 

Enlarged dorsals separated by a vertebral series whose width is less than that 
of largest of dorsal scales; prefrontals 2, rarely 3 (by inclusion of an azy- 
gous); frontonasals 5; general color variable, but usually dark brown or 
gray with dark cross bands, and heavily stippled, spotted, or blotched ven- 
trally; abdominal patches in males dark blue to indigo; average head 
length/head width ratio 70.6 percent; average length, snout to vent, 46.4 mm. 

ornatus linearis (p. 143) 
ORNATUS GROUP 


Species—Three species and 11 forms are included; all but two 
subspecies of ornatus occur or may be expected to occur in Mexico. 
Range.—Utah, central Texas, and southern Nevada southward to 
extreme northern Mexico, reaching farther southward only along the 
coast to southern Sinaloa; Clarion Island. 
UROSAURUS CLARIONENSIS (Townsend) 
Uta clarionensis TOWNSEND, Proc. U. S. Nat. Mus., vol. 13, 1890, p. 148. 
Urosaurus clartonensis, MirTLEMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, p. 
149, pl. 8 (type). 
Type—U.S.N.M. No. 15904; C. H. Townsend collector. 
Type locality—Clarion Island, Revillagigedo Islands, Mexico. 
Range.—Known only from the type locality (administered by 
Colima). 
UROSAURUS ORNATUS ORNATUS (Baird and Girard) 
Uta ornata (part) Barrp and Grrarp, Proc. Acad. Nat. Sci. Philadelphia, vol. 6, 
1852, p. 126. 
Uta ornata ornata (part), Scumipt, Amer. Mus. Nov., No. 22, 1921, p. 6. 
Urosaurus ornatus ornatus, Mirtteman, Bull. Mus. Comp. Zool., vol. 91, 1942, 
pp. 133-135. pl. 1 (cotypes).—Smitu, Handbook of lizards, 1946, pp. 264- 
266, pl. 65. 


CHECKLIST OF REPTILES OF MEXICO 143 


Type.—U.S.N.M. No. 2750, two cotypes, male and female. 

Type locality —Restricted to Rio San Pedro [Devils River], Val 
Verde County, Tex. 

Range.—Central and southern Texas, south to northern Coahuila. 
Has been taken along the Rio Grande River on the American side, 
and very probably occurs in Coahuila. 

UROSAURUS ORNATUS CAERULEUS (Smith) 


Uta caerulea Situ, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 172-178, pl. 26. 
Urosaurus ornatus caeruleus, M1irTLEMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, 
pp. 186-137, pl. 9, lower fig. 


Type.—Univ. Kansas Mus. Nat. Hist., No. 19237 (Smith and Dun- 
kle field No. 132); Hobart M. Smith and David H. Dunkle collectors. 

Type locality —Thirty miles north of Chihuahua City, Chihuahua. 

Range.—Central Chihuahua. Reported only from the type locality 
and from 20 miles south of Chihuahua. 


UROSAURUS ORNATUS CHIRICAHUAE (Mittleman) 


Uta ornata chiricahuae MirTLEMAN, Proc. Biol. Soc. Washington, vol. 54, 1941, 
p. 165. 

Urosaurus ornatus chiricahuae, MirTLEMAN, Bull. Mus. Comp. Zool., vol. 91, 
1942, pp. 189-142.—Smiru, Handbook of lizards, 1946, p. 266, pl. 66. 


Type.—Mus. Vert. Zool. Univ. California, No. 7751. 

Type locality.—Pinery Canyon, Chiricahua Mountains, 6,000 feet, 
Cochise Co., Ariz. 

Range.—So far as known, restricted to the Chiricahua and Dos 
Cabezas Mountains, Ariz.; of probable occurrence in Mexico. 


UROSAURUS ORNATUS LINEARIS (Baird) 


Uta ornata var. linearis Batrp, United States and Mexican boundary survey, 
1859, vol. 2, pt. 2, p. 7. 

Uta ornata linearis, Scumipt, Amer. Mus. Nov., No. 22, 1921, p. 6. 

Urosaurus ornatus linearis, MirrTLEMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, 
pp. 187, 139, pl. 3 (neotype).—Smiru, Handbook of lizards, 1946, pp. 268-271, 
pl. 67. 

Type.—U.S.N.M. No. 2759 (now lost); Caleb Kennerly collector. 
Neotype: U.S.N.M. No. 62077, Los Nogales, Sonora, Mexico; F. J. 
Dyer, collector. 

Type locality. —Los Nogales, Sonora, Mexico. 

Range.—Southern Arizona and southern New Mexico, southward 
to northern Sonora and Chihuahua. 


UROSAURUS ORNATUS SCHMIDTI (Mittleman) 


Uta ornata schmidti MirrtemMan, Herpetologica, vol. 2, pt. 2, 1940, pp. 33-34, 
pl. 3, fig. 1 (1941). 

Urosaurus ornatus schmidti, MirrLEMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, 
pp. 135-136, pl. 2 (type).—Smirn, Handbook of lizards, 1946, pp. 271-272, 
pl. 68). 


144 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type.—U.S.N.M. No. 32929; V. Bailey collector. 

Type locality.—Fort Davis, Jeff Davis County, Tex. 

Range. —Southwestern Texas and northern Chihuahua. Reported 
in Mexico only from Chihuahua: Samalayuca. 


UROSAURUS ORNATUS SCHOTTII ® (Baird) 


Uta schottit Barrp, Proc. Acad. Nat. Sci. Philadelphia, vol. 10, 1858, p. 252. 

Uta ornata schottii, MirrLEMaAN, Copeia, 1941, pp. 1386-138. 

Urosaurus ornatus schottii, MirrLeMaAN, Bull. Mus. Comp. Zool., vol. 91, 1942, 
pp. 149-151. 

Uta (Phymatolepis) lateralis BoULENGER, Ann. Mag. Nat. Hist., ser. 5, vol. 11, 
1883, p. 342 (Brit. Mus. Nat. Hist.; type locality restricted to Presidio, 
Sinaloa). 

Uta lateralis, BOULENGER, Catalogue of the lizards of the British Museum, vol. 
2, 1885, p. 214. 

Uta ornata lateralis, VAN DENBURGH, Occ. Pap. California Acad. Sci., No. 10, 
1922, pp. 199-201.—O.IvEr, Copeia, 1948, pp. 97-107, 1 fig. 

Uta gularis Craain, Bull. Washburn Lab. Nat. Hist., vol. 1, 1884, p. 7 (type 
now lost, formerly in Washburn College collection, Topeka, Kans.; Guay- 
mas, Sonora; Prof. Lovewell collector). 


Type.—U.S.N.M. No. 2761 (now lost); A. Schott collector. 
Type local’ty.—‘‘Sta. Madelina, Cal.-Mex. Boundary survey’”= 
?Maedalena, Sonora, Mexico. 


Range.—Central Sonora and southward to southern Sinaloa, and 
the Tres Marias Islands; Tiburén Island. 


UROSAURUS ORNATUS SYMMETRICUS (Baird) 


Uta symmetrica Barrp, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 253. 

Uta ornata symmetrica, Scumipt, Amer. Mus. Nov., No. 22, 1921, p. 6. 

Urosaurus ornatus symmetricus, MirrLEMAN, Bull. Mus. Comp. Zool., vol. 91, 
1942, pp. 142-144, pl. 4 (neotype).—Smirxa, Handbook of lizards, 1946, 
pp. 273-274, pl. 69. 


Type.—U.S.N.M. No. 2760 (lost). Neotype: U.S.N.M. No. 2744; 
Fort Yuma, Imperial County, Calif.; M. Thomas collector. 

Type locality Fort Yuma, Imperial County, Calif. 

Range.—In United States: southern California and adjacent parts 
of Arizona south to western Sonora and northern Baja California. 
Reported from Sonora: 2 miles south of Nogales, Pinetos Camp, 
32 miles south of Nogales, Duros Millos, Gran Desierto; Baja 
California: “Colorado River valley and desert to Delta.” 


UROSAURUS GRACIOSUS Hallowell 


Uro-saurus graciosus HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, vol. 7, 
1854, p. 92.—Smiru, Handbook of lizards, 1946, pp. 259-262, pl. 63. 

Uta graciosa, Batrp, United States and Mexican boundary survey, 1859, vol. 2, 
pt. 2, p. 7-—Van DeEenBuRGH, Occ. Pap. California Acad. Sci., No. 10, 1922, 
pp. 212-216, pl. 17. 


82 The status of this form is questioned by Oliver, loc, cit., who feels that the name should be Uta ornata 
lateralis Boulenger. 


CHECKLIST OF REPTILES OF MEXICO 145 


Urosaurus ornatus graciosus, MITTLEMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, 
pp. 144-145, pl. 7 (cotypes). 

Type.—Acad. Nat. Sci. Philadelphia Nos. 8550-8551. 

Type locality——‘“Lower California’? [southern California], here 
restricted to Winterhaven (—Fort Yuma), Calif. 

Range.—Southern Nevada, western Arizona, southern California, 
and northern Baja California. Reported from Baja California: 85 
miles south of Mexicali, San Felipe. It may occur also in Sonora but 
is not recorded from there. 


NIGRICAUDUS GROUP 


Species.—Four. 
Range.—Jalisco, Michoacan, and Baja California. 


UROSAURUS NIGRICAUDUS (Cope) 


Uta nigricauda Core, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 176.—Van DEN- 
BURGH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 216-219. 
Urosaurus nigricaudus, MirrLEMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, pp. 
157-159, pl. 10. 
Type.—U.S.N.M. No. 5307, 12 cotypes; John Xantus collector. 
Type locality Cape San Lucas, Baja California. 
Range.—Southern tip of Baja California, with adjacent coastal 
islands, Espiritu Santo, Ballena, San José, and 7Magdalena. 


UROSAURUS MICROSCUTATUS (Van Denburgh) 


Uta microscutata VAN DenspuracH, Proc. California Acad. Sci., ser. 2, vol. 4, 1894, 
p. 298; Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 219-221. 
Urosaurus microscutatus, MirTLEMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, 
pp. 159-163.—Smirn, Handbook of lizards, 1946, pp. 262-264, pl. 64. 
Uta parviscutata Corn, Ann. Rep. U. S. Nat. Mus., 1898 (1900), pp. 324-325, 
fig. 45 (substitute name for Uta microscutata Van Denburgh). 
Type.—Stanford Univ. Mus. No. 1221; J. M. Stowell collector. 
Type locality —San Pedro Ma4rtir Mountains, Baja California. 
Range.—Extreme southern California through all of Baja California 
except the southern fifth. Reported from Baja California: Puerto 
Escondido, San Xavier, San Quintin, San Pedro Ma4rtir, Mount San 
Matias, and the Islands of San Francisco, San José, Danzante, Coro- 
nado, Carmen, San Marcos, Santa Magdalena. 


UROSAURUS GADOVI (Schmidt) 


Uta gadovt Scumipt, Amer. Mus. Nov., No. 22, 1921, pp. 3-4. 
Urosaurus gadovi, MirTLeMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, pp. 154-156, 
pais 
Type.—Amer. Mus. Nat. Hist. No. 20355; Hans Gadow collector. 
Type locality—Cofradia, Jalisco, Mexico. 
Range.—Pacific slopes of Michoacin and Jalisco. Reported from 
Michoacdén: Apatzingian, Acahuato; Jalisco: Cofradia. 


146 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


UROSAURUS IRREGULARIS (Fischer) 


Phymatolepis (Uia) irregularis Fiscoer, Abh. Nat. Ver. Bremen, vol. 7, 1882, pp. 
232-234, pl. 17, figs. 1-4. 
Uta irregularis, BOULENGER, Catalogue of the lizards in the British Museum, 
ed. 2, vol. 2, 1885, p. 216. 
Urosaurus irregularis, MirrLEMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, pp. 
156-157, pl. 12 (upper fig.). 
Type —Municipal Nat. Hist. Coll. Bremen, Germany, No. 437; 
collector unknown. 
Type locality —‘‘Aus dem Hochlande von Mexico.” 
Range.—Mexico highlands. No specific locality known. 


BICARINATUS GROUP 


Species.—Three, with a total of six forms. 
Range.—Pacific slopes from southwestern Chihuahua south to 
Chiapas; Socorro Island. 


UROSAURUS AURICULATUS (Cope) 


Uta auriculata Corr, Proc. Boston Soc. Hist., vol. 14, 1871, p. 303.—VaNn 
DENBURGH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 37, 197-199. 
Urosaurus auriculatus, MirrLemaNn, Bull. Mus. Comp. Zool., vol. 91, 1942, pp. 
163-164, pl. 16 (type). 
Type —U.S.N.M. No. 7027; Grayson collector. 
Type locality —Socorro Island, Revillagigedo Islands. 
Range.—Known only from the type locality. 


UROSAURUS BICARINATUS BICARINATUS (Duméril) 


Phymatolepis bi-carinatus Dumérit, Arch. Mus. Nat. Hist. Nat. Paris, vol. 8, 
1856, p. 549, pl. 23, figs. 2, 2a, 2b. 

Uta bicarinata, Corn, Proc. Acad. Nat. Sci. Philadelphia, vol. 16, 1864, p. 177. 

Uta bi-carinata bi-carinata, MitTLEMAN, Journ. Washington Acad. Sci., vol. 31, 
1941, pp. 70-71, figs. 1G, 2. 

Urosaurus bicarinatus bicarinatus, MitTLEMAN, Bull. Mus. Comp. Zool., vol. 91, 
1942, pp. 164-166, pl. 13. 


Type—Mus. Hist. Nat. Paris, two cotypes; A. Boucard collector. 

Type locality—Mexico, here restricted to Cuernavaca, Morelos. 

Range.—Pacific slopes from Michoac4n to central Guerrero, and 
up the basin of the Rio Balsas to southern Puebla. Reported from 
Puebla: ‘Puebla,’ Iztcar de Matamoros, Tlapanala, Chiautla; 
Morelos: Cuernavaca; Guerrero: Agua del Obispo, Cocoyul, Chil- 
pancingo, Mesquititlan, Iguala, Rio Balsas, etc.; Michoacdn: San 
Blas, Jorullo. 


UROSAURUS BICARINATUS ANONYMORPHUS (Mittleman) 


Uta anonymorpha MirtLteMan, Herpetologica, vol. 2, 1940, pp. 34-38, pl. 3, fig. 2. 
Uta bicarinata anonymorpha, MirTLeMAN, Journ. Washington Acad. Sci., vol. 
31, 1941, pp. 71-72, figs. 1G, 2. 


CHECKLIST OF REPTILES OF MEXICO 147 


Urosaurus bicarinatus anonymorphus, MirrLEMAN, Bull. Mus. Comp. Zool., 
vol. 91, 1942, pp. 166-168, pl. 14 (type). 

Type-—U.S.N.M. No. 46988; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality — Tehuantepec, Oaxaca. 

Range.—Pacific slopes in eastern Guerrero, Oaxaca and western 
Chiapas. Reported from Chiapas: Tonali; Oaxaca: Tehuantepec, 
Juchitan, San Gerdénimo, Ixtepec, Totolapam, Tres Cruces, Mount 
Guengola, Cerro Arenal, etc.; Guerrero: Tierra Colorada. 


UROSAURUS BICARINATUS NELSONI (Schmidt) 


Uta nelsoni Scumipt, Amer. Mus. Nov., No. 22, 1921, p. 4. 

Uta bicarinata nelsoni, MiTTLEMAN, Journ. Washington Acad. Sci., vol. 31, 1941, 
pp. 72-73, figs. 1H, 2. 

Urosaurus bicarinatus nelsoni, MIrTLEMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, 
p. 168, pl. 15. 


Type.—U.S.N.M. No. 46836; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality.—Cuicatlan, Oaxaca. 

Range.—Known only from the type locality. 


UROSAURUS BICARINATUS TUBERCULATUS (Schmidt) 


Uta tuberculata Scumipt, Amer. Mus. Nov., No. 22, 1921, p. 4. 

Uta bi-carinata tuberculata, MirrLEMAN, Journ. Washington Acad. Sci., vol. 31, 
1941, pp. 73-74, figs. LE, 2.—Oniver, Copeia, 1943, p. 105. 

Urosaurus bicarinatus tuberculatus, MirrLEMAN, Bull. Mus. Comp. Zool., vol. 91, 
1942, pp. 169-170, pl. 12, lower fig. 


Type.—Amer. Mus. Nat. Hist. No. 13737; Paul D. R. Riithling 
collector. 

Type locality —Colima, Colima, Mexico. 

Range.—Southern Sonora southward to Jalisco. Sonora: Guirocoba; 
Sinaloa: Presidio de Mazatlan; Jalisco: ?‘‘North of Rio Santiago”’; 
Colima: Colima, Villa Alvarez, Paso del Rio. 


UROSAURUS UNICUS (Mittleman) 


Uta unica MirrLeman, Journ. Washington Acad. Sci., vol. 31, 1941, pp. 74-76, 
fig. 3. 

Urosaurus unicus, MirrLEMAN, Bull. Mus. Comp. Zool., vol. 91, 1942, pp. 170- 
173.—Smiru and MirrLeman, Trans. Kansas Acad. Sci., vol. 46, 19438, pp. 
243-249. 

Type.—U.S.N.M. No. 14248; Edward Wilkinson collector. 
Type locality.—Chihuahua (Batopilas?). 
Range.—Known only from the type locality. 


Genus UTA Baird and Girard 


Uta Barrp and Grrarp, in Stansbury, Exploration and survey of the valley of the 
Great Salt Lake of Utah . . ., 1852, pp. 344-345. 


148 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Genotype.— Uta stansburiana Baird and Girard. 

Range.—Western Texas to California, north to Washington, south 
through Baja California and adjacent islands, and into the northern 
tier of Mexican states from Sonora to Coahuila. 

Species.—Nine are here listed, with a total of eleven forms; only one 
form (U. s. stansburiana) is extralimital, making a total for the genus 
of 12 forms. The entire genus is badly in need of careful revision, 
however; considerable shuffling of the nominal species and subspecies 
is to be expected eventually. 


10. 


KEY TO MEXICAN SPECIES OF UTA 


. Dorsal scales shorter, not imbricate, not mucronate, often with intervening 


oraniles-lardarksblotchybehind axillas: sess = as ae eee ee 2 
Dorsal scales larger, imbricate at least centrally, usually without intervening 
granules; caudal scales imbricate, keeled, and strongly mucronate._.._ 4 

. Scales on base of tail not imbricate; not unicolor above___-_----_-------- 3 


Scales on base of tail imbricate, strongly keeled and mucronate; no dark or 
light markings above, except rarely a few pale blue dots. 
nolascensis (p. 151) 


. Basal caudals weakly -keeled, not or but shortly mucronate; about 110-125 


dorsals from interparietal to back of thighs; 26-30 of largest dorsals equal 
length of head to back of interparietal plate; gular region bluish 

stellata (p. 150) 

Basal caudals keeled and strongly mucronate; about 106-116 dorsals from 

interparietal to backs of thighs; 21-25 of largest dorsals equal length of head 


to back of interparietal plate; gular region blackish_-_---- palmeri (p. 151) 

. Dorsal scales from interparietal to rear of thighs 70-81_.squamata (p. 149) 
Dorsalliscalest82ror sores Se sees a Seek o ae ae ep 5 

. No stripes whatever in dorsal pattern of either young or adults; dorsals 
92-117,’ generally’ more'than 1002-2222. 22. 28s. ot taylori (p. 150) 
Stripes present in young and some adults; dorsals larger or smaller_---_---- 6 

. No distinct dark blue blotch behind axilla__________-_ mannophorus (p. 149) 
A distinct. dark blue spot or blotch behind axilla_.____._.._..___._.-___--- ih 

. Two rows of postrostrals; both internasals separated by 2 scales from 
OSU oles ee see tee ees eee ee are a Tr ene rte wer eee ede ee 8 
One row of postrostrals; one or both internasals separated from rostral by 
onlysoneyscaless26) Si 2 pisses = fps ee eee yh. Se 10 
Dorsal scales generally 17-23 in a head length (from snout to rear edge of 
iInvéerparictalje LF Ree see Le eee ey ee A ETE Re area 9 
Dorsal scales generally 23-28 in a head length_stansburiana hesperis (p. 149) 
size larger (64 mm. snout to ‘vent)22.___2 282 22a i2s martinensis (p. 149) 
Sizersmaller’(48iinm-) aes Seen. sr eae ee he ie concinna (p. 150) 


Hind leg longer, 74-85 percent of body length__stansburiana elegans (p. 148) 
Hind leg shorter, 65-79 percent of body length. 
stansburiana stejnegeri (p. 150) 


UTA STANSBURIANA ELEGANS Yarrow 


Uta elegans Yarrow, Proc. U. S. Nat. Mus., vol. 5, 1882, p. 442.—Scumipt, 


Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 656-657. 


Uta stansburiana elegans, VAN DeNBURGH, Occ. Pap. California Acad. Sci., No. 


10, 1922, pp. 240-247 (part). 


CHECKLIST OF REPTILES OF MEXICO 149 


Type.—U.S.N.M. No. 12666. 

Type locality —La Paz, Baja California. 

Range.—Southern two-thirds of Baja California and on Smiths, 
Mejia, Angel dela Guarda, Isla Partida, Isla Raza,Sal Si Puedes, North 
San Lorenzo, South San Lorenzo, Tortuga, San Marcos, Ildefonso, 
East and West Las Galeras, Monserrate, San José, San Francisco, 
Espiritu Santo, and Ballena Islands. 


UTA MANNOPHORUS Dickerson 


Uta mannophorus DickERson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, pp. 
470—-471.—Van DeEnBurGH, Occ. Pap. California Acad. Sci., No. 10, 1922, 
pp. 252-254. 
Uta elegans, Scumipt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 656-657 
(part). 
Type.—U.S.N.M. No. 64260; C. H. Townsend collector. 
Type locality—Carmen Island, Gulf of California. 
Range.—Known only from type locality and Danzante and Coro- 
nado Islands, Baja California. 


UTA SQUAMATA Dickerson 


Uta squamata Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, p. 471.— 
Van DEnBURGH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 249-251.— 
Scumipt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, p. 656. 

Type.—U.S.N.M. No. 64259; C. H. Townsend collector. 
Type locality — Santa Catalina Island, Gulf of California. 
Range.—Known only from type locality. 


UTA STANSBURIANA HESPERIS Richardson 


Uta stansburiana hesperis RicHarpDsoN, Proc. U. 8. Nat. Mus., vol. 48, 1915, pp. 
415-418.— Van DeEnBuRGH, Occ. Pap. California Acad. Sci., No. 10, 1922, 
pp. 233-239.—ScumiptT, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, p. 655.— 
Smitu, Handbook of lizards, 1946, pp. 281-288, pl. 72. 
Uta parva Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, p. 471 
(U.S.N.M. No. 64258; San Bartolomé Bay, Baja California). 
Type.—Univ. California Mus. Vert. Zool. No. 892; J. Grinnell 
collector. 
Type locality—Arroyo Seco Cafion, near Pasadena, Los Angeles 
County, Calif. 
Range.—San Joaquin Valley of central California southward on the 
Pacific slope through northwestern Baja California to San Bartolomé 
Bay; Los Coronados Islands. 


UTA MARTINENSIS Van Denburgh 


Uta martinensis VAN DrENBURGH, Proc. California Acad. Sci., ser. 3, vol. 4, 
1905, pp. 18-19, pl. 6; Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 
247-249.—Scumipt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, p. 655. 


Type.—California Acad. Sci. No. 4698; R. H. Beck collector. 


150 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type locality—San Martin Island, Baja California. 
Range.—Known only from the type locality. 


UTA CONCINNA Dickerson 


Uta concinna DickErson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, p. 470.— 
Scumipt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 655-656. 
Type.—U.S.N.M. No. 64257; C. H. Townsend collector. 
Type locality—Cerros [= Cedros] Island, Baja California. 
Range.—Known only from the type locality, and the Natividad 
Islands, Baja California. 


UTA STELLATA Van Denburgh 


Uta stellata VAN DENBURGH, Proc. California Acad. Sci., Zool., ser. 3, vol. 5, 1905, 
pp. 21-22, pl. 8; Oce. Pap. California Acad. Sci., No. 10, 1922, pp. 225-227. 
Type.—California Acad. Sci. No. 4704, R. H. Beck collector. 
Type locality —San Benito Islands, Baja California. 
Range.—Known only from the type locality. 


UTA STANSBURIANA STEJNEGERI Schmidt 


Uta stansburiana stejnegeri Scumipt, Amer. Mus. Nov., No. 15, 1921, pp. 1-2; 
Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 654-655.—Smiru, Handbook 
of lizards, 1946, pp. 283-286, pl. 73. 

Uta stansburiana elegans, VAN Denpurau, Occ. Pap. California Acad. Sci., No. 
10, 1922, pp. 240-247 (part), pl. 18. 

Type.—Amer. Mus. Nat. Hist. No. 384; A. G. Ruthven collector. 

Type locality—Mouth of Dry Cafion, Alamogordo, Otero County, 
N. Mex. 

Range.—Western Texas through southern Nevada to eastern 
California, southward into northeastern Baja California and the 
northern tier of Mexican states from Sonora to western Coahuila. 
Reported from Baja California: Cocopah Mountains, Volcano Lake, 
etc.; Sonora: Costa Rica Ranch, Sonoyta, Pozo de Luis, Tepoca Bay, 
San Pedro Bay, Ortiz, Batamotal, Guaymas, Empalme; Islands of 
Tiburén, Patos, Pelican, and San Esteban; Coahuila: 10 miles east of 
Torreén, San Pedro, 5 miles south of San Pedro, 30 miles west of La 
Rosa, Jaral, Torreén; Durango: Avilco, 5 miles north of Conejos, 
25 miles north of Bermejillo, Lerdo; Chihuahua: 15 miles south of 
Juarez. 

UTA TAYLORI Smith ® 
Uta taylorit Smiru, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 158-166, pl. 26, fig. 3. 


Type.-—EHT-HMS No. 10692 (originally field No. 320a), male; E. 
H. Taylor collector. 
Type locality —Ten miles northwest of Guaymas, Sonora. 


88 The possibility of intergradation between Uta taylori and Uta stansburiana, as recently suggested, is 
too remote to consider. Ufa stansburiana extends throughout the known range of taylori. 


CHECKLIST OF REPTILES OF MEXICO 151 


Range-—Known only from the region near the type locality. 
Reported from Sonora: La Posa, 54 miles southwest of Hermosillo. 


UTA PALMERI Stejneger 


Uta palmeri STEJNEGER, North Amer. Fauna, No. 3, 1890, p. 106.—Van DEn~ 
BURGH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 221-223. 


Type.—U.S.N.M. No. 16002; Edward Palmer collector. 
Type locality—San Pedro Martir Island, Sonora. 
Range.—Known only from the type locality. 


UTA NOLASCENSIS Van Denburgh and Slevin 


Uta nolascensis VAN DENBURGH and S.teEvin, Proc. California Acad. Sci., ser. 
4, vol. 11, 1921, pp. 395-396.—Van DensBureu, Occ. Pap. California Acad. 
Sci., No. 10, 1922, pp. 223-225. 


Type.—California Acad. Sci. No. 50508; Joseph R. Slevin collector. 
Type locality San Pedro Nolasco Island, Sonora. 
Range.—Known only from the type locality. 


Family XANTUSIIDAE Baird 


Xantusiidae Barrp, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 254. 


Genera.—Four genera known; the only extralimital genus is Cri- 
cosaura of Cuba. 
Range.—Southwestern United States to Panama; Cuba. 


KEY TO MEXICAN GENERA OF THE XANTUSIIDAE 


1. Supraoculars present, as small scales above eyes (medial to superciliaries) ; 


dorsaliscales uniformly, granular ==) 22-2 == ==> = Xantusia (p. 154) 
Supraoculars absent, no small scales above eyes (except superciliaries) ; dorsal 
EcalestheterogcencOusMNisIZes sates | ate oe See ee eee eee 2 


2. Distinct, vertical rows of well-differentiated, enlarged, keeled scales on sides of 
body, separated from each other by granular area_._Lepidophyma (p. 151) 
No distinct, vertical rows of enlarged scales on sides of body_-Gaigeia (p. 153) 


Genus LEPIDOPHYMA A. Duméril 


Lepidophyma A. Dum£&RIL, in Duméril and Duméril, Catalogue méthodique de la 
collection des reptiles, 1851, pp. 137-138. 

Poriodogaster Smiru, tn Gray, Proc. Zool. Soc. London, 1863, p. 154 (type, 
P. grayit Smith). 

Akleistops Mituurr, Verh. Naturf. Ges. Basel, vol. 6, 1878, p. 390 (type, 
A. guatemalensis Miller). 


Genotype—Lepidophyma flavimaculatus A. Duméril. 

Range.—From Hidalgo, Mexico, south to Costa Rica. 

Species.—Two, with five forms, one of which (flavimaculata obscu- 
ra) is extralimital. 


152 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


KEY TO MEXICAN SPECIES OF LEPIDOPHYMA 


1. All (except basal) whorls of enlarged scales on tail separated from each other 
dorsally by 4 rows of scales; femoral pores 15 to 22; median prefrontal nor- 
mally present, sometimes absent___-flavimaculata flavimaculata (p. 152) 

All whorls of enlarged scales on tail separated from each other dorsally by 3 
rows of scales (rarely feeble evidence of a fourth row); femoral pores less 


than 14; median prefrontal present or absent._._____-_-_____-_-_---_ 2 
2. ‘Avmedian prefrontal’ 222222 2 eee smithii smithii (p. 152) 
Noimedian prefrontals eA o2 4a ees etn oP eae ee a yk eh ea 3 


8. Scales in posterior temporal region (anterior to ear) minute and very uniform 
in size, except for a series of relatively large, projecting auricular lobules and 

a row of larger scales beside the upper temporals; whorls on tail relatively 
little differentiated..220. 225323344 S44 535 See 2 smithii occulor (p. 153) 
Seales in posterior temporal region (anterior to ear) larger, irregular in size; 
auricular lobules poorly defined or absent; no scales bordering upper tem- 
porals; tail whorls strongly differentiated___-___- smithii tehuanae (p. 152) 


LEPIDOPHYMA FLAVIMACULATA FLAVIMACULATA A. Duméril 


Lepidophyma flavimaculatus A. Dum&riL, in Duméril and Duméril, Catalogue 
méthodique de la collection des reptiles (Paris Museum), 1851, pp. 1388-139. 

Lepidophyma flavomaculatum flavomaculatum, Smitu, Proc. U. S. Nat. Mus., vol. 
92, 1942, p. 379. 

Type.—Mus. Hist. Nat. Paris; P. M. A. Morelet collector. 

Type locality—Province de Petén, Guatemala, here restricted to 
Rio de la Pasidén. 

Range.—Atlantic slopes from southern Veracruz to British Hondu- 
ras, excluding the Yucatan Peninsula. Recorded in Mexico from 
Veracruz: Rio de las Playas; Tabasco: Rio de las Playas; Chiapas: 
across the Usumacinta River from Piedras Negras, Petén, Guatemala. 


LEPIDOPHYMA SMITHII SMITHII Bocourt 


Lepidophyma Smithii Bocourt, Journ. Zool., Paris, vol. 5, No. 5, 1876, pp. 
402-403; Mission scientifique au Mexique . . ., Etudes sur les reptiles, livr. 
5, 1878, pp. 309-312, pl. 20F, figs. 3-3b, and pl. 20G, figs. 2-2b. 

Lepidophyma smithii smithii, Smitu, Proc. U. 8. Nat. Mus., vol. 92, 1942, p. 380. 

Akleistops guatemalensis Miuuer, Verh. Naturf. Ges. Basel, vol. 6, 1878, pp. 
390-398, pls. 1, 2 (Nat. Mus. Basel; Mazatenango, Guatemala). 


Type.—Mus. Hist. Nat. Paris, six cotypes; ‘‘Commission Scien- 
tifique”’ collector.** 

Type locality —‘Tehuantepec, and west coast of Guatemala,’ 
stricted to ‘Guatemala,’ here restricted to Mazatenango. 

Range.—Pacific slopes of Chiapas and western Guatemala. Re- 
corded in Mexico only from Chiapas: La Esperanza near Escuintla. 


’ re- 


LEPIDOPHYMA SMITHII TEHUANAE Smith 


Lepidophyma smithii tehuanae Smiru, Proc. U. 8S. Nat. Mus., vol. 92, 1942, pp, 
377-378. 


84 On the basis of the figures given of the types, Smith believes they represent a population characteristic 
of Chiapas and Guatemala, and not of the region of Tehuantepec city. 


CHECKLIST OF REPTILES OF MEXICO 153 


Type.—U.S.N.M. No. 111488; H. M. Smith collector. 

Type locality—Cerro Arenal, 30 kilometers west of Tehuantepec, 
Oaxaca. 

Range.—Pacific slopes of Oaxaca and Chiapas (?) in the vicinity of 
the Isthmus of Tehuantepec. Reported from Tres Cruces, El Limén, 
Cerro Arenal, La Concepcién, Santa Efigenia, mountains near Santo 
Domingo, near Tehuantepec, Cafetal Concordia; ? Chiapas: Tonala. 


LEPIDOPHYMA SMITHIT CCCULOR Smith 


Lepidophyma smithit occulor Smiru, Proc. U. 8. Nat. Mus., vol. 92, 1942, pp. 
378-379. 

Type —U.S.N.M. No. 47133; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality —Jalpan, Querétaro. 

Range.—Atlantic slopes presumably from northern Veracruz into 
southern Tamaulipas. Recorded only from San Luis Potosi: Tama- 
zunchale; Querétaro: Jalpan. 


Genus GAIGEIA Smith 


Gaigeia SmitH, Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, 1939, p. 24. 


Genotype —Lepidophyma gaigeae Mosauer. 
Range.—Hidalgo and Oaxaca. 
Species.—Four. 


KEY TO MEXICAN SPECIES OF GAIGEIA 


1. All whorls on tail complete, none restricted to dorsal surface_gaigeae (p. 154) 
Some of scale whorls on tail restricted to dorsal surface________________~_ 2 
2. Only one row of scales on ventral surface between whorls of enlarged scales 
(i. e., every third whorl restricted to dorsal surface)_.._.._..----_-__- 3 
Proximally 2 and distally 3 rows of scales on ventral surface between whorls 
Gtaenlargeduscaless—= 22. 32 ea eae eae os See ea sylvatica (p. 154) 
3. Numerous, very closely approximated, enlarged, keeled scales, separated by 
small granules, present on sides of body; 2 or 3 rows of granules in vertebral 
SERN cig gn te a ch Rat as Ny NS radula (p. 153) 
Dorsal scales practically uniform in size; 4 rows of granules in vertebral region. 
dontomasi (p. 153) 

GAIGEIA DONTOMASI Smith 


Gaigeia dontomasi Smitu, Proc. U. 8. Nat. Mus., vol. 92, 1942, pp. 374-376. 


Type.—U.S.N.M. No. 111473; Thomas MacDougall collector. 
Type locality —Lachiguiri, Oaxaca, 7,100 feet elevation. 
Range.—Known only from the type locality. 


GAIGEIA RADULA Smith 


Gaigeia radula Smitu, Proc. U. 8. Nat. Mus., vol. 92, 1942, pp. 378-379. 
Type —U.S.N.M. No. 111472; H. M. Smith collector. 


154 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type locality—San José Manteca, 5 kilometers from San Carlos 


Yautepec, Oaxaca. 
Range.—Known only from the type locality. 


GAIGEIA SYLVATICA (Taylor) 


Lepidophyma sylvatica TayLor, Copeia, 1939, pp. 131-138, figs. 1, 2. 
Gaigeia sylvatica, Smitu, Proc. U. 8. Nat. Mus., vol. 92, 1942, p. 380. 
Type -—EHT-HMS No. 16259; E. H. Taylor collector. 
Type locality —Seven miles north of Zacualtipan, Hidalgo. 
Range.—Known only from the type locality. 


GAIGEIA GAIGEAE (Mosauer) 


Lepidophyma gaigeae MosavurEr, Herpetologica, vol. 1, 1936, pp. 3-5, pl. 2. 
Gaigeia gaigeae, SmitTuH, Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, 1939, p. 24. 
Type.—Mus. Comp. Zool. No. 42145; Walter Mosauer collector. 

Type locality —Durango, Hidalgo. 
Range.—The immediate vicinity of the type locality. 


Genus XANTUSIA Baird 


Xantusia Barrp, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 255. 
Zablepsis Corr, Amer. Nat., vol. 29, 1895, p. 758 (type, Xantusta henshawi Stejne- 


ger). 
Amoebopsis Corr, Amer. Nat., vol. 29, 1895, p. 758 (type, Xantusia gilberti Van 
Denburgh). 
Genotype.—Xantusia vigilis Baird. 
Range.—Southern California, Arizona, southern Utah, Baja Cali- 


fornia, and adjacent islands. 
Species.—Five, one of which includes two subspecies; three species 


occur in Mexico. 


KEY TO MEXICAN SPECIES OF XANTUSIA 


DiVienttralipol ates? my 2 Ser Cs as te aes ee hae ne ae om ore 2 
Ventralyplates in 04 seriess 2. Oo So oe ee eee henshawi (p. 155) 
2A single troutal- eye large = arn. ee ee” eae 2 ek eee vigilis (p. 154) 
A pair. on srontals, (eve eimall 2 = 5 es ee ee gilberti (p. 155) 


XANTUSIA VIGILIS Baird 


Xantusia vigilis Batrp, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 255.—Van 
DenxsuRGH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 477-482, pl. 
49.—Smitu, Handbook of lizards, 1946, pp. 330-333, pl. 91. 

Type—U.S.N.M. No. 3063 (three cotypes); John Xantus collector. 

Type locality —‘ Fort Tejon,” California. 

Range.—Southwestern Utah and eastern California southward into 
northern Baja California as far as San Felipe Bay and San Matias Pass. 


CHECKLIST OF REPTILES OF MEXICO 155 


XANTUSIA HENSHAWI Stejneger 


Xantusia henshawi STEJNEGER, Proc. U. 8S. Nat. Mus., vol. 16, 1893, p. 467.— 
Van Denzsuray, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 484-486, 
pl. 50.—Smiru, Handbook of lizards, 1946, pp. 325-327, pl. 88. 
Xantusia picta Corr, Amer. Nat., vol. 29, 1895, pp. 859, 939 (Tejon Pass, Calif.; 
probably Poway, San Diego County, Calif., fide Van Denburgh). 
Type.—U.S.N.M. No. 20339; H. W. Henshaw collector. 
Type locality——Witch Creek, San Diego County, Calif., elevation 
2,700 feet. 
Range.—Extreme southwestern California and northern Baja Calif- 
fornia, in the San Pedro M&rtir Mountains. 


XANTUSIA GILBERTI Van Denburgh 


Xantusia gilberti VAN DENBURGH, Proc. California Acad. Sci., vol. 5, 1895, p. 121, 
pl. 11; Oce. Pap. California Acad. Sci., No. 10, 1922, pp. 482-484. 
Type.—California Acad. Sci. No. 401; Gustav Eisen collector. 
Type locality —San Francisquito, Sierra Laguna, Baja California. 
Range.—Cape region of southern Baja California. 


Family SCINCIDAE Gray 
Scincidae Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 201. 


Genera.—About 60, of which three occur in Mexico. Of these three, 
none are confined to the Western Hemisphere. Only two other genera 
occur in the Americas: Cryptoblepharus (South America), a wanderer 
from the Pacific Islands, and Neoseps (Florida). 

Range.—Southern Canada south through Central America and 
South America to Argentina; West Indies; Africa and Madagascar; 
south Asia; China; Japan; Philippine Islands; Dutch Indies; Pacific 
Islands and Australia. 


KEY TO MEXICAN GENERA OF SCINCIDAE 


1. Two supranasals between rostral and frontonasal; frontoparietal always di- 
vided; lower eyelid with a translucent disk or not; palatine bones in contact 
OTsnotseeew So ee ee ee SSeS Bee eo ce eee 2 

Supranasals absent, the single frontonasal in contact with rostral; frontoparietal 
divided or not; an undivided translucent disk on lower eyelid; palatine bones 
ANSCONLAC EON DAGC Tsp lire eee ee eet yar ee ra Scincella (p. 156) 

2. An undivided translucent disk on lower eyelid; palatine bones in contact on 

median: line. —- ee ee ee ees Sate Mabuya (p. 155) 
Lower eyelid without a translucent disk, or if so it is divided into several parts; 
palatine bones separated on median line___------------- Eumeces (p. 160) 


Genus MABUYA Fitzinger 
Mabuya Fitzincrer, Neue Classification der Reptilien, 1826, pp. 23, 52. 


Genotype.—Lacertus mabouya Lacépéde (by absolute tautonymy, 
fide M. A. Smith, Fauna British India, etc., 1935, Reptilia and Am- 
861316—50——11 


156 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


phibia, vol. 2, p. 257). This is contrary to opinion 92 of the Inter- 
national Commission on Zoological Nomenclature, which states that 
Scincus sloanii of Daudin is the type. Fitzinger did not list that form.* 
Range.—Southern Mexico on both coasts; Central America, South 
America; Africa, Madagascar; southern Asia, Philippines, East 
Indies. 
Species.—About 85 species, only one of which occurs in Mexico. 


MABUYA MABOUYA MABOUYA (Lacépéde) 


Lacertus Mabouya Lacf&ripe, Histoire naturelle des quadrupédes ovipares et des 
serpens, vol. 2, 1788, p. 378, pl. 24 (part). 

Mabuya mabouya mabouya, Dunn, Proc. Acad. Nat. Sci. Philadelphia, vol. 87, 
1935, pp. 537-546 (part). 

Scincus agilis Rapp1, Mem. Soc. Ital. Modena, vol. 19, No. 18, 1823, p. 62 (type 
unknown; Rio de Janeiro). 

Mabuia alliacea Corr, Journ. Acad. Nat. Sci. Philadelphia, ser. 2, vol. 8, 1875, 
p. 115, pl. 6, fig. 1 (U. 8S. N. M. Nos. 30619-20: Costa Rica) .% 

Type.—Apparently unknown. 

Type locality.—Restricted by Dunn to the Lesser Antilles; here 
restricted to St. Vincent. 

Range.—Coastal and foothill areas from southern Veracruz and 
Colima southward to Ecuador, Bolivia, and southern Brazil. Re- 
corded from numerous localities in the states of Colima, Michoacan, 
Guerrero, Oaxaca, Veracruz, Tabasco, Campeche, Yucatén, Quintana 
Roo (Cob4), and Chiapas. 


Genus SCINCELLA Mittleman 
Leiolopisma Avct. in part, nec Duméril and Bibron, Erpétologie générale, vol. 5, 
1839, p. 742 (type, Scincus telfairii Desjardines). 
Scincella ** MirrLeEMAN, Herpetologica, vol. 6, 1950, p. 19. 

Genotype.—Scincus lateralis Say. 

Range.—Greater part of eastern United States east of the Rocky 
Mountains; Mexico from Hidalgo southward through Central America 
to Panama. Ceylon, southern Asia, China, Philippines, East Indies, 
Tasmania, Africa, New Zealand. 

Species.—About 50 species. Seven species and 10 forms are known 
in the Americas, all except S. incertum of Guatemala occurring in 
Mexico (even this exception may actually occur). 

KEY TO MEXICAN SPECIES OF SCINCELLA 
1. Frontoparietal divided; many median subcaudals in contact on each side with 


SCO he ee eee eh a ea a 2 od Ss ay re 2 
Frontoparietal single; median subcaudals in contact with only 1 scale on 
Ped SU (0 = dey lM YR, |G eh PR A UN a IT AE a ol a 6 


8 Also see Travassos, Bol. Mus. Nac. Brasil, new ser., Zoo]., No. 37, 1945, pp. 1-7. 

86 This is the only name based upon continental (as opposed to island) specimens north of South America. 
It may be revived, for undoubtedly geographically segregated variants of taxonomic magnitude do exist 
within the enormous range now included within this supposedly single race. 

87 Adoption here of this name is the decision of solely the senior author. 


CHECKLIST OF REPTILES OF MEXICO 157 


2. Usually 3 or more pairs of nuchals; upper tertiary temporal (occasionally 
split) in contact with parietal; 26-28 scale rows around middle of body; 
limbs short, separated by 10-12 scales when adpressed__-_laterale (p. 159) 

Two pairs of nuchals, or fewer; upper tertiary temporal separated from parietal 
by contact of upper secondary temporal and nuchal_______________-_- 3 

3. Limbs, when adpressed, touching or overlapping in adults, forelimb reaching 

rear corner of eye; scale rows at middle of body often 30 or 32_______- 4 
Limbs, when adpressed, separated from each other by 1 or more scale lengths; 
scale rows at middle of body rarely 30 and apparently never 32_______-_ 5 

4, “Usually 1 pair of nuchals or less; lamellae under fourth toe 19 or fewer; 

dorsolateral light stripe continuous”. (Smith)----- caudaequinae (p. 158) 
Usually 2 pairs of nuchals; lamellae under fourth toe 20 (constant?); dorso- 
lateral light stripe dark-spotted, interrupted____---_-- silvicolum (p. 158) 

5. Usually nuchals on the two sides 1-2 or more (89 percent); lateral light line 
along head and neck poorly defined; axilla-groin/snout-vent percentage 59; 
65 mm. maximum snout-vent measurement. 

gemmingeri gemmingeri (p. 159) 
Usually nuchals on the two sides 1-1 or less (88 percent); lateral light line 
along head and neck fairly well defined; axilla-groin/snout-vent percentage 

56; 54 mm. maximum snout-vent measurement. 
gemmingeri forbesorum (p. 159) 
GeSCuesrOws i UrOr more: Uva hols ee hs te SPS eee 7 
DEAE LOWS IES tNAn GOs. 20 2h eee Be etl oe ee ee rN 9 

7. Tail blue or blue-gray, with traces of a cross-banded pattern evidenced by 
feeble dark bars visible laterally or by transverse series of light spots; legs 
longer aisuallyvoverlappinge invadults- === ===" 9s = eee ee se ee 8 

Tail pink or reddish, with no traces of a cross-banded pattern, but instead with 

a longitudinal dark line on each side at least at base; legs shorter, never over- 
lapping in adults; dorsal scales 65 to 79, usually 69 or more. 

assatum assatum (p. 160) 

8. Dorsal scales 65 to 72, average 69.1_____.___-_--_--- cherriei stuarti (p. 158) 

Dorsal scales 59 to 67, average 63.2_____-_-------- cherriei cherriei (p. 157) 

9. Tail blue or blue-gray, with traces of a cross-banded pattern evidenced by 
feeble dark bars visible laterally or by transverse series of light spots; legs 
longer, usually overlapping in adults; dorsal scales 54 to 60, average 57. 

cherriei ixbaac (p. 158) 

Tail pink or reddish, with no traces of a cross-banded pattern, but instead with 

a longitudinal dark line on each side at least at base; legs shorter, never 
overlapping in adults; dorsal scales 58 to 72, average over 66. 

assatum taylori (p. 160) 


SCINCELLA CHERRIEI CHERRIEI (Cope) 


Mocoa cherriet Corn, Proc. Amer. Philos. Soc., vol. 31, 1893, p. 339. 

Lygosoma assatum cherriei, Stuart, Occ. Pap. Mus. Zool. Univ. Michigan, No. 
421, 1940, pp. 13-14. 

Lygosoma cherriei cherriet, SMiru, Proc. Biol. Soc. Washington, vol. 54, 1941, pp. 
181-182. 

Leiolopisma cherriei cherriei, SmirH, Herpetologica, vol. 3, 1946, p. 111. 

Scincella cherriet cherriei, MiTTLEMAN, Herpetologica, vol. 6, 1950, p. 20. 


Type—Amer. Mus. Nat. Hist. No. 9531; George K. Cherrie 
collector. 
Type locality — Palmar, Costa Rica. 


158 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Range.—Tabasco and northern Chiapas on Atlantic slopes, and 
Costa Rica on Pacific slopes, eastward to Panama. Recorded in 
Mexico only from Tabasco: Teapa; Chiapas: Palenque. 


SCINCELLA CHERRIEI IXBAAC (Stuart) 


Lygosoma assatum izbaac Stuart, Occ. Pap. Mus. Zool. Univ. Michigan, No. 421, 
1940, pp. 8-10. 

Leiolopisma cherriei ixbaac, Smitu, Herpetologica, vol. 3, 1946, p. 111. 

Scincella cherriet txbaac, MirrLEMAN, Herpetologica, vol. 6, 1950, p. 20. 


Type-—Univ. Michigan Mus. Zool. No. 80820; Milton Trautman. 

Type locality —Chichen Itz4, Yucatan. 

Range.—The peninsula of Yucatan, southward as far as Campeche 
and northern Petén, Guatemala. Recorded in Mexico only from 
Yucatdn: Chichen Itzi; Campeche: Tres Brazos. 


SCINCELLA CHERRIEI STUARTI (Smith) 
Leiolopisma cherriet, Smita, Proc. Biol. Soc. Washington, vol. 52, 1939, pp. 
191-192, 193, pl. 2, fig. 7. 


Lygosoma cherriet stuarti Smits, Proc. Biol. Soc. Washington, vol. 54, 1941, 
pp. 81-82. 


Leiolopisma cherriet stuarti, Smita, Herpetologica, vol. 3, 1946, p. 111. 
Scincella cherriet stuarti, MirrTLEMAN, Herpetologica, vol. 6, 1950, p. 20. 

Type —U.S.N.M. No. 115174; H. M. Smith collector. 

Type locality —Potrero Viejo, Veracruz. 

Range.—Central Veracruz, in foothills, southward to the Isthmus 
of Tehuantepec; recorded only from the states of Veracruz and 
Oaxaca (between Rio Grande and La Gloria). 


SCINCELLA CAUDAEQUINAE (Smith) 
Leiolopisma caudaequinae Smitu, Univ. Kansas Sci. Bull., vol. 34, 1950, p. —. 


Type—Univ. Illinois Mus. Nat. Hist. No. 10131; J. P. Craig. 

Type locality.—Salto Cola de Caballo, 25 miles south of Monterrey, 
Nuevo Leén. 

Range.—Eastern foothills from central Nuevo Leén southward to 
southern San Luis Potosi (known only from the type locality and 10 
miles west of Naranjo, San Luis Potosi). 


SCINCELLA SILVICOLA (Taylor) 


Leiopisma silvicolum Taytor, Copeia, 1937, No. 1, pp. 5-7.—Smiru, Journ. 
Washington Acad. Sci., vol. 39, 1949, p. 41; Univ. Kansas Sei. Bull., vol. 
34, 1950, p. —. 

Scincella silvicola, MirrLemMan, Herpetologica, vol. 6, 1950, p. 20. 


Type—EKHT-HMS No. 10083, adult female; E. H. Taylor collector. 


Type locality Forested hill about 10 miles southeast of Cérdoba, 
near San Lorenzo, Veracruz. 


CHECKLIST OF REPTILES OF MEXICO 159 





Range.—Atlantic slopes in central Veracruz and in the headwaters 
of the Rio Papaloapam in Oaxaca. Recorded from Veracruz: San 
José de Gracia; Oaxaca: Cuicatlin. 


SCINCELLA GEMMINGERI GEMMINGERI (Cope) 


Oligosoma gemmingeri Corr, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 180. 

Lygosoma (Mocoa) Gemmingeri, Bocourt, Mission scientifique au Mexique.. . 
Etudes sur les reptiles, livr. 7, 1881, pp. 449-450. 

Leiolopisma gemmingeri, TayLor, Copeia, 1937, pp. 7-8.—Smiru, Journ. Washing- 
ton Acad. Sci., vol. 39, 1949, pp. 40, 41. 

Leiolopisma gemmingert gemmingeri, SmitH, Univ. Kansas Sci. Bull, vol. 34, 
1950, p. —. 

Lygosoma aeke (part), BouLENGER, Catalogue of the lizards in the British 
Museum, vol. 3, 1887, p. 263. 

Type—U.S.N.M. No. 6331, four cotypes; Francois Sumichrast 
collector. 

Type locality —Orizaba, Veracruz. 

Range.—Eastern Hidalgo, central and southern Veracruz, and 
Oaxaca south to Tehuantepec, on the slopes of the plateau and in 
lowlands. Recorded only from Veracruz: Tequeyutepec, La Perla, 
Rio Verde, Jalapa, Orizaba; Hidalgo; Zacualtipan (possibly referable 
to L. g. forbesorum); Oaxaca: Ixcuintepec, Tres Cruces, San José 
Manteca, 17 miles north of Niltepec, Cafetal Concordia. 


SCINCELLA GEMMINGERI FORBESORUM (Taylor) 


Leiolopisma forbesorum Taytor, Copeia, 1937, pp. 8-11. 
Leiolopisma gemmingeri forbesorum, SmitH, Univ. Kansas Sci. Bull., vol. 34, 
1950, p. —. 

Type —EHT-HMS No. 10043, E. H. Taylor collector. 

Type locality —La Placita, Hidalgo, 8 miles south of Jacala, elevation 
7,000 feet. 

Range—Known only from higher mountains of Hidalgo, in the 
vicinity of the type locality. 


SCINCELLA LATERALE (Say) 


Scincus laieralis Say, in Long’s Expedition to the Rocky Mountains, vol. 2, 1823, 
. 234. 
Tehibkevia laterale, JoRDAN, Manual of the vertebrates of the northern United 
States, ed. 8, 1899, p. 324.—Cors, Ann. Rep. U.S. Nat. Mus., 1898 (1900), 
pp. 622-624, fig. 123.—Smiru, Handbook of lizards, 1946, pp. 337-340, pl. 92. 
Scincella laterale, MirrLEMAN, Herpetologica, vol. 6, 1950, p. 19. 
Type-—U.S.N.M. No. 3152. 
Type locality —Mississippi River, below Cape Girardeau, Mo. 
Range.—New Jersey and southward to the Gulf and to central 
Coahuila, excluding the lower Rio Grande Valley. Reported from 
Coahuila: Nogales (Sabinas River near Mtzquiz). 


160 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


SCINCELLA ASSATA ASSATA (Cope) 


Lampropholis assatus Corn, Proc. Acad. Nat. Sci. Philadelphia, 1864, pp. 179-180. 
Leiolopisma assatum, Burt, Trans. American Micr. Soc., vol. 54, 1935, p. 177. 
Leiolopisma assatum [assatum], OuIveR, Occ. Pap. Mus. Zool. Univ. Michigan, 
No. 360, 1937, p. 12.—Smirn, Herpetologica, vol. 3, 1946, p. 111; Journ. 
Washington Acad. Sci., vol. 39, 1949, p. 40. 
Lygosoma assatum assatum, Stuart, Occ. Pap. Mus. Zool. Univ. Michigan, 
No. 421, 1940, pp. 12-13. 
Scincella assata assata, MirrLeMAN, Herpetologica, vol. 6, 1950, p. 20. 
Type.—Acad. Nat. Sci. Philadelphia No. 9465. 
Type locality.—Volcan Isalco, El Salvador. 
Range.—Pacific slopes, southeastern Chiapas southward to Hon- 
duras. Recorded in Mexico from Chiapas: Huehuetan, Escuintla. 


SCINCELLA ASSATA TAYLORI (Oliver) 


Leiolopisma assatum taylori OtivER, Occ. Pap. Mus. Zool. Univ. Michigan, No. 
360, 1937, pp. 12-15.—Smiru, Journ. Washington Acad. Sci., vol. 39, 1949, 
. 40. 
Scineclla assata taylort, MirrLEMAN, Herpetologica, vol. 6, 1950, p. 20. 

Type.—Mus. Zool. Univ. Michigan No. 80107; A. Bakewell and 
J. A. Oliver collectors. 

Type locality Santiago, Colima. 

Range.—Colima to southwestern Chiapas, in lower areas. Recorded 
from Colima: Santiago, Paso del Rio; Guerrero: Between Rincén and 
Cajones, Mazatlan, Chilpancingo, Agua del Obispo, Tierra Colorada; 
Jalisco: Tenacatita; Chiapas: Tonala; Oaxaca: Matias Romero, Santa 
Efigenia, Tapanatepec. 


Genus EUMECES Wiegmann 


Eumeces W1EGMANN, Herpetologia Mexicana, 1834, pt. 1, p. 36. 

Lamprosaurus HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, 1852, p. 206 
(type, Lamprosaurus guttulatus Hallowell=Plestiodon obsoletum Baird and 
Girard). 

Platypholis Ducts, La Naturaleza, ser. 2, vol. 1, 1887, p. 486 (type, Eumeces 
altamirani Dugés). 

Genotype.—Scincus pavimentatus Geoffroy=Eumeces pavimentatus 

(see Wiegmann, Archiv fiir Naturg., vol. 2, 1835, p. 288). 

Range.—Southern Canada southward through the United States, 
Mexico, and Central America to Nicaragua; Bermuda; North Africa; 
southwestern Asia; China; French Indo-China; Japan. 

Species.—About 58 species and 76 forms are recognized; 23 forms 
of 22 different species occur in Mexico. 


KEY TO MEXICAN SPECIES OF EUMECES 


1. Median dorsal scales at least three times the width of adjoining scales__. 2 
Dorsal scales not greatly widened, rarely twice as wide as adjoining scales___ 3 

2. Scales in 21 rows about middle of body; 3 broad black stripes begin on snout, 
pass back and break up on middle of back__________- schwartzei (p. 162) 


10. 


1. 


12. 


13. 


14. 


CHECKLIST OF REPTILES OF MEXICO 161 


Scales in 19 rows about middle of body; no stripes, but scales with black 


LO tis ee ae ee ne dae gD ee eo altamirani (p. 163) 
. Three supraoculars; no postnasal; 1 postmental_____.._---------------- 4 
Four supraoculars; postnasals and postmentals variable____-_---------- 5 


. Parietals enclose interparietal; scale rows about middle of body, 24; seventh 


labial in contact with upper secondary temporal______-__- dugesii (p. 169) 
Parietals not enclosing interparietal; scale rows usually 24; seventh labial 
not in contact with upper secondary temporal__lynxe furcirostris (p. 163) 


aePostmentalssnormallyic29-- he sos oe a ds Pe ok eee 6 


Postmentalsnorma lily, dy wes fon es eels ses eee a eae Pee 12 


. Body without white stripes at any time; lateral scale rows generally oblique; 


young black with a blue tail; head with cream spots; adults olive with 
darker areas on scales; limbs large, overlapping when adpressed; scales 
26-26 LOWS.» 125mm. snout) tO Venten==- ==225-25es225 obsoletus (p. 165) 
Body with white stripes in young, retained in adult, or approaching uniform 
olive coloration in adults, especially old males; lateral scale rows parallel; 
young never black but usually with blue or pink (red) or orange tail_-__ 7 


. Dorsolateral light stripes distinct and confined to third scale row. 


multivirgatus (p. 165) 

Dorsolateral light stripes not involving third scale row at all, or else including 
also, sccond scales 0 wi--es 4 a et en ee 8 
Median light line beginning between arms running forward bifurcating on 
head; or median line lost and the bifurcating lines only, remain on head. 9 
No trace of a median light line or bifurcating lines on head; usually 4 light 
lines; 2plateralands2 dorsolateraless= == == =e eee eee 10 
Parietals enclose interparietal; scale rows usually 28 at middle of body; post- 
nasal usually present; median white line to shoulder; limbs short, not 
TOUCHINe Swhentad pressedssss= =e eee ae eae a callicephalus (p. 164) 
Parietals not enclosing interparietal; bifurcating lines on head, but median 
line obsolete; postnasal absent; limbs touch in young when adpressed, in 
adults separated by 3 or 4 scale lengths; postmental single or divided; 26-28 
scale rows; Maximum size about 70 mm_____---_-- tetragrammus (p. 165) 
Parietals enclose interparietal; 24 scale rows; seventh labial broadly in contact 
with upper secondary temporal; tail orange in young; 4 light lines retained 
in adults; limbs fail to touch when adpressed__-_------- lagunensis (p. 167) 
Parietalsido.moteencloselimterpanietale sss ase = see ee oes et 


Snout-vent length greater than 75 mm., or interparietal nearly parallel- 
sided; specimens under 50 mm. snout-vent length with pink tail (no blue) ; 
8 supralabials on each side (98 percent) ___gilberti rubricaudatus (p. 167) 
Snout-vent length not more than 75 mm.; interparietal bluntly wedge-shaped; 
juveniles, usually adults, with blue tail; 7 supralabials on one or both sides 
(G4. DeLCenht) a= seo ka aes = ee ee ee skiltonianus (p. 167) 


Aemedian light line biturcatineom frontalea== == ee ee 13 
Median light line present or absent, if present bifurecating behind frontal. 14 
Median line from middle of body bordered with darker, and bifurcating on 
anterior half of frontal; limbs separated when adpressed; usually 24 scales 
BbOl bs DOGyA — 2 eee Oe NE es ee te lynxe lynxe (p. 163) 
Median light line bifurcating on the posterior part of frontal, or all lines lost 
in adult olive coloration; limbs overlapping when adpressed; subcaudals 
widened; maximum snout to vent length, 100 mm___sumichrasti (p. 164) 


Parietalsienckasing: interparietalusics 2h oi. . ls Sa See oe eee ee 15 
iParietals omene Oslin INverparietal.. 922-6 ee eee ee a 17 


162 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


15. Limbs long, overlapping when adpressed; strong, wide dorsolateral light lines; 
seventh labial broadly in contact with secondary upper labial; no primary 
temporal; frontoparietals narrowly separated; scale rows, 28; snout to 
vent,-65.mm2 oe See eB Se ata BA ee colimensis (p. 169) 

Limbs shorter, separated when adpressed_____-______---------______- 16 

16. Seventh labial not touching upper secondary temporal; primary temporal as 
large as upper secondary temporal; 24 scale rows; maximum length snout to 
Vent toil mins ete Ue Je a ee eee ee parvulus (p. 166) 

Seventh labial broadly in contact with upper secondary temporal; primary 
temporal present, smaller than upper secondary temporal; 22—24 scale rows 
around middle of body; maximum length snout to vent, 66 mm. 

indubitus (p. 168) 

17. Primary temporal absent; limbs short, not touching when adpressed; seventh 
labial in contact with upper secondary temporal; 22 scale rows; dorsolateral 
linés growing dim posterior] y240 97 ae oo ee dicei (p. 169) 

Primary temporal present! 10 Bho Vieicag sy .eUP 8 Te Bee sen 18 

18. Four narrow dark dorsal lines and narrow dorsolateral light lines; 22-24 
scale rows; no postnasal; limbs widely separated when adpressed; maximum 
snout moosventslength«Gnmin ae ay Se ee ee eee copei (p. 166) 

Dorsalpattern' not of 4 narrow black lines!: 222. 225205 Seas. Lees eres 19 

19. Bifurcating lines on head; lateral and dorsolateral light lines rarely reaching 
farther back than arm; scale rows, 26-28; maximum snout to vent length, 
GG rar SE A BEET ER LET EST ES EN ANE brevilineatus (p. 164) 

INo biftrcating dines on"head iso ith, Fo _Dehhh ose he ENe DA Seeht 7 20 

20. A postnasal; usually bronze olive, with lateral brown stripe, with sometimes 
trace of a dorsolateral light line; 26 scale rows; the limbs adpressed, 
the toes*may orimay not.‘touch 22 22246" 22 ee humilis (p. 166) 

ING! POStISSA ING os oy ne ne os ae tee AERO ST Rs 21 

21. Scale rows 20 around body; ear small, median dorsal scales widened. 

parviauriculatus (p. 166) 

Scaletrowse MorelthanltZO was FAR Ue IO DL Onsen es ie Me 22 

22. Scale rows 22 (rarely 24); body slender, dorsolateral lines distinct to tail; 
seventh labial usually touching upper secondary temporal; snout to vent 
SGhamnaik Pees Ae ee EE See Be ee RE a ochoterenai (p. 169) 

Seale rows usually 24, rarely 22; dorsolateral lines to tail or not; parietal 
usually not enclosed but occasionally may be enclosed, variable. 

brevirostris (p. 168) 
EUMECES SCHWARTZEI Fischer 

Eumeces schwartzet Fiscuer, Abh. Nat. Ver. Hamburg, vol. 8, 1884, pp. 3-5, 

pl. 7, fig. 1.—Tayuor, Uniy. Kansas Sci. Bull., vol. 23, pt. 1, 1935 (1936), 

pp: 94-101, figs. 5,6, pl. 1. 


Type.—Naturh. Mus., Hamburg, Germany.® 

Type locality —‘Einer kleinen Insel in der Laguna de Términos 
(Campeche Bai).”’ 

Range.-—Campeche, Tabasco, Yucat4én; in Central America it is 
known from Guatemala and British Honduras. Reported from 
Campeche: Tres Brazos, Encarnacién, Isla de Carmen; Tabasco: 
Tenosique; Yucatdn: Chichen Itza. 

88 This specimen was actually taken aboard ship in a cargo of dyewood and collected on board ship, alive, 


collector unknown. It was sent to the Zoologischen Garten in Hamburg. At its death it was sent to the 
Naturhistorischen Museum of Hamburg. 


CHECKLIST OF REPTILES OF MEXICO 163 


EUMECES ALTAMIRANI Dugés 

Eumeces aliamirani Ducts, La Naturaleza, ser. 2, vol. 1, 1887-1890 (1891), 
pp. 485, 486, pl. 22, 6 figs —Taytor, Proc. Biol. Soc. Washington, vol. 49, 
1936, pp. 55-58; Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 102-108, 
fig. 6, pl. 2—Smiru and Necker, Anal. Esc. Nac. Cience. Biol., vol. 3, 1943, 
pp. 190-192. 

Type—In Museo “Alfredo Dugés’” in Colegio del Estado de 
Guanajuato in Guanajuato (without number); Federico Altamirano 
collector. 

Type locality—Regiones CA4lidas del Estado de Michoac4n” 
(later designated by Dugés as Apatzing4n de la Constitucién, Mich- 
oacan). 

Range.—Known only from Michoacdn, on the southern slopes of 
the plateau. Reported from El Sabino, Apatzingan. 


EUMECES LYNXE LYNXE (Wiegmann) 

Scincus quinquelineatus var., W1EGMANN, Isis von Oken, 1828, p. 373 (non 
Linnaeus). 

Plestiodon quinquelineatum, Dum&rRit and Brsron, Erpétologie générale, vol. 5, 
1839, pp. 707-708 (part). 

Euprepes lynze W1EGMANN, Herpetologia Mexicana, 1834, pp. 36-387. 

Eumeces lynxe, Peters, Monatsb. Akad. Wiss. Berlin, 1864, p. 484.—Smirs, 
Journ. Washington Acad. Sci., vol. 39, 1949, p. 41. 

Eumeces lynxe lynxe, Taytor, Univ. Kansas Sci. Bull., vol. 23, pt. 1, 1935 (1936), 
pp. 163-173, pl. 41, figs. 18, 19, fig. B, 

Piestiodon Bellit Gray, Catalogue of the specimens of lizards in the collection of 
the British Museum, 1845, p. 92 (type locality not designated, here restricted 
to El Chico, Hidalgo). 

Type.—Zool. Mus., Berlin; F. Deppe coilector. 

Type locality —‘Specimena nostra prope Chico invenit Deppe.” 
Very probably (and here restricted to) El Chico (or Mineral £1 Chico) 
near Pachuca, Hidalgo, where the species has been found to be abun- 
dant. 

Range-—Known definitely from Hidalgo, Veracruz, San Luis 
Potosi and Puebla; recorded, probably erroneously, from Guerrero, 
Guanajuato and Michoacan. Reported from Hidalgo: Zacualtipan, 
San Miguel, Guerrero, Durango, El Chico; Veracruz: Jalapa, Mount 
Orizaba; San Luis Potost: Alvarez; Puebla: Zacatlan. 


EUMECES LYNXE FURCIROSTRIS Cope 
Eumeces furcirostris Corn, Proc. American Philos. Soc., vol. 22, 1885, pp. 169— 
170.—Taytor, Univ. Kansas Sci. Bull., vol. 23, pt. 1, 1935 (1936), pp. 173- 
178, figs. 18, 20; Univ. Kansas Sci. Bull., vol. 29, 1948, pp. 272-273. 
Type.—Acad. Nat. Sci. Philadelphia No. 11327; Dr. Flohr collector. 
Type locality.—Jalapa, Veracruz. 
Range.—Puebla, central northern Veracruz, and possibly eastern 
Hidalgo. Reported from Puebla; Tezuitlan; Veracruz: Toxtlacuaya, 
Jalapa, Las Vigas, Cofre de Perote; Hidalgo: Zacualtipan. 


164 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 
EUMECES SUMICHRASTI (Cope) 


Plistodon sumichrasti Corr, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 321. 

Eumeces sumichrasti, Bocourt, Mission scientifique au Mexique . . ., Etudes sur 
les reptiles, livr. 6, 1879, p. 422.—Taytor, Univ. Kansas Sci. Bull., vol. 23, 
1935 (1936), pp. 178-186, figs. 21-23, pl. 12—Smirx and Necker, Anal. Esc. 
Nac. Cienc. Biol., vol. 3, 1948, pp. 192-194, pl. 3, fig. 4-—Smirx, Journ. 
Washington Acad. Sci., vol. 39, 1949, p. 41. 

Eumeces rovirosae Ducks, La Naturaleza, ser. 2, vol. 2, 1895-1896 (1895), pp. 
298-299, pl. 13 (type locality, Mineral de Santa Fé, Chiapas; type in Alfredo 
Dugés Museum, Colegio del Estado de Guanajuato). 

Eumeces schmidti Dunn, Proc. Acad. Nat. Sci. Philadelphia, vol. 84, Mar. 22, 
1932, pp. 30-31 (type locality, Lancetilla, Honduras; Acad. Nat. Sci. Phila- 
delphia No. 19877). 


Type.—U.S.N.M. No, 6601; Francois Sumichrast collector. 

Type locality.—“‘Orizava” ex errore, = Potrero, Veracruz. The 
specimen bears Sumichrast’s original tag, ‘‘Potrero, No. 4, F. Sumi- 
chrast.’”’? Later Sumichrast published notes that he had found the 
species “en los encinales de Potrero, cerca de Cordoba a una altura 
de 590 metros.” 

Range.—Veracruz to Chiapas in lowlands; Central America; Hon- 
duras, British Honduras, and Guatemala. Reported from Veracruz: 
Potrero, Jalapa; Chiapas: Palenque, La Esperanza, Mineral de Santa 
Fé. 


EUMECES BREVILINEATUS Cope 


Eumeces brevilineatus Corr, U. 8S. Nat. Mus. Bull. 17, 1880, pp. 18-19, 44, 46.— 
Taytor, Univ. Kansas Sci. Bull., vol. 28, 1935 (1936), pp. 283-290, figs. 
41, 42, 48, pl. 22. 

Type.—Lectotype, designated by Taylor, loc. cit., U.S.N.M. No. 
10159, one of four cotypes; G. W. Marnock collector. 

Type locality.—Helotes, Bexar County, Tex. 

Range.—Tamaulipas and Nuevo Leon; in the United States: 
southern Texas. Reported from Tamaulipas: 26 kilometers north of 
El Limén; Nuevo Leén: 4 miles west of Sabinas Hidalgo, 31 miles 
south of Sabinas Hidalgo. 


EUMECES CALLICEPHALUS Bocourt 


Eumeces callicephalus Bocourt, Mission scientifique au Mexique . . ., Etudes sur 
les reptiles, livr. 6, 1879, pp. 431-433, pl. 22D, figs. 2-2c, pl. 22E, fig. 2.— 
Taytor, Univ. Kansas Sci. Bull., vol. 23, 1985 (1936), pp. 290-298, figs. 44, 
45, pl. 23. 

Type.—Mus. Nat. Hist. Nat. Paris; Alfredo Dugés collector. 
Type locality.—Guanajuato [Guanajuato]. 
Range.—Sonora (Alamos), Chihuahua, Durango, Zacatecas, Jalisco, 

Guanajuato, Michoac4n, Querétaro (Huaxteca Potosina), Nayarit; in 

United States: Arizona (southeastern). 


CHECKLIST OF REPTILES OF MEXICO 165 


EUMECES TETRAGRAMMUS (Baird) 


Plestiodon tetragrammus Batrp, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 256. 

Eumeces tetragrammus, Corr, U. 8S. Nat. Mus. Bull. 1, 1875, p. 45—Taytor, 
Univ. Kansas Sci. Bull., vol. 23, pt. 1, 1985 (1936), pp. 298-304, fig. 46; 
tbid., vol. 29, 1943, p. 274. 

[Eumeces tetragrammus] var. funebrosus Cope, Ann. Rep. U. S. Nat. Mus., 1898. 
(1900), p. 661 (type locality, Matamoros, Tamaulipas; U. 8. N. M. No. 3120, 
two cotypes). 

Type—U. 5S. N. M. No. 3124; Darius Nash Couch collector. 

Type locality —‘‘Lower Rio Grande.” (Later indicated by Baird, 
as ‘“Matamoras, Mex.’”’ A second specimen was at hand collected by 
Dr. Kennerly at “Salado River,” U.S. N. M. No. 3139.) 

Range.—Tamaulipas, San Luis Potosi, and Veracruz, in lowlands; 
in the United States: southern Texas. Reported from Yamaulipas: 
3% miles west of El Forlén, San José, Matamoros; San Luis Potosi: 
Antiguo Morelos, Ebano; Veracruz: south of Tampico. 


EUMECES OBSOLETUS (Baird and Girard) 


Plestiodon obsoletum Barrp and GiraRbD, Proc. Acad. Nat. Sci. Philadelphia, vol. 
6, 1852, p. 129. 

Eumeces obsoletus, Corr, U. S. Nat. Mus. Bull. 1, 1875, p. 45.—Tayuor, Univ. 
Kansas Sci. Bull., vol. 23, pt. 1, 1935 (1936), pp. 305-320, figs. 47, 48, pl. 24. 

Lamprosaurus guttulatus HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, 1852, 
pp. 206-207 (type locality, “Jornada del Muerte,’’ New Mexico; Acad. Nat. 
Sci. Philadelphia). 

Type —U.S. N. M., No. 3133; John H. Clark collector. 

Type locality —‘‘Valley of the Rio San Pedro of the Rio Grande del 
Norte,” now Devils River, Tex. 

Range.—Tamaulipas, Nuevo Leén, Coahuila, and Chihuahua. In 
United States: Arizona, New Mexico, and Texas, north to Utah, 
Colorado, Nebraska, and Kansas. Reported from Tamaulipas: 
Matamoros; Nuevo Leén: Santa Catarina; Chihuahua: Chihuahua 
(city); Coahuila: Cuatro Ciénegas. 


EUMECES MULTIVIRGATUS (Hallowell) 


Plestiodon multivirgatum HAaLLOWELL, Proc. Acad. Nat. Sci. Philadelphia, 1857, 
p. 215. 

Eumeces multivirgatus, Cop, U. 8. Nat. Mus. Bull. 1, 1875, p.45.—Tayrtor, Univ. 
Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 341-358, figs. 54-56, pls. 27, 28. 


Type.—Acad. Nat. Sci. Philadelphia No. 9371; Dr. Hammond 
collector. 

Type locality —‘‘Posa Creek, 460 miles west of Fort Riley, Kan- 
sas’ (=Cow Creek,** Larimer County, Colo.). 

Range.—The central high plains and proplateau areas south from 
southwestern Nebraska to extreme western Texas, northern Arizona, 


8 The original tag attached to the specimen definitely states Cow Creek. The ‘‘Posa’”’ is a bad interpreta- 
tion of poor chirography. 


166 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


and presumably western Chihuahua. Recorded in Mexico only from 
Chihuahua: no specific locality.” 


EUMECES HUMILIS Boulenger 


Eumeces Bocourtii (non Brocchi) BouLENGER, Ann. Mag. Nat. Hist., ser. 5, vol. 11, 
1883, p. 342. 

Eumeces humilis BouLENGER, Catalogue of the lizards in the British Museum, 
vol. 8, 1887, p. 377 (new name for Eumeces Bocourtii)—TayYuor (part), 
Univ. Kansas Sci. Bull., vol. 23, 1985 (1936), pp. 358-363, figs. 58, 59.— 
SmituH, Proc. New England Zool. Club, vol. 21, 1942, pp. 94, 95. 

Type.—Brit. Mus. Nat. Hist. Nos. 83,4, 5,33-34 (two cotypes); 

Alfonso Forrer collector. 

Type locality —Presidio [Sinaloa]. There is a strong presumption 
that the locality is in Sinaloa although not known positively to be so. 

Range.—Sinaloa. Known only from the type locality. (Specimens 
previously referred to this species from Texas and New Mexico have 
been referred to another form, Humeces taylori, by Smith.) 


EUMECES PARVULUS Taylor 


Eumeces parvulus Taytor, Proc. Biol. Soc. Washington, vol. 46, 1933, pp. 175- 
178, fig. 1; Univ. Kansas Sci. Bull., vol. 28, 1985 (1936), pp. 363-367, figs. 
59, 60, pl. 31, figs. 3, 4.—Smitru, Copeia, 1948, p. 250. 
Type.—U.S.N.M. No. 56903; collector unknown. Originally in the 
Julius Hurter collection. 
Type locality.—Tepic, Nayarit. 
Range.—Colima, Nayarit and Sinaloa. Reported from Colima: 
Paso del Rio; Nayarit: Tepic, Compostela, Minim4n; Sinaloa: Plomo- 


sas. 
EUMECES PARVIAURICULATUS Taylor 


Eumeces parviauriculatus Taytor, Proc. Biol. Soc. Washington, vol. 46, 1933, pp* 
178-181, fig. 2; Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 368-371, 
fig. 59, pl. 31, fig. 5. 
Type.—U.S.N.M. No. 47536; E. A. Goldman collector. 
Type locality.—Near Alamos, Sonora. 
Range.—Sonora and western Chihuahua. Reported from Sonora: 
Near Alamos; Chihuahua: Mojarachic. 
EUMECES COPEI Taylor 
Eumeces copet Tayutor, Proc. Biol. Soc. Washington, vol. 46, 1933, pp. 133-137; 
Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 387-394, figs. 64, 65, pl. 33. 
Type.—EHT-HMS No. 29717 (EHT field No. 3859); H. M. Smith 
and EK. H. Taylor collectors. 
Type locality.—Ten miles southeast of Asuncién, in the western part 
of the state of México. 
8 This specimen, U. 8. N. M. No. 30833, very likely represents a distinct species (see Taylor, loc. cit., 
p. 353), but we hesitate to attempt a characterization of it because of certain obvious abnormalities the speci- 


men possesses, and because of its intimate and entirely uncertain relationship with populations farther to 
the north in Arizona and New Mexico. 


CHECKLIST OF REPTILES OF MEXICO 167 


Range.—Puebla, Distrito Federal, Morelos, México, and Michoacan. 
Reported from Puebla: Near Rio Frio (México); Mézico: Rio Frio, 
near Salazar, 10 miles southeast of Asuncién, 8 miles west of Villa 
Victoria, 15 kilometers west of Toluca, 3 miles west of Zinacatepec; 
Morelos: Tres Marias, Lagunas de Zempoala; Distrito Federal: Santa 
Lucia; Michoacdén: Cerro de Tecolote, Mount Tancitaro. 


EUMECES SKILTONIANUS (Baird and Girard) 


Plestiodon skiltonianum Batrp and Grrarp, Proc. Acad. Nat. Sci. Philadelphia, 
1852, p. 69. 

Eumeces skiltonianus, Corr, U. S. Nat. Mus. Bull. 1, 1875, p. 45.—Roperrs and 
Fitcu, Univ. California Publ. Zool., vol. 48, 1947, pp. 193-198, pl. 8a (col.). 

Eumeces skiltonianus skiltonianus, Taytor, Univ. Kansas Sci. Bull., vol. 23, pt. 
1, 1935 (1936), pp. 415-428, fig. 68, pls. 35, 36, figs. 2, 3, 4. 

Eumeces skiltonianus amblygrammus Cores, Ann. Rep. U.S. Nat. Mus., 1898 (1900), 
p. 643 (type locality, Fort Humboldt, Calif.; U.S.N.M. No. 3166). 


Type.—U.S.N.M. No. 3172, two cotypes, smaller one designated as 
lectotype; Rev. George Gary (or Geary) collector. 

Type locality.—Oregon, here restricted to The Dalles. 

Range.—Southern British Columbia to northwestern Baja California 
and adjacent islands, and eastward to western Montana and Utah. 
Reported in Baja California from Los Coronados Islands, Todos Santos 
Islands; Ensenada, San José, Rancho San José, Alcatraz, San Pedro 
Martir Mountains, Arroyo Encantada, San Quintin, etc. 


EUMECES GILBERTI RUBRICAUDATUS Taylor 


Eumeces quadrilineatus HALLOWELL (nec Blyth), Reports of explorations and 
surveys, to ascertain the most practicable and economical route for a railroad 
from the Mississippi River to the Pacific Ocean, 1853-6, vol. 10, pt. 4, Zool., 
Rept., 1859, p. 10, pl. 9, figs. 3a, b, ¢, d (type locality, ‘“Upper California, 
near Mohave river and in San Bernardino Valley,”’ here restricted to Mojave 
River; U. 8. Nat. Mus.). 

Eumeces gilberti rubricaudatus Taytor, Univ. Kansas Sci. Bull., vol. 23, 1935 
(1936), pp. 446-451, figs. 72-73, pl. 39, vol. 29, 1943, p. 277; Rodgers and 
Fitch, Univ. California Publ. Zool., vol. 48, 1947, pp. 203-205, pl. 8e (col.). 


Type.—California Acad. Sci. No. 39002. 
Type locality— Tehachapi Mountains, Calif. 
Range.—Southern California and extreme northwestern Baja 


California. Reported in Mexico only from Baja California: San 
Antonio del Mar, North Coronado Island. 


EUMECES LAGUNENSIS Van Denburgh 


Eumeces lagunensis VAN DEeNBURGH, Proc. California Acad. Sci., ser. 2, vol. 5, 
1895, pp. 79, 1384-135, pl. 18.—Taytor, Univ. Kansas Sci. Bull., vol. 23, 
1935 (1936), pp. 481-437, figs. 69, 70, pl. 36, fig. 1—RopcrErs and Fircn, 
Univ. California Publ. Zool., vol. 48, 1947, pp. 205-206. 


168 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Plestiodon lagunensis, VAN DENBURGH and SLEVIN, Proc. California Acad. Sci., 
ser. 4, vol. 11, 1921, pp. 28, 40, 44, 52. 

Plestiodon skiltonianus lagunensis, NEutson, Mem. Nat. Acad. Sci., vol. 16, 1921, 
pp. 114-115. 

Eumeces skiltonianus lagunensis, LinspAuLE, Univ. California Publ. Zool., vol. 38, 
19382, p. 374. 

Eumeces skiltonianus, LovERIDGE, Copeia, No. 173, 1930, pp. 111-112. 


Type.—Originally California Acad. Sci. Nos. 400 and 402, cotypes 
(destroyed in the earthquake and fire in 1906); Gustav Eisen collector. 
Neotype.—U. S. N. M. No. 67398; W. M. Mann collector; February 
1924, ‘‘on the Trail between Loreto and Comondt.” 

Type locality—San Francisquito, Sierra de la Laguna, Baja 
California. 

Range.—Confined to the southern third of Baja California in the 
mountains. Reported from Baja California: Comondt; San Francis- 
quito, Sierra de la Laguna; between Loreto and Comondu. 


EUMECES BREVIROSTRIS (Giinther) 


Mabouia brevirostris GinrueEr, Proc. Zool. Soc. London, 1860, pp. 316-317. 
Eumeces brevirostris, Bocourt, Mission scientifique au Mexique . . ., Etudes sur 
les reptiles, livr. 6, 1879, pp. 489-440, pl. 22A, figs. 7, 7a, 7b, and pl. 22E, 
fig. la.——Taytor, Univ. Kansas Sci. Bull., vol. 23, pt. 1, 1935 (1936), pp. 
459-466, figs. 76, 77, pl. 41. 
Type.—Brit. Mus. Nat. Hist.; Auguste Sallé collector. 
Type locality Oaxaca, here restricted to the city of Oaxaca. 
Range.—Durango to Oaxaca and east to Veracruz, in highlands. 
Reported from Guerrero: Omilteme; Cazaca: Cerro San Felipe, La 
Parada, San José Lachiguiri, Tehuantepec; Veracruz: Laguna, Perote, 
San Bernardino, Totalco, Orizaba; Puebla: Near Rio Frio (México), 
Texmelucan, El Seco (Km. 205); Durango: Ciudad, El Salto, Coyote; 
Jalisco: La Cumbre de los Arrastrados, Talpa, Mascota, Sierra de 
Juanocatlan, La Laguna; Michoacdn: No specific record. 


KUMECES INDUBITUS Taylor 


Eumeces indubitus Taytor, Univ. Kansas Sci. Bull., vol. 21, 1933 (1934), pp 
257-267, pls. 24, 25; vol. 23, pt. 1 (1936), pp. 466-472, figs. 76, 78, pl. 42. 


Type.-—EKHT-HMS No. 29715 (EHT field No. 1731); E. H. Taylor 
and H. M. Smith collectors. 

Type locality.—Kilometer 63 (Mexico-Cuernavaca highway), near 
Cuernavaca, Morelos. 

Range.—Known from the states of Morelos, México, and Michoacan. 
Reported from Morelos: near Cuernavaca, Tepoztlan, Lagunas de 
Zempoala, Huajintlan; México: Asuncién; Michoacdn: 15 miles south- 
east of Zitacuaro, Puerto Hondo. 


CHECKLIST OF REPTILES OF MEXICO 169 


EUMECES DUGESII Thominot 


Eumeces (Plestiodon) Dugesii THominot, Bull. Soc. Philom. Paris, ser. 7, vol. 7 
(1882-1883), 1883, pp. 138-139.—Taytor, Univ. Kansas Sci. Bull., vol. 23, 
pt. 1, 1935 (1936), pp. 472-478, figs. 76, 79, pl. 43. 

Type.—Mus. Hist. Nat. Paris; Alfredo Dugés collector. 

Type locality —‘ Province Guanajuato,” here restricted to the city 
of Guanajuato. 

Range.—Guanajuato, Jalisco (?), and Michoacan. Reported from 
Guanajuato: Guanajuato; Michoacén: Carapa, Rancho San José, 9 
miles west of Zacapt, Tangancicuaro, Patamban, El Soledad (Tan- 
citaro), Apatzing4an; Jalisco: Nevado de Colima.” 


EUMECES COLIMENSIS Taylor 
Eumeces colimensis Taytor, Publ. Field Mus. Nat. Hist., zool. ser., vol. 20, 1935, 
pp. 77-80, fig. 7; Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 478-482, 
figs. 76-80, pl. 40, fig. 3. 
Type.—Chicago Nat. Hist. Mus. No. 1649; collector unknown. 
Type locality —Colima, Colima. 
Range.—Known only from the type locality. 


EUMECES DICEI Ruthven and Gaige 


Eumeces dicei RuTHVEN and GaicE, Occ. Pap. Mus. Zool. Univ. Michigan, No. 
260, 1933, pp. 1-3.—Taytor, Univ. Kansas Sci. Bull., vol. 238, 1985 (1936), 
pp. 482-485, figs. 76-81; vol. 29, 1943, pp. 280-282. 
Type-—Mus. Zool. Univ. Michigan No. 69253; Lee R. Dice 
collector. 
Type locality —Marmolejo, Tamaulipas. 
Range——Tamaulipas and Nuevo Leén. Reported from Nuevo 
Leén: Pablillo, Cieneguillas south of Galeana; Tamaulipas: Mar- 


molejo. 
EUMECES OCHOTERENAI Taylor 


Eumeces ochoterenae Taytor, Proc. Biol. Soc. Washington, vol. 46, 1933, pp. 
129-133, 2 figs.; Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 485-489, 
figs. 76, 82, pl. 43. 

Type-—EHT-HMS No. 29716 (EHT Field No. 1015); E. H. 
Taylor and H. M. Smith collectors. 

Type locality —Mazatlan; ‘4 mi. north of Chilpancingo, Guerrero”’ 
(actually about 15 km. south of Chilpancingo). 

Range—Known only from Guerrero; probably confined to the 
Sierra Madre del Sur. Reported from Agua del Obispo, 7 miles east 
of Chilpancingo, Mazatlan, vicinity of Chilpancingo, between Rincén 
and Cajones, Chilapa. 

% Gadow (Proc. Zool. Soc. London, 1905, pp. 195, 218-219) records a “‘fuscirostris”’ (=furcirostris) from 
this locality. Inasmuch as the most distinctive feature of d. furcirostris is the possession of three supra- 


oculars, it seems highly probable that Gadow observed this character in his specimen from the Nevado. 
In such case the specimen would be dugesii, the only species in that area that possesses this character. 


170 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Family ANELYTROPSIDAE Cope 


Anelytropsidae Corn, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 228; Ann. 
Rep. U. 8. Nat. Mus. for 1898 (1900), p. 665. 


Genera.—A single genus, Anelytropsis Cope, is known. 
Range.—F oothills of the plateau in central eastern Mexico. 


Genus ANELYTROPSIS Cope 


Aunelytropsis Corr, Proc. Amer. Philos. Soc., vol. 22, 1885, p. 380; Ann. Rep. 
U.S. Nat. Mus. for 1898 (1900), p. 667. 


Genotype.—Anelytropsis papillosus Cope. 
Range.—Central eastern part of Mexico. 


Species.—One. 
ANELYTROPSIS PAPILLOSUS Cope 


Anelytropsis papillosus Corr, Proc. Amer. Philos. Soc., vol. 22, 1885, pp. 380-381, 
pl. (unnumbered), fig. 9 (opposite p. 184).—Smiru, Univ. Kansas Sci. Bull., 
vol. 22, 1935, p. 146, pl. 24, figs. 2, 3. 


Type.—Unknown; originally two cotype specimens. 

Type locality.—Near Jalapa, Veracruz. 

Range.—Eastern San Luis Potosi and eastern central Veracruz. 
Reported from San Luis Potosi: 20 miles south of Valles; Veracruz: 
Jalapa, Motzorongo. 


Family TEIIDAE Gray 
Teiidae Gray, Philos. Mag., ser. 2, vol. 2, 1827, p. 55. 


Genera.—About 40 genera, 3 of which, Cnemidophorus, Ameiva, 
and Gymnophthalmus, occur in Mexico. 


KEY TO MEXICAN GENERA OF TEIIDAE 


1. Anterior nasal plates not separated by a frontonasal; frontoparietal present; 
inner finger well developed; scales granular; eyelids present________---- 2 
Nasal plates widely separated by a frontonasal; inner finger absent; no eyelids; 
scales cycloid, quincuncial; prefrontals present but no frontoparietals. 

Gymnophthalmus (p. 192) 
2. Central gular scales not or scarcely enlarged as compared with adjacent lateral 
scales; tongue somewhat widened posteriorly, and notched behind; no basal 
sheath between base of tongue and larynx_____-- Cnemidophorus (p. 174) 
Central gular scales enlarged, often abruptly so; tongue not, or less widened 

posteriorly, a basal sheath evident between the base of tongue and larynx. 
Ameiva (p. 170) 

Genus AMEIVA Meyer 


Ameiva Mryer, Synopsis reptilium ... sistema generum methodum . 
1795, p. 27. 


* ey 


Genotype.—Lacerta americana Seba [=Ameiva ameiva (Linnaeus)]. 

Range.—Tamaulipas and Jalisco to Brazil; West Indies. 

Species.—About 13, with some 28 forms. Two species with 11 
subspecies occur in Mexico. 


10. 


CHECKLIST OF REPTILES OF MEXICO 171 


KEY TO MEXICAN SPECIES OF AMEIVA 


. Outer row of ventrals considerably smaller than the others. 


festiva edwardsii (p. 174) 


Outer row of ventrals as large as the others___.__._____..-___________ 2 
Ipreanaluscalestiny2 LOWS = = =a = om ae ee ees Se ee 3 
Preanal scales in one row or with no more than 1 posterior scale divided__ 9 


. Two rows of granules between third supraocular and superciliaries; third 


supraoculars generally completely separated from frontoparietals by 
granulése ee 355 = = ot (ek pepe obese 5. taney ie of undulata parva (p. 173) 
A single row of granules between third supraoculars and superciliaries; third 
supraoculars in contact with frontoparietals anteriorly____..__...___-- 4 


peViedian rulars abruptly enlarcedsss = =e aes meen een en yee eee ees 6 


Median gulars little enlarged, gradually merging with lateral gulars______ 5 


. Upper lateral vertical light lines from axilla to groin, 12 or more light lines. 


undulata gaigeae (p. 172) 

Upper lateral vertical lines from axilla to groin, 11 or less light lines. 
undulata hartwegi (p. 171) 
Median gulars irregular or no more than 2 regular (87 percent); lamellae 
under fourth toe 28 or more; no upper lateral light stripe, although large 
spots take its place in adult males____________ undulata podarga (p. 172) 
At least 3 median gulars regular (100 percent in all except u. amphigramma, 
with 87 percent); lamellae variable; upper lateral light stripe present or 
absento=. = = Ste wie Se Be ese CANE SERRE oo oe ao 


. Dorsolateral dark stripes present except in some large adults, in which the 


upper lateral light spots if present are much narrower than the spaces between 
them; lamellae on the fourth toe usually (88 percent) 27 or less. 
undulata stuarti (p. 173) 
No dorsolateral dark stripes; upper lateral light spots as wide as or wider 
than spaces between, or represented by a continuous upper lateral light 
stripe lamellae variables? 2.2 oe ae eee 8 
Upper lateral light spots or lines not contacting dorsolateral light area or 
line in adult males, separated by a narrow dark area; lateral gulars not or 


BCArceliyfeM lang ces ae eee ee eee undulata amphigramma (p. 172) 
Upper lateral light spots merged with dorsolateral light line in adult males; 
lateral gulars markedly enlarged_____________ undulata thomasi (p. 173) 


Last preanal scale generally (86 percent) divided; lateral markings showing 
little tendency to be arranged vertically; middorsal markings greatly re- 


Guced=. Sia Re eee, BEG Ae £9 EGY undulata dextra (p. 173) 
Last preanal scale generally entire; lateral markings tending to be arranged 
vertically; middorsal markings well developed -_-_-_------------------ 10 


Upper lateral light spots in adult males large, wider than intervening dark 
spaces; generally (95 percent) 5 or less rows of preanals. 

undulata sinistra (p. 174) 

Upper lateral light spots in adult males small, narrower than intervening 

dark spaces in all males; frequently (65 percent) 6 or more rows of preanals 

undulata undulata (p. 174) 


AMEIVA UNDULATA HARTWEGI Smith 


Ameiva undulata hartwegi Smiru, Proc. Biol. Soc. Washington, vol. 53, 1946, 


p. 55.—Smirxu and Laursg, Univ. Kansas Sci. Bull., vol. 31, 1946, pp. 35-30, 
pl. 2, fig. b. 
861316—50——12 


172 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type.—U.S.N.M. No. 108600; H. M. Smith collector. 

Type locality —‘‘Across the Rio Usumacinta from Piedras Negras, 
Guatemala, in Chiapas, Mexico.” 

Range.—Atlantic slopes of Mexico and Guatemala from the vicinity 
of the southeastern end of Laguna de Términos south and eastward 
across the base of the Yucatan Peninsula to northwestern Honduras. 
Recorded in Mexico only from Chiapas (the type locality) ; it probably 
occurs also in Campeche and Quintana Roo. 

AMEIVA UNDULATA GAIGEAE Smith and Laufe 
Ameiva undulata gaigeae SmMirH and Lavurs, Univ. Kansas Sci. Bull., vol. 31, 
1946, pp. 37-39, fig. 1C, pl. 2C. 

Type—EHT-HMS No. 11927; Progreso, Yucatan; Hobart M. 
Smith collector. 

Type locality.—Progreso, Yucatan. 

Range.—Northern half of the Yucatan Peninsula and southward 
to the island Carmen along the extreme eastern coast. Reported 
from Yucatdén: Chichen Itza, La Vega, Tunkas; Quintana Roo: 
Coba, Mujeres Island; Campeche: Champoton. 

AMEIVA UNDULATA PODARGA Smith and Laufe 
Ameiva undulata podarga SmitH and Laurer, Univ. Kansas Sci. Bull., vol. 31, 
1946, figs. 1D, 2A, pp. 40-43. 

Type.-EHT-HMS No. 14471; Hobart M. Smith and David H. 
Dunkle collectors. 

Type locality—Seven miles west of Ciudad Victoria, Tamaulipas. 

Range.—Known from southern Tamaulipas and eastern San Luis 
Potosi and probably extends into Veracruz and Hidalgo lowlands. 
Reported from Tamaulipas: Alta Mira, Victoria, Hacienda La 
Clementina near Forlén, Antiguo Morelos; San Luis Potosi: near 
Ciudad del Maiz, Rio Guayala near Magiscatzin, near Valles, Huichi- 
huayan, Tamazunchale. 


AMEIVA UNDULATA AMPHIGRAMMA Smith and Laufe 


Ameiva undulata amphigramma Smita and Laure, Trans. Kansas Acad. Sci., 
vol. 48, 1945, pp. 338-344; Univ. Kansas Sci. Bull., vol. 31, 1946, pp. 43-47, 
pled, figs: ©; (D: 

Type.—EKHT-HMS No. 11983; H. M. Smith collector. 

Type locality —San Andrés Tuxtla, Veracruz. 

Range.—Northern Veracruz southward at low elevations to the 
Isthmus of Tehuantepec, westward into valleys extending into extreme 
eastern Oaxaca and probably northeastern Puebla. Reported from 
Veracruz: Atoyac, Boca del Rio, Cuatotolapam, Lake Catemaco, 
Potrero Viejo, Rodriguez Clara, San Andrés Tuxtla, etc.; Oaxaca: 
Cosolapa, Matias, Agua Fria, Tuxtepec; Tabasco: La Venta; Puebla: 
1 mile northeast of Huauchinango. 


CHECKLIST OF REPTILES OF MEXICO 173 


AMEIVA UNDULATA THOMASI Smith and Laufe 


Ameiva undulata thomast SmirH and Laure, Univ. Kansas Sci. Bull., vol. 31, 
1946, pp. 47-50. 


Type.-—EHT-HMS No. 15327, H. D. Thomas, collector. 

Type locality—lLa Libertad, Chiapas, near Rio Cuilco where it 
crosses the Guatemalan border. 

Range.—Upper tributaries of Rio Grijalva in the interior of Chiapas 
and adjacent Guatemala. Recorded in Mexico only from the type 
locality. 

AMEIVA UNDULATA STUARTI Smith 


Ameiva undulata stuartt Smiru, Proc. Biol. Soc. Washington, vol. 53, 1940, pp. 
55-56.—SmirH and Laure, Univ. Kansas Sci. Bull., vol. 31, 1946, pp. 50-51, 
fig. 1B; pl. 1, fig. B. 

Type.—U.S.N.M. No. 108601, H. M. Smith collector. 

Type locality Palenque, Chiapas. 

Range.—Atlantic slopes of Mexico from the middle of the Isthmus 
of Tehuantepec eastward in the lowlands to the southern borders of 
Laguna de Términos and to Tenosique, Tabasco; southward up the 
valley of the Rio Grijalva at least as far as Tuxtla Gutiérrez, Chiapas. 
Reported from Chiapas: Palenque, San Ricardo, Tuxtla Gutiérrez; 
Campeche: Balchacaj, Tres Brazos; Tabasco: Tenosique, Frontera, 
Teapa; Oaxaca: Isthmus of Tehuantepec (intergrades?). 


AMEIVA UNDULATA PARVA Barbour and Noble 


Amewa undulata parva Barsour and Nosiez, Bull. Mus. Comp. Zool., vol. 59, 
1915, pp. 476-477. —Smiru and Laursg, Univ. Kansas Sci. Bull., vol. 31, 1946, 
pp. 51-54, fig. 1A, pl. 2, fig. A. 
Type.-—Mus. Comp. Zool. No. 5831; Van Patten collector. 
Type locality Guatemala (restricted by Smith and Laufe to 
Mazatenango). 
Range.—Pacific slopes from the Isthmus of Tehuantepec in Oaxaca 
to Costa Rica. Reported in Mexico from Oazaca: near Niltepec; 
Chiapas: Tapachula, Tonalé, La Esperanza, Huehuetan. 


AMEIVA UNDULATA DEXTRA Smith and Laufe 


Ameiva undulata dextra Smiru and Laure, Univ. Kansas Sci. Bull., vol. 31, 1946, 
pp. 54-59, fig. 2C. 

Type.-—EHT-HMS No. 11966; E. H. Taylor and H. M. Smith 
collectors. 

Type locality—Near Rincén, Guerrero. 

Range.—Southern slope of the Sierra Madre del Sur. Known from 
Oaxaca and Guerrero. Reported from Oazaca: Juquila; Guerrero: 
Chilpancingo, Cocoyul, Los Cajones, Rincén, Organos, El Treinta, 
Acapulco, 8 miles east of Coyuca. 


174 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


AMEIVA UNDULATA SINISTRA Smith and Laufe 


Ameiva undulata sinistra Smiru and Lavurs, Univ. Kansas Sci. Bull., vol. 31, 1946, 
pp. 59-62. 

Type —EHT-HMS No. 11908; H. M. Smith collector. 

Type locality —Manzanillo, Colima. 

Range.—Pacific slopes from Jalisco to the arid Balsas basin in 
Michoacan, thence inland along the northern drainage of the Rio 
Balsas to Puebla. Reported from Colima: Colima, Queseria, Hda. 
Paso del Rio, Salvador, Pascuales, Periquillo, Manzanillo; Jalisco: 
Tenacatita, Ixtapa, Tenacatita Bay; Michoacén: Uruapan; Puebla: 
Chiautla; Morelos: Puente de Ixtla. 





AMEIVA UNDULATA UNDULATA (Wiegmann) 


Cnemidophorus undulatus WiEGMANN, Herpetologia Mexicana, 1834, p. 27. 

Ameiva undulata, Gray, Catalogue of the specimens of lizards in the British 
Museum, 1845, p. 20. 

Ameiva undulata undulata, Smira and Laure, Univ. Kansas Sci. Bull., vol. 31, 
1946, pp. 62-64, fig. 2B. 


Type.—Zool. Mus. Berlin; F. Deppe collector. 

Type locality Mexico, by inference. Restricted by Smith (Proc. 
Biol. Soc. Washington, vol. 53, 1940, p. 56) to Tehuantepec. 

Range.—Pacific slopes of the Isthmus of Tehuantepec as far west 
as Puerto Angel, and eastward to Niltepec. Known only in the state 
of Oaxaca: El] Limén, Palmar, Tres Cruces, Cafetal Concordia, Juchi- 
tan, Puerto Angel, Totontepec, Tehuantepec, Rancheria La Manga. 


AMEIVA FESTIVA EDWARDSII Becourt 


Ameiva edwardsii Bocourt, Ann. Sci. Nat., ser. 5, vol. 17, 1873, art. 17, pp. 1-2. 

Ameiva festiva edwardsii, Sruart, Occ. Pap. Mus. Zool. Univ. Michigan, No. 471, 
1943, p. 21, fig. 7—Smiru and Laurs, Univ. Kansas Sci. Bull., vol. 31, 1946, 
pp. 64-66. 


Type.—Mus. Hist. Nat. Paris; Commission Scientifique collector. 

Type locality—tIzabal and Santa Maria de Panzés, near Rio 
Polochic, Guatemala; here restricted to Panzés. 

Range.—Atlantic foothills from the Isthmus of Tehuantepec 
through Honduras, in heavy forests. Recorded in Mexico only from 
Chiapas: Ixtacomitan, Palenque, across the Rio Usumacinta from 
Piedras Negras (Petén, Guatemala). 


Genus CNEMIDOPHORUS Wagler 


Cnemidophorus WaGiER, Natiirliches System der Amphibien . . ., 1830, p. 154.— 
Burt, U. 8. Nat. Mus. Bull. 154, 1931.—Buremr, Chicago Acad. Sci. Nat. 
Hist. Misc., No. 65, 1950, pp. 1-9. 

Aspidoscelis FITzINGER, Systema reptilium, 18438, p. 20 (type, Lacerta serlineata 
Linnaeus). 

Verticaria Corr, Proc. Amer. Philos. Soc., vol. 11, 1869, p. 158 (type, Cnemi- 
dophorus hyperythrus Cope). 


CHECKLIST OF REPTILES OF MEXICO 175 


Genotype.—Seps murinus Laurenti. 

Range.—North America south from Oregon, Wisconsin, and Mary- 
land through Central America to southern Brazil and Bolivia. 

Species.—About 18, and a total of about 47 species and subspecies; 
15 species and 41 forms, as here recognized, occur in Mexico. These 
18 species represent 5 distinct groups, all but one of which (the 
lemniscatus group, restricted to Central and South America, of 3 
species and 5 forms as recognized by Burt, op. cit.), occur in Mexico. 


KEY TO MEXICAN SPECIES OF CNEMIDOPHORUS 


ia nly; One frontoparietal scalee ese. . a. ee ec et ee 2 
ERIS PE TASTE IDEN TLC ORS 2 eae sha a te 3h ten ke WR ee 9 
2. Dorsal surface unicolor, not striped or reticulated_hyperythrus pictus (p. 186) 
Dorsal surface striped or reticulated, not unicolor_.__.__._...__....__._____-- 3 
3. Lateral stripes more or less broken; at least a few spots or cross bars present; 
body often tessellated; adults large.__._._...__._______- ceralbensis (p. 188) 
Lateral stripes distinct and in even, continuous lines; spots and cross bars 
St Seld SEO PAM OTH eo) Peel oe a kee: S Hep gee a FOE ae Ye 4 
4. Usually 2 or 3 longitudinal light lines in vertebral area____.___.________-- 5 
Usually 1 median dorsal light line, forked anteriorly or not_________---- 6 
5. Usually 3 light lines along middie of back; second supraocular usually without 
granules between it and frontal_______ hyperythrus hyperythrus (p. 187) 
Usually 2 light lines; second supraocular usually at least in part separated 
fromifrontal by granules. _. ... =... - hyperythrus beldingi (p. 186) 
6. Dorsal stripe not of same intensity and width as lateral stripes but fainter and 
ABSA E pA OE? ec tee ane os A ee bets ee gs ee 7 
Dorsal stripe of same intensity and width as lateral stripes____.___-__--- 8 
7. More than half of second supraocular usually in contact with frontal. 

hyperythrus schmidti (p. 187) 

Half or less than half of second supraocular usually in contact with frontal. 
hyperythrus beldingi (p. 186) 
8. Ground color of sides (between the 2 distinct lateral stripes) reddish gray; 
dorsal ground color gray or black, abruptly contrasted with lateral ground 
COON a ee oe hyperythrus danheimae (p. 186) 
Ground color of sides not usually reddish gray, but often reddish or brownish; 
dorsal ground color of same general shade as that of the sides and not in 


SHATOICONETASe LOUt =A) =e hyperythrus caeruleus (p. 186) 

9. Supraoculars normally 3; striped at least in the young______---------- 10 
Supraoculars normally 4, or, if 3, no stripes at any stage, but a pattern of small, 
scattered light spots on a dark background____.___._........._---= 15 

10. Scales between midventral scutes and anus 4 to 8; snout-vent measurement 
REV ER CXECCOIN GO ODIs =n oe = ee eee ed 12 
Seales between midventral scutes and anus 9 to 15; snout-vent measurement 
TEACHING pl Sern Tne ees ee ay ee A fee gen tae Sk Te et eee 11 


11. Adults andsubadults spotted, the spots often dim_guttatus guttatus (p. 179) 
Adults and subadults (as well as young) striped, the stripes sometimes broken 
LIP COIS Os eee, Se se guttatus immutabilis (p. 180) 


1 The species possessing this character were formerly regarded as members of the genus Verticaria Cope. 
Burt (U. S. Nat. Mus. Bull. 154, 1931, p. 14) has concluded, however, that the group is not a natural one, 
since one species (ceralbensis) is related to the tesselatus group, while the others are related to the serlineatus 
group. The character is, furthermore, subject to a slight variation. 


176 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


12. 


13. 


14. 


15. 


16. 


ie 


18. 


19. 


20. 


21. 


22. 


23. 


24. 


An accessory scute between frontoparietal and parietal. 
deppii cozumelus (p. 179) 


INO SUCCESSOR Yas S UG C2 saan ae spe teu ee gs ta eg ce ee ley 13 
Femoral pores usually 17 or fewer (81-82 percent)____________________ 14 
Femoral pores usually 18 or more (94 percent) _______- deppii deppii (p. 178) 
Lower preocular generally (79 percent) in contact with loreal; a broad mid- 

dorsal light bande sas< 2) oe Ses ee ee deppii lineatissimus (p. 179) 
Lower preocular generall (73 percent) not in contact with loreal; narrow light 

stripesion back as\on sides 2. 2-222 sa 4e deppii oligoporus (p. 179) 


Enlarged scales immediately preceding gular fold (mesoptychials) small (not 
more than 3 times diameter of smallest scales in median part of pregular 
fold); or, if larger, no evidence whatever of longitudinal light lines, or else 
at least a few dark spots on throat below level of ear openings_-____-__ 16 

Mesoptychials larger; a lined pattern always present in the young, generally 
in adults; no dark spots on throat below level of ear openings; or, if less than 
3 times diameter of smallest scales in median part of pregular fold, 7 light 
lines and intervening dark spaces absolutely straight, continuous and 


CLES COMMING GGG ee sate ae ee a a iN eRe ep 28 
Mesoptychials moderately or considerably enlarged, more than 3 times 
diameter of smallest scales in median part of pregular fold__________ 17 
IMesoptychialstemiall ens. crc se one sees a ere ne nae 19 
No evidence of a longitudinal arrangement of lines on back___________ 18 
Dorsal'surface distinctly lined=22-2=_ =) == ae eee tesselatus (p. 188) 


No spotting in area above level of tympanum and anterior to insertion of 
forearm, spots relatively well defined, very small, not yellowish. 
bacatus (p. 187) 
Spots present in area above tympanum and anterior to insertion of forearm; 
spots often obscure and diffuse, usually orange or yellowish. 
catalinensis (p. 188) 
Ventral surfaces of tail and limbs more or less suffused with red or pink. 20 
ING SOstr ae tee etc careneht earn Pind ea = maT ee el pee ea 21 
Black markings on temporal regions absent or few or ill-defined; dorsal and 
lateral markings on body showing both transverse and longitudinal arrange- 
ment in at least large specimens; each dark unit or spot square or rectangular 
in outline, not noticeably rounded_______________-_ tigris rubidus (p. 191) 
Black markings on temporal region well defined; dorsal and lateral markings 
on body showing a predominantly longitudinal arrangement; back covered 
by alternate longitudinal chains of black and white, individual spots in 
black chain rarely square or rectangular_________ tigris celeripes (p. 190) 
Dorsal markings consisting of either fine reticulations or light spots, and these 
markings without longitudinal arrangement_________________-_-_-- 22 
Dorsal markings with at least some indication of longitudinal arrange- 
NV CTR Uses cers eae ea ene ee ae ee cae me 2 ore el are 23 
Labials deep or moderate brown or black_________- tigris martyris (p. 191) 
Labials light gray, slate or light brown_________-_-__-_ tigris canus (p. 190) 
Dorsal light lines only 3 on each side (exclusive only of a pair of lateral lines), 
Uwice as wide as Interspaces=—- no tana eee ee ee maximus (p. 188) 
Dorsal light lines more numerous or narrower, or both_____._--------- 24 
Dorsal pattern of 6-8 persistent longitudinal light stripes.__..__.___-- 25 
Dorsal pattern of fewer than 6 unbroken light stripes; at least lower lateral 
stripe on each side broken by dark vertical bars__________-____---- 27 


CHECKLIST OF REPTILES OF MEXICO 177 


25. Dark fields between dorsal light stripes unspotted and unbroken throughout 


life; entire ventrum black in adults____________-_ tigris aethiops (p. 189) 
At least some of dark fields broken into rows of spots; ventrum not entirely 
black imeadultsh- 6 tees 15. ele teh 2 Fe ler et Sa IE te 26 


26. Middorsal dark field or row of spots distinctly wider than paravertebral light 
stripes; throat white, spotted or barred with black. 
tigris multiscutatus (p. 191) 
Middorsal dark field or row of spots as wide as or slightly narrower than 
paravertebral light stripes; throat and chest black; belly spotted. 
tigris gracilis (p. 188) 
27. Four longitudinal light stripes persisting unbroken in middorsum of adults, 
the dark fields between represented by rows of black spots; sides with 
bold-verticaltbars==". 5. ae es = care es eee tigris tigris (p. 189) 
Six longitudinal light stripes persisting unbroken in adults, or, if any broken, 
dorsal pattern generally light and without any bold markings. 
tigris marmoratus (p. 190) 


28. Anterior nasal in contact with second upper labial; femoral pores never more 


eRe TN AEE cs sre br Mo 7S ees SEN he al Fp pa gee Mires AS Gem gs 42 
Anterior nasal usually separated from second upper labial; if in contact, 
femoral pores; usually; morewthany W4es. eee ee ee ee 29 

29. Enlarged scales preceding gular fold relatively small, frequently grading into 
cranularscalesioffold: 28225 tees oe Se ee pa ee ee ee 30 
Enlarged scales preceding gular fold relatively large, abruptly differentiated 
FLOMUSTANULAT SCALES OLmeb INTO Beate mee ee ye ee er ee 31 


30. Postantebrachials enlarged; ventral surfaces light bluish in adults; juvenile 
striped pattern retained throughout life; maximum snout-vent length 70 
mm.; dorsal scales relatively large, 50 to 70 in a transverse row between 
enlarcedibellyaplatess:.- 5 =. See te CP pe inornatus (p. 184) 

Postantebrachials not enlarged; belly, chest, or gular region dark (black) in 
adults; snout-vent length reaching 95 mm.; dorsal scales 85 or more in a 
(EEATISVCESC REO Was = 2 es ee 8 tigris aethiops ” (p. 189) 


31. In addition to a dorsolateral light stripe on tail, a lateral light stripe present, 
bordered below bya black line (near base of tail) continuing onto otherwise 
unicolor posterior surface of thigh; no light spots in dark field between 
stripes in adults; postantebrachials not enlarged in either sex; neither sex 
darkablueoriorange belowWae === — see. ee eee eee sexlineatus (p. 184) 


No lateral light stripe on tail, at least not distinguishable from ventral tail 
color; thighs mottled or unicolor, no posterior light stripe except sometimes 
feebly indicated at insertion of leg; light spots often present in dark fields 
between stripes; postantebrachials distinctly enlarged generally, sometimes 
less so in females; males frequently dark blue on belly or chest, or else cream, 
REGED PAGE SUMMONS Ee ee 2 aa eo ee eh 8 Bw se 32 

32. Only 5 light lines, including a vertebral, 2 dorsolateral and 2 lateral lines 
through the ear; ventral surfaces cream even in males, never blue; light lines 
very broad, more than half width of intervening dark spaces in adults; no 
marking between light lines, except series of small spots in dark spaces in 
juveniles (disappearing in adults)_.__..........._..----.-- burti (p. 185) 

Wot Sosnid tien St eee is po at had ee te a EL 33 


Young only key out here; adults key out through couplet 16 to couplet 25. 


178 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


33. Space between paravertebral light stripes (i. e., those originating even with 
parietal scales, not with median interparietal) narrower or at least no wider 
than space between dorsolateral and upper lateral light stripes; if stripes not 


visible, they are replaced by dorsal pattern of dark cross bars__-_-_-_--_- 34 

Space between paravertebral light stripes wider; if stripes not visible, they are 

replaced by numerous scattered light spots____._.._.__...._...._.__-- 37 

34, Aqultsiwithoutya crosssbarred. pattern 4525422082222 22 ee ee 35 
Aduiltsierosss barred ils: 2s yo 00 lot Neel see De Neyo Ae Soe eye 3 


35. Dark interspaces or rows of dark spots continuing to area above hind legs. 
sackii stictogrammus (p. 183) 
Dark interspaces or rows of spots disappearing anterior to area above hind 
Leo Sey: ee. Mare un oa Neer RSL ea sackii semifasciatus (p. 184) 
36. Postantebrachials generally scutelike_____._________ sackii scalaris (p. 182) 
Postantebrachials generally polygones, little enlarged_sackii australis (p. 181) 
37. Hind legs mottled in adults, but not with small light spots; dorsal pattern 
ENOSS= 10 ATE! Clay CULT ys ees ee nc sackii sackii (p. 180) 
Hind legs with small light spots in adults; dorsal pattern not cross-barred_ 38 
38. Stripes absent in adult, replaced by numerous, scattered light dots especially 


towerdercarthalty ofebOdy sea ane eee sackii bocourti (p. 181) 
Sbripes presen Gym Ace tyes se ee ek Sakon nk Sy Re ene teats Eee eee 39 

39. Collar black in adults; stripes obsolete except for lateral stripes; keels on 
caudal scutes nearly parallel___.____..__.-_.._- sackii mariarum (p. 183) 
Collar light; stripes not obsolete; keels on caudal scales strongly oblique__ 40 

40. Ventral surfaces of tail and limbs pink_________ sackii angusticeps (p. 183) 
Ventral surfaces of tail and limbs cream or white____...____-.-__-___- 41 

41. Stripes becoming rows of light spots in adults__sackii communis (p. 182) 
Stripesnpersistentsg--s4 hewn) ae od sackii gularis (p. 183) 

42. Femoral pores 11-15 in known specimens__________-_____- labialis (p. 185) 
Femoral pores 19-20 in known specimens_______-__--_------ gadovi (p. 185) 


DEPPIT GROUP 


Species.—Six forms, belonging to two species, now recognized. 
Range.—Central Veracruz on the Atlantic coast and Nayarit on the 
Pacific southward to Costa Rica. 


CNEMIDOPHORUS DEPPII DEPPII Wiegmann 


Cnemidophorus deppii W1EGMANN, Herpetologia Mexicana, 1834, p. 29.—Bo- 
court, Mission scientifique au Mexique . . . , Etudes sur les reptiles, livr. 
3, 1874, p. 281, pl. 20c, fig. 5, pl. 20d, fig. 1. 

Cnemidophorus deppei [deppei], Gapow, Proc. Zool. Soc. London, 1906, pp. 809- 
316, figs. 71 D-G, 72 A-G. 

Cnemidophorus deppii deppiit, Burt, U.S. Nat. Mus., Bull. 154, 1931, pp. 56-63 
(part), fig. 14, right—Harrweea and Otiver, Occ. Pap. Mus. Zool. Univ. 
Michigan, No. 359, 1937, pp. 1-3.—Smirx, Ann. Carnegie Mus., vol. 30, 
1944, pp. 90-91. 

Cnemidophorus decemlineatus HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, 
1860, p. 482 (U.S.N.M. No. 6058, 12 cotypes; Nicaragua; C. Wright collector). 

Cnemidophorus lativittis Corn, Proc. Amer. Philos. Soc., vol. 17, 1877, p. 94 
(U.S.N.M. No. 30227; Juchitén, Oaxaca; F. Sumichrast collector). 

Cnemidophorus alfaronis Corr, Proc. Acad. Nat. Sci. Philadelphia, 1894, p. 199 
(Amer. Mus. Nat. Hist. No. 16315; San Mateo, Costa Rica; A. Alfaro 
collector). 


CHECKLIST OF REPTILES OF MEXICO 179 


Type.—Zool. Mus. Berlin; F. Deppe collector. 

Type locality —Mexico, restricted to Tehuantepec, Oaxaca. 

Range.—Southern Guerrero on Pacific slopes and the Salama Basin 
in Guatemala on Atlantic slopes, south to Costa Rica. Recorded in 
Mexico only from the states of Guerrero, Oaxaca, and Chiapas. 


CNEMIDOPHORUS DEPPII COZUMELUS Gadow 


Cnemidophorus deppei cozumela Gavow, Proc. Zool. Soc. London, 1906, p. 136. 
Cnemidophorus deppii cozumelus, Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 
63-65, fig. 14 left —Sruart, Occ. Pap. Mus. Zool. Univ. Michigan, No. 292, 
1934, pp. 12-13, fig. 1. 
Type.—Brit. Mus. Nat. Hist. Nos. 1886.4.15.17—20, four cotypes. 
Type locality —Cozumel Island, east coast of Quintana Roo. 
Range.—The eastern portion of the Yucatan Peninsula. Recorded 
in Mexico only from Quintana Roo (Mujeres and Cozumel Island). 


CNEMIDOPHORUS DEPPII LINEATISSIMUS Cope 


Cnemidophorus lineatissimus Corps, Proc. Amer. Philos. Soc., vol. 17, 1877, p. 94. 
Cnemidophorus deppii lineatissimus, HARTWEG and Ouiver, Occ. Pap. Mus. Zool. 
Univ. Michigan, No. 359, 1937, p. 2. 


Type.—U.S.N.M. Nos. 24937-40, 32299-32314, 20 cotypes; Hége 
collector. 

Type locality—Colima and Guadalajara (Jalisco), here restricted 
to Colima, Colima. 

Range.—Pacific slopes from Nayarit to central and perhaps southern 
Guerrero. The range interdigitates with that of C. d. depp in central 
Guerrero. Recorded only from Nayarit, Jalisco, Colima, Michoacan, 
Guerrero, and Puebla (Chiautla). 


CNEMIDOPHORUS DEPPII OLIGOPORUS Smith 


Cnemidophorus deppii oligoporus Smita, Publ. Field Mus. Nat. Hist., zool. ser., 
vol. 24, 1939, pp. 26-27; Smrra and Burcer, Trans. Kansas Acad. Sci., vol. 
53, 1950, pp. 173-174. 


Type.—Chicago Nat. Hist. Mus. No. 29145; Julius Friesser. 

Type locality —Perez, Veracruz. 

Range.—Atlantic slopes from northern Veracruz to Campeche. 
Recorded only from the states of Veracruz and Campeche (Ciudad del 
Carmen). 

CNEMIDOPHORUS GUTTATUS GUTTATUS Wiegmann 

Cnemidophorus guttatus WieaMANN, Herpetologia Mexicana, 1834, p. 29.— 
Bocourt, Mission scientifique au Mexique ..., Etudes sur les reptiles, 
livr. 3, 1874, p. 285, pl. 20c, fig. 4-—Burt, U.S. Nat. Mus. Bull. 154, 1931, 
pp. 66-74 (part). 

Cnemidophorus guttatus guttatus, Copr, Trans. Amer. Philos. Soc., vol. 17, 1892, 
p. 32.—Gapow, Proc. Zool. Soc. London, 1906, pp. 309, 325-326, fig. 75.— 
Hartwee and Ouiver, Oce. Pap. Mus. Zool. Univ. Michigan, No. 359, 1937, 
p. 3. 


180 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Cnemidophorus unicolor Corr, Proc. Amer. Philos. Soc., vol. 17, 1877, p. 93 
(U. S. Nat. Mus.; ‘‘West Tehuantepec,”’ Oaxaca; F. Sumichrast collector). 


Type.—Zool. Mus. Berlin; F. Deppe collector. 
Type locality —Mexico, here restricted to Veracruz, Veracruz. 
Range.—Atlantic slopes, from central Veracruz to the Isthmus of 
Tehuantepec. Recorded only from the states of Veracruz and 
Oaxaca. 
CNEMIDOPHORUS GUTTATUS IMMUTABILIS Cope 


Cnemidophorus immutabilis Corn, Proc. Amer. Philos. Soc., vol. 17, 1877, p. 93 

Cnemidophorus guttatus immutabilis, Corr, Trans. Amer. Philos. Soc., vol. 17 
1892, p. 31.—Gapow, Proc. Zool. Soc. London, 1906, pp. 309, 326-327 
figs. 74, 75.—HartTWEa and OLIvER, Occ. Papers Mus. Zool. Univ. Michigan 
No. 359, 1937, pp. 3-7. 

Cnemidophorus microlepidopus Cops, loc. cit. (U.S.N.M. No. 30187; “West 
Tehuantepec,” here restricted to the city of Tehuantepec, Oaxaca; F. 
Sumichrast collector). 

Cnemidophorus guttatus striatus Gapow, Proc. Roy. Soc. London, vol. 72, 1903, 
p. 115 (Brit. Mus. Nat. Hist., “Isthmus of Tehuantepec,” here restricted to 
the city of Tehuantepec, Oaxaca; Hans Gadow collector). 


Type.—U.S.N.M. No. 30141; F. Sumichrast collector. 

Type locality — ‘West Tehuantepec,’ Oaxaca, here restricted to 
the city of Tehuantepec. 

Range.—Pacific slopes from Colima to Chiapas. Recorded only 
from the states of Colima (Manzanillo), Michoac4n, Guerrero, Morelos, 
Oaxaca, and Chiapas (Tonala). 


SEXLINEATUS GROUP 


Species.—Six, with a total of 16 forms as at present recognized. 

Range.—Southern California, southeastern Wyoming, Wisconsin, 
and Maryland southward to the Gulf of Mexico, to Guatemala and 
into northwestern Baja California. 

Remarks.—The arrangement presented herewith for this group is 
far from final. We have, however, made an attempt to correlate 
zoogeography with variations pointed out by others, and the tentative 
conclusions we trust will at least be a step forward. The present pic- 
ture does make some geographic sense, as previous arrangements 
have not. 

CNEMIDOPHORUS SACKII SACKII Wiegmann 


Cnemidophorus sackii W1EGMANN, Herpetologia Mexicana, 1834, pp. 28-29.— 
Burt, U.S. Nat. Mus. Bull. 154, 1931, pp. 249-251.% 

Cnemidophorus sexlineatus sackii, Burt, Proc. Biol. Soc. Washington, vol. 44, 
1931, pp. 73-78. 

Cnemidophorus sackit sackit, Smiru, Journ. Washington Acad. Sci., vol. 39, 1949, 
pp. 41-42. 


§3 See also discussions in Taylor, Univ. Kansas Sci. Bull., vol. 24, 1938, pp. 520-523; Schmidt and Stuart, 
Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, 1941, pp. 244-255; and Stuart, Mise. Publ. Mus. Zool. Univ. 
Michigan, No. 69, 1948, pp. 57-59. 


CHECKLIST OF REPTILES OF MEXICO 181 


Cnemidophorus mexicanus Perers, Monatsh. Akad. Wiss. Berlin, 1869, pp. 62-63 
(Zool. Mus. Berlin, three cotypes; Uhde collector; ‘““Mexico,” ® here restricted 
to Matamoros, Puebla). 

Cnemidophorus costatus Corr, Proc. Amer. Philos. Soc., vol. 17, 1877, pp. 95-96 
(U.S.N.M. No. 31859; Boucard collector; ‘‘Mexico,’’ here restricted to 
Cuernavaca, Morelos). 

Cnemidophorus mexicanus balsas Gapow, Proc. Zool. Soc. London, 1906, pp. 363- 
367, fig. 83 (many cotypes, Chicago Nat. Hist. Mus. and Brit. Mus. Nat. 
Hist.; lectotype here designated as Brit. Mus. Nat. Hist. No. 1906.6.1.71; 
Hans Gadow collector; type locality here restricted to Cuernavaca, Morelos). 


Type.—Zool. Mus. Berlin No. 884; F. Deppe collector. 

Type locality.—‘‘ Mexico,” restricted to Cuernavaca, Morelos. 

Range.—The Rio Balsas Valley. Recorded from numerous locali- 
ties in Morelos, Guerrero, Puebla, and Michoacan. 


CNEMIDOPHORUS SACKII BOCOURTI Boulenger 


Cnemidophorus bocourti BouLENGER, Catalogue of the lizards in the British 
Museuni, vol. 2, 1885, p. 367. 

Cnemidophorus communis bocourti, Gapow, Proc. Zool. Soc. London, 1906, pp. 
356-358, fig. 80. 

Cnemidophorus sackii bocourti, SmirH, Rev. Soc. Mex. Hist..Nat., vol. 7, 1946, 
pp. 69-70. 

Cnemidophorus sexlineatus sackii, Bocourt, Mission scientifique au Mexique. . ., 
Etudes sur les reptiles, livr. 4, 1874, pp. 276-277, pl. 20e, fig. 6. 

Cnemidophorus sackii, Scumipt and Stuart, Publ. Field Mus. Nat. Hist., zool. 
ser., vol. 24, 1941, pp. 244-245. 

Cnemidophorus communis copeit, GADow, Proc. Zool. Soc. London, 1906, pp. 350— 
351 (specimen from Santo Domingo, Oaxaca, only). 

Cnemidophorus motaguae Sackett, Notulae Nat., No. 77, 1941, pp. 1-4 (Acad. 
Nat. Sci. Philadelphia No. 22143; J. T. Sackett collector; Motagua River 
Valley 10 kilometers northeast of Zacapa, Zacapa, Guatemala). 


Type.—Brit. Mus. Nat. Hist., three cotypes; No. 1857.10.28.81 here 
designated lectotype. 

Type locality —‘ Mexico” and ‘California,’ here restricted to Oax- 
aca, Oaxaca. 

Range.—Uplands from central Oaxaca through central Chiapas 
into central Guatemala; range perhaps discontinuous. Recorded in 
Mexico only from QOaraca: Oaxaca, Santo Domingo; and Chiapas: 
Piedra Parada. 


CNEMIDOPHORUS SACKII AUSTRALIS Gadow 


Cnemidophorus communis australis Gapow, Proc. Zool. Soc. London, 1906, pp. 
352-356, figs. 62C, D, 64C, D, 65F, 79B, C. 

Cnemidophorus mexicanus typica Gapow, Proc. Zool. Soc. London, 1906, pp. 
360-363, fig. 81A—D, fig. 82. 


% Gadow (op. cit., p. 358) states that Peters’s cotypes are immature and therefore not readily placed. The 
original description is of Jittle assistance. We do not follow Gadow’s allocation of the name, with a distinct 
southern race, for two reasons: (1) The name can simply be disposed of, if the types are unidentifiable, by 
placing it in synonymy; and (2) we believe it quite probable that the types actually came from southwestern 
Puebla in the Balsas Basin, where a number of Peters’s species are known to be restricted. This latter area 
is occupied by C. s. sackii, of which Peters’s name can reasonably be held a synonym. 


182 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Type.—Numerous cotypes, Brit. Mus. Nat. Hist. and Chicago Nat. 
Hist. Mus.; lectotype here designated as Brit. Mus. Nat. Hist. No. 
1906.7.19.11. 

Type locality —Not specified, Lagunas or Cuicatlan, Oaxaca; here 
restricted to Cuicatlan, Oaxaca. 

Range.—Foothills about the central Oaxacan highlands, excluding 
the Balsas Basin, in the upper headwaters of the Rfo Papaloapam 
and Rio Tehuantepec, and near the Isthmus of Tehuantepec. 


CNEMIDOPHORUS SACKII COMMUNIS Cope 


Cnemidophorus communis Corr, Proce. Amer. Philos. Soc., vol. 17, 1877, p. 95. 

Cnemidophorus communis occidentalis Gapow, Proc. Zool. Soc. London, 1906, pp. 
339-346, fig. 69, 77A—-F, 78B, 79A (many cotypes, Brit. Mus. Nat. Hist. 
and Chicago Nat. Hist. Mus.; lectotype here designated as Brit. Mus. Nat. 
Hist. No. 1892.2.8.33, from Ixtlin, Nayarit). 

Cnemidophorus communis copei Gapow, Proc. Zool. Soc. London, 1906, pp. 346—- 
352, fig. 78A, C, E (cotypes in Brit. Mus. Nat. Hist. and Chicago Nat. Hist. 
Mus.; lectotype here designated as Brit. Mus. Nat. Hist. No. 1906.7.19.5, 
from Colima, Colima; a cotype from Santo Domingo de Guzm4n we refer to 
C. s. bocourti). 


Type.—Numerous cotypes, all lost. 

Type locality —Not definitely stated: Colima, Guadalajara, Cér- 
doba, Coban (Guatemala) or San Antonio (Texas); here restricted to 
Colima, Colima. 

Range.—Most of the central plateau of Mexico, from Chihuahua 
and presumably Nuevo Leén southward to central Puebla; the Pacific 
coast from the Balsas basin northward to southern Sinaloa and perhaps 
Sonora. Recorded from ? Sonora, Chihuahua, Durango, Coahuila, 
Tamaulipas, San Luis Potosi, Zacatecas, Hidalgo, Querétaro, Guana- 
juato, Aguascalientes, Jalisco, Sinaloa, Nayarit (including Isabel Is- 
land), Colima, Michoacan, México, Distrito Federal, Puebla, and 
Veracruz. 

CNEMIDOPHORUS SACKII SCALARIS Cope 
Cnemidophorus sexlineatus tigris Cope (nec Baird and Girard), Proc. Amer. 


Philos. Soc., vol. 23, 1886, p. 283 (types not designated; Chihuahua, 
Chihuahua). 


Cnemidophorus gularis scalaris Corr, Trans. Amer. Philos. Soc., vol. 17, 1892, 
p. 47.—Smira and Mirrteman, Trans. Kansas Acad. Sci., vol. 46, 1943, 
pp. 246-247. 


Cnemidophorus gularis gularis verus Copn, op. cit., p. 45 (no types designated; 
Chihuahua, Chihuahua). 


Cnemidophorus gularis gularis obsoletus Cops, loc. cit. (as above). 
Type-—U.S.N.M. Nos. 8319, 14302, seven cotypes (No. 14302a, 
lectotype); John Potts collector. 
Type locality —Not specifically designated, either ‘‘ Mexican plateau 


south of Chihuahua,” or ‘City of Chihuahua;” here restricted to 
Chihuahua, Chihuahua. 


CHECKLIST OF REPTILES OF MEXICO 183 


Range.—Central and eastern Chihuahua, western Coahuila and 
northern Durango. Recorded from each state mentioned. 


CNEMIDOPHORUS SACKII STICTOGRAMMUS Burger 


Cnemidophorus sackii stictogrammus, BurGER, Chicago Acad, Sci. Nat. Hist. Misc., 
No. 65, 1950, p. 5. 


Cnemidophorus gularis octolineatus, SmiTH (nec Baird), Handbook of lizards, 1946, 
pp. 409-412, figs. 126-127, pl. 114 (part). 

Type.—U.S.N.M. (Hensley-Burger No. 768). 

Type locality—Yank Springs, 6 miles southeast of Ruby, Santa 
Cruz County, Ariz. 

Range.—Central Arizona and Mexico southward through Sonora 
and probably northern Sinaloa, and the northern parts of Chihuahua 
and Coahuila. Recorded in Mexico from Sonora and Chihuahua. 


CNEMIDOPHORUS SACKII MARIARUM Giinther 


Cnemidophorus mariarum GUNTHER, Biologia Centrali-Americana, Reptilia and 
Batrachia, 1885, p. 28.—STEsnecER, North Amer. Fauna, No. 14, 1899, pp. 
67-68.—Gapvow, Proc. Zool. Soc. London, 1906, pp. 328-330. 


Type.—Brit. Mus. Nat. Hist. Nos. 1881.10.1.81-82, 86-88, five 
cotypes. 
Type locality —Tres Marias Islands. 


Range.—Restricted to the Tres Marias Islands, on each one of which 
it has been taken. 


CNEMIDOPHORUS SACKII ANGUSTICEPS Cope 


Cnemidophorus angusticeps Corr, Proc. Amer. Philos. Soc., vol. 17, 1877, p. 
95.—Smiru, Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, 1939, pp. 24-25. 


Type.—U.S.N.M. Nos. 24876-24878, three cotypes. 

Type locality.—‘ Yucatan,” here restricted to Chichen Itza. 

Range.——The northern and western portions of the Yucatan 
Peninsula. Recorded from Yucatdén: Chichen Itza, Progreso, Tunkas; 
Quintana Roo: Coba, Cozumel Island; Campeche: Champotén. 


CNEMIDOPHORUS SACKII GULARIS Baird and Girard 


Cnemidophorus gularis Batrp and Girarp, Proc. Acad. Nat. Sci. Philadelphia, 
1852, p. 128.. 

Cnemidophorus gularis gularis, Copr, Trans. Amer. Philos. Soc., vol. 17, 1892, p. 
334.—Smitu, Handbook of lizards, 1946, pp. 406-409, fig. 121, pl. 113. 

Cnemidophorus guttatus HALLOWELL (nec Wiegmann), Proc. Acad. Nat. Sci., 
Philadelphia, 1854, p. 192 (types lost; Texas, here restricted to Brownsville). 

Cnemidophorus gularis sealous Copr, Amer. Nat., vol. 26, 1892, p. 522 (nomen 
nudum). 

Cnemidophorus gularis sericeus Copr, Trans. Amer. Philos. Soc., vol. 17, 1892, 
p. 48 (U.S.N.M. No. 15650; Wm. Taylor collector; San Diego, Tex.). 


184 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Cnemidophorus gularis meeki Gapow, Proc. Zool. Soc. London, 1906, pp. 332-334 
(numerous cotypes, Brit. Mus. Nat. Hist. and Chicago Nat. Hist. Mus.; 
lectotype here designated as Chicago Nat. Hist. Mus. No. 1294, the larger 
male specimen, from Montemorelos, Nuevo Leén). 

Type.—U.S.N.M. Nos. 3022, 2989, 14 cotypes (lectotype No. 
3022a); Colonel Graham collector. 
Type locality.—‘‘Indianola and the Valley of the Rio Grande del 

Norte,” restricted to mouth of Devils River, Tex. 

Range.—Oklahoma and most of Texas southward through Tamau- 
lipas and eastern Nuevo Leén to northern Veracruz. Recorded in 

Mexico from Tamaulipas, San Luis Potosi, Veracruz, and Nuevo Leén. 


CNEMIDOPHORUS SACKII SEMIFASCIATUS Cope 


Cnemidophorus gularis semifasciatus CorpE, Trans. Amer. Philos. Soe., vol. 17, 
1892, p. 49. 

Cnemidophorus sackii semifasciatus, BURGER, Chicago Acad. Sci. Nat. Hist. Misc., 
No. 65, 1950, pp. 4-5. 

Cnemidophorus septemvittatus Corr, Trans. Amer. Philos. Soc., vol. 17, 1892, p. 40 
(“El Dorado County, California,” in error; restricted to Marfa, Presidio 
County, Tex.; U.S.N.M. No. 42141). 

Cnemidophorus octolineatus Scumipt and SmirH (nec Baird), Publ. Field Mus. 
Nat. Hist., zool. ser., vol. 29, 1944, pp. 85-86. 


Type.—U.S.N.M. No. 9248; Lieutenant Couch collector. 

Type locality Agua Nueva, Coahuila. 

Range.—Big Bend area of Texas and adjacent Coahuila. Recorded 
in Mexico only from the state of Coahuila. 


CNEMIDOPHORUS INORNATUS Baird 


Cnemidophorus inornatus Barirp, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 
255.—Scumipt and Owens, Publ. Field Mus. Nat. Hist., zool. ser., vol. 
29, 1944, pp. 106-107; BurcEerR, Chicago Acad. Sci. Nat. Hist. Misc., 1950 
(in press). 

Cnemidophorus octolineatus Batrp, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 
255 (Pesqueria Grande, Nuevo Leén; U.S.N.M. No. 3009; Lieutenant 
Couch collector). 

Cnemidophorus perplecus (nec Batrp and GrrARD), VAN DeNnBuRGH, Occ. Pap. 
California Acad. Sci., No. 10, 1922, pp. 495-497.—Scumipr and Smriru, 
Publ. Field Mus. Nat. Hist., zool. ser., vol. 29, 1944, pp. 86-87.—Smiru, 
Handbook of lizards, 1946, pp. 412-414, pl. 15. 

Cnemidophorus arizonae VAN DENBURGH, Proc. California Acad. Sci., ser. 2, 
vol. 6, 1896, p. 344 (Stanford Univ. No. 2631; W. W. Price, collector; Fair- 
bank, Cochise County, Ariz.). 

Cnemidophorus gularis velox SPRINGER, Copeia, No. 169, 1928, p. 102 (Butler 
Univ. No. 848; Pueblo Bonito, San Juan County, N. Mex., by present 
restriction). 


Type.—U.S.N.M. No. 3032; Lieutenant Couch collector. 

Type locality—Pesqueria Grande (=Garcia), Nuevo Le6én. 

Range.—Western Texas to extreme southeastern Arizona, south- 
ward into Chihuahua, Coahuila, and Nuevo Leén. Recorded in 
Mexico only from the states cited. 


CHECKLIST OF REPTILES OF MEXICO 185 


CNEMIDOPHORUS BURTI Taylor 


Cnemidophorus burtt Taytor, Univ. Kansas Sci. Bull., vol. 24, 1938, pp. 485-487, 
pl. 42, fig. 2. 


Type.—EHT-HMS No. 13117; E. H. Taylor collector. 
Type locality.—La Posa, 10 miles northwest of Guaymas, Sonora. 
Range.—Southwestern Sonora. Known only from the vicinity 
of Guaymas. 
CNEMIDOPHORUS SEXLINEATUS (Linnaeus) 


Lacerta sexlineata LinNaxEvs, Systema naturae, ed. 12, vol. 3, 1766, p. 364. 

Cnemidophorus sexlineatus, DuMi&RIL and Brsron, Erpétologie générale, vol. 5, 
1839, p. 131.—Taytor, Univ. Kansas Sci. Bull., vol. 24, 1939, pp. 520-522.— 
Smitu, Handbook of lizards, 1946, pp. 415-418, figs. 13, 122, pl. 116. 

Cnemidophorus sexlineatus sexlineatus, Corr, Ann. Rep. U. 8. Nat. Mus., 1898 
(1900), pp. 598, 603-605.— Burt, U.S. Nat. Mus., Bull. 154, 1931, pp. 76-97, 
fig. 20. 

?Cnemidophorus perplecus BatrD and GrrarD, Proc. Acad. Nat. Sci. Philadelphia, 
1852, p. 128 (U. S. N. M. No. 3060); Wm. Gambell collector; type locality 
unknown). 


Type.—Not known. 

Type locality —‘‘ Carolina,” here restricted to Charleston, 8. C. 

Range.—Eastern United States from Wisconsin, southeastern 
Wyoming, and Maryland southward to the Gulf as far west as, per- 
haps, the mouth of the Rio Grande. No definitely reliable records 
are known for Mexico, but we are informed by Bryce Brown that the 
species occurs near Brownsville, Tex., and may be expected across the 
river in Tamaulipas. Records available from Matamoros, Tamauli- 
pas, may or may not belong here. 


CNEMIDOPHORUS GADOVI Burger 


Cnemidophorus gadovii Burcer, Chicago Acad. Sci. Nat. Hist. Misc., No. 65, 
1950, p. 2. 


Type—U.S.N.M. No. 40042. 
Type locality—Hermosillo, Sonora. 
Range.—Known only from the type locality. 


CNEMIDOPHORUS LABIALIS Stejneger 


Cnemidophorus labialis StpsnEcER, Proc. U. S. Nat. Mus., vol. 12, 1890, pp. 
643-644.—Corr, Ann. Rep. U. S. Nat. Mus., 1898 (1900), pp. 610-612, fig. 
122.— Burt, U.S. Nat. Mus. Bull. 154, 1931, pp. 141-146.—Tevis, Copeia, 
1944, pp. 16-17. 


Type.—U.S.N.M. No. 15596; L. Belding collector. 
Type locality.—‘‘Cerros Island,’”’ Pacific coast, Baja California. 


Range.—Northwestern Baja California between San Quintin and 
Punta Eugenia, Cedros Island. 


186 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


HYPERYTHRUS GROUP 


Species.—One, represented by six forms as at present recognized. 
Range.—Extreme southwest California south throughout the penin- 
sula of Baja California and on adjacent islands. 


CNEMIDOPHORUS HYPERYTHRUS BELDINGI (Stejneger) 


Verticaria beldingi StesNEGER, Proc. U. 8. Nat. Mus., vol. 17, 1894, p. 17. 

Verticaria hyperythra beldingi, VAN DENBURGH, Occ. Pap. California Acad. Sci., 
No. 10, 1922, pp. 560-563, pl. 55. 

Cnemidophorus hyperythrus beldingi, GRINNELL and Camp, Univ. California Publ. 
Zool., vol. 17, 1917, p. 175. 


Type.—U.S.N.M. No. 11980; L. Belding collector. 

Type locality —‘Cerros” Island, Baja California. 

Range.—Extreme southwestern California southward on Pacific 
slopes (not Gulf) to Cedros Island. 


CNEMIDOPHORUS HYPERYTHRUS CAERULEUS (Dickerson) 


Verticaria caerulea DicxERSoN, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, p. 472. 
Cnemidophorus hyperythrus caeruleus, Burt, U.S. Nat. Mus. Bull. 154, 1931, pp. 
240-242. 


Type.—Amer. Mus. Nat. Hist. No. 5517; C. H. Townsend collector. 
Type locality —Carmen Island, Baja California. 
Range.—Known only from the type locality. 


CNEMIDOPHORUS HYPERYTHRUS DANHEIMAE Burt 


Verticaria sericea VAN DernxpurRGH, Proc. California Acad. Sci., ser. 2, vol. 5, 
1895, pp. 132-133, pl. 12. 

Cnemidophorus hyperythrus danheimae Burt, Proc. Biol. Soc. Washington, vol. 42, 
1929, p. 154 (new name for Verticaria sericea Van Denburgh, 1895, pre- 
occupied by Cnemidophorus gularis sericeus Cope, 1892); U. S. Nat. Mus. 
Bull. 154, 1931, pp. 244-246. 


Type.—California Acad. Sci. No. 435; Walter E. Bryant collector. 
Type locality.—San José Island, Gulf of California, Baja California. 
Range.—Known only from the type locality. 


CNEMIDOPHORUS HYPERYTHRUS PICTUS (Van Denburgh and Slevin) 


Verticaria picta VAN DenxpurcH and S.LEviNn, Proc. California Acad. Sci., ser. 4, 
vol. 11, 1921, p. 98. 

Cnemidophorus hyperythrus pictus, Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 
242-244, 


Type.—California Acad. Sci. No. 49155; Joseph R. Slevin collector. 
Type locality —Monserrate Island, Baja California. 
Range.—Known only from the type locality. 


CHECKLIST OF REPTILES OF MEXICO 187 


CNEMIDOPHORUS HYPERYTHRUS SCHMIDTI (Van Denburgh and Slevin) 


Verticaria hyperythra schmidt VAN DrENBURGH and SLEVIN, Proc. California 
Acad. Sci., ser. 4, vol. 11, 1921, p. 397.— Van Densureu, Occ. Pap. California 
Acad. Sci., No. 10, 1922, pp. 563-566. 

Cnemidophorus hyperythrus schmidti, LinspaLEe, Univ. California Publ. Zool., 
vol. 38, 1932, p. 373. 


Type.—California Acad. Sci. No. 50512; Joseph R. Slevin collector. 

Type locality—San Marcos Island, Gulf of California, Baja Cali- 
fornia. 

Range.—The central fifth of the peninsula of Baja California from 
about lat. 28° south to 25° 30’ N. (the Vizcaino Desert south to 
Comondi), and adjacent islands. 


CNEMIDOPHORUS HYPERYTHRUS HYPERYTHRUS Cope 


Cnemidophorus hyperythrus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1863, p. 103. 

Verticaria hyperythra hyperythra, VAN DensureH, Occ. Pap. California Acad. 
Sci., No. 10, 1922, pp. 556-559. 

Cnemidophorus hyperythrus hyperythrus, Burt, Proc. Biol. Soc. Washington, 
vol. 42, 1929, p. 154 (part); U. S. Nat. Mus., Bull. 154, 1931, pp. 226-240 
(part).—Linspauxz, Univ. California Publ. Zool., vol. 38, 1932, pp. 372-373. 

Verticaria espiritensis VAN DENBURGH and SLEVIN, Proc. California Acad. Sci., 
ser. 4, vol. 11, 1921, p. 397 (Calif. Acad. Sci. No. 50511; Espfritu Santo 
Island, Baja California; J. R. Slevin collector).—Van DrENBURGH, op. cit., 
pp. 566-568. 

Verticaria franciscensis VAN DENBURGH and SLEvIN, loc. cit. (Calif. Acad. Sci. 
No. 50513; San Francisco Island, Baja California; J. R. Slevin collector).— 
Van DENBURGH, op. cit., pp. 568-570. 


Type.—U.S.N.M. No. 5299; John Xantus collector. 

Type locality —Cape San Lucas, Baja California. 

Range.—The southern third of the peninsula of Baja California 
and its adjacent islands on both the Gulf and Pacific sides, south 
from about lat. 25°30’. 

TESSELATUS GROUP 

Species.—Six, one of which is represented by eleven subspecies, as 
now recognized. 

Range.—Western United States from Idaho and Oregon south 
through Baja California and western Texas to southern Coahuila, 
Chihuahua, and Sonora. 


CNEMIDOPHORUS BACATUS Van Denburgh and Slevin 


Cnemidophorus bacatus VAN DenBURGH and SLEvin, Proc. California Acad. Sci., 

ser. 4, vol. 11, 1921, p. 97.—Van DeEnsBouRGH, Occ. Pap. California Acad. Sci., 
No. 10, 1922, pp. 544-546.—Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 
211-213. 

Type.—California Acad. Sci. No. 49152; Joseph R. Slevin collector. 

Type locality—San Pedro Nolasco Island, Sonora. 

Range.—Known only from the type locality. 

861316—50-——13 


188 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 
CNEMIDOPHORUS CATALINENSIS Van Denburgh and Slevin 


Cnemidophorus catalinensis VAN DENBURGH and SLEvin, Proc. California Acad. 
Sci., ser. 4, vol. 11, p. 396.—Van DeEnrurau, Occ. Pap. California Acad. Sci., 
No. 10, 1922, pp. 542-544.—Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 
213-216, fig. 29. 
Type.—California Acad. Sci. No. 50507; Joseph R. Slevin collector. 
Type locality.—Santa Catalina Island, Baja California. 
Range.—Known only from the type locality. 


CNEMIDOPHORUS CERALBENSIS (Van Denburgh and Slevin) 


Verticaria ceralbensis VAN DENBURGH and SLEviN, Proc. California Acad. Sci. 
ser. 4, vol. 11, 1921, p. 396.—Van Drnpurau, Occ. Pap. California Acad: 
Sci., No. 10, 1922, pp. 554-556. 

Cnemidophorus ceralbensis, Burt, U. 8. Nat. Mus. Bull. 154, 1931, pp. 216-218. 


- Type.—California Acad. Sci. No. 50510; Joseph R. Slevin collector. 
Type locality. —Ceralbo Island, Baja California. 
Range.—Known only from the type locality. 


CNEMIDOPHORUS TESSELATUS (Say) 


Ameiva tesselata Say, in Long’s Expedition to the Rocky Mountains, vol. 2, 1823, 
. 60. 

Cae tesselatus, SmirH and Buraer, Bull. Chicago Acad. Sci., vol. 8, 
1949, p. 282. 

Cnemidophorus grahamiit Baird and Grirarp, Proc. Acad. Nat. Sci. Philadelphia, 
1852, p. 128 (U.S.N.M. No. 3036, 2 cotypes; Colonel Graham collector; 
between San Antonio and El Paso, Texas, here restricted to Fort Davis).— 
SrrecKER, Baylor Univ. Bull. No. 13, 1910, pp. 8-13, pl. 1.—Smirxs, Hand- 
book of lizards, 1946, pp. 419-421, pl. 117. 

Type.—Lost. 

Type locality.k—Arkansas River, near Castle Rock Creek, Colo. 
(= Beaver Creek, Fremont County). 

Range.—Western Texas, eastern New Mexico, and probably southern 
Colorado, southward into adjacent northern Chihuahua and Coahuila. 
No reliable Mexican records are known to us; Yarrow (U.S. Nat. Mus. 
Bull. 24, 1883, p. 43) records it from “between Panos and St. Luis, 
Mexico,” and Cope (U. S. Nat. Mus. Bull. 32, 1887, p. 45) cites it 
from ‘‘between Mexico City and Chihuahua,” but neither record 
definitely refers to the species as now understood, and even if they did, 
the locality of collection is very uncertain. 


CNEMIDOPHORUS MAXIMUS Cope 


Cnemidophorus maximus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1863, p. 104; 
Ann. Rep. U. S. Nat. Mus., 1898 (1900), pp. 570-571, fig. 104.—Van DEn- 
BURGH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 506—508.—Burt, 
U.S. Nat. Mus. Bull. 154, 1931, pp. 218-221. 


Type.—U.S.N.M. No. 5297; John Xantus collector. 


CHECKLIST OF REPTILES OF MEXICO | 189 


Type locality.—Cape San Lucas, Baja California. 
Range.—Southern Baja California, southward from La Paz Bay, or 
perhaps Magdalena Bay. 


CNEMIDOPHORUS TIGRIS TIGRIS Baird and Girard 


Cnemidophorus tigris BatrD and GriraRp, Proc. Acad. Nat. Sci. Philadelphia, 
1852, p. 69. 

Cnemidophorus tigris tigris, Camp, Univ. California Publ. Zool., vol. 17, 1916, 
pp. 71-72.—Smira and Buraer, Bull. Chicago Acad. Sci., vol. 8, 1949, 
p. 282. 

Cnemidophorus tessellatus tessellatus, Cope (nec Say), U. S. Nat. Mus. Bull. 1, 
1875, p. 46.—Van DensBurGH, Occ. Pap. California Acad. Sci., No. 10, 
1922, pp. 508-516, pl. 23.—Burt, U. 8S. Nat. Mus. Bull. 154, 1922, pp. 
146-199 (part).—Smiru, Handbook of lizards, 1946, pp. 421-424, pl. 118. 


Type.—U.S.N.M. No. 4103; Capt. Howard Stansbury collector. 

Type locality.—‘“‘Valley of the Great Salt Lake,’’ Utah, restricted to 
Salt Lake City, Utah. 

Range.—Southern Idaho and southeastern Oregon south through 
western Utah and southeastern California to extreme northeastern 
Baja California, and extreme northwestern Sonora. Recorded in 
Mexico only from Baja California and Sonora (between Sonoyta and 
Puerto Pefiasco). 


CNEMIDOPHORUS TIGRIS GRACILIS Baird and Girard 


Cnemidophorus gracilis Barrp and GrirarD, Proc. Acad. Nat. Sci. Philadelphia, 
1852, p. 69. 

Cnemidophorus tigris gracilis, BurerR, Chicago Acad. Sci. Nat. Hist. Misc., No. 
65, 1950, p. 6. 

Cnemidophorus melanostethus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1863, 
p. 104 (U.S.N.M. No. 3067, 12 cotypes; H. B. Mollhausen collector; “region 
of the Colorado, California,” here restricted to Yuma, Ariz.). 


Type.—U.S.N.M. No. 3034; J. L. LeConte collector. 

Type locality.—Desert of Colorado,” restricted to Yuma, Ariz. 

Range.—Extreme eastern California in Colorado River Valley, 
southern Arizona south of the plateau, extreme southwestern New 

Mexico, northern Sonora. Recorded in Mexico only from the state 

of Sonora. 

CNEMIDOPHORUS TIGRIS AETHIOPS Cope 

Cnemidophorus tessellatus aethiops Corr, Ann. Rep. U. S. Nat. Mus., 1898 (1900), 
p. 582.—Smiru, Handbook of lizards, 1946, pp. 424-426, pl. 119 (part). 

Cnemidophorus tigris aethiops, SmirH and Burcer, Bull. Chicago Acad. Sci., 
vol. 8, 1949, p. 282. 

Cnemidophorus disparilis Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, 
p. 473 (U.S.N.M. No. 64445; C. H. Townsend collector; Tiburén Island, 
Sonora). 

Cnemidophorus estebanensis DicKERSON, op. cit., p. 474 (U.S.N.M. No. 64446; 
C. H. Townsend collector; San Esteban Island, Sonora). 

Cnemidophorus punctilineatus DicKERSON, op. cit., p. 475 (U.S.N.M. No. 64447; 
C. H. Townsend collector; Tiburén Island, Sonora). 


190 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Cnemidophorus melanostethus, VaN DernBuRGH, Occ. Pap. California Acad. 
Sci., No. 10, 1922, pp. 529-533 (part). 


Type-—U.S.N.M. Nos. 64240-5, cotypes; Jenkins and Evermann 
collectors. 

Type locality—Hermosillo, Sonora. 

Range.—Southern Sonora and adjacent islands (Tiburén, San 
Esteban). 


CNIMIDOPHORUS TIGRIS MARMORATUS Baird and Girard 


Cnemidophorus marmoratus Batrp and Grirarp, Proc. Acad. Nat. Sci. Philadel- 
phia, 1852, p. 128. 

Cnemidophorus tigris marmoratus, BurRGER, Chicago Acad. Sci. Nat. Hist. Misc., 
No. 65, 1950, p. 7. 

Cnemidophorus variolosus Corr, Proc. Acad. Nat. Sci. Philadelphia, 1863, p. 39 
(U.S.N.M. No. 3066; Lieutenant Couch collector; Parras, Coahuila). 


Type.—U.S.N.M. No. 3024; two cotypes; J. H. Clark collector. 

Type locality —‘Between San Antonio and El Paso,’ restricted to 
El Paso, Tex. 

Range.—Southwestern New Mexico to southwestern Coahuila. 
Recorded in Mexico only from Chihuahua, Durango (5 km. west of 
Torreén), and Coahuila. 


CNEMIDOPHORUS TIGRIS CANUS Van Denburgh and Slevin 


Cnemidophorus canus VAN DENBURGH and SLEvIN, Proc. California Acad. Sci., 
ser. 4, vol. 11, No. 6, 1921, p. 97.—Van DENnBuRGH, Occ. Pap. California Acad. 
Sci., vol. 10, 1922, pp. 540-542. 

Cnemidophorus tessellatus canus, Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 
208-211. 

Cnemidophorus tigris canus, SMitH and BurGER, Bull. Chicago Acad. Sci., vol. 8, 
1949, p. 282. 


Type.—California Acad. Sci. No. 49153; Joseph R. Slevin collector. 

Type locality—Sal Si Puedes Island, Gulf of California, Baja 
California. 

Range.—The type locality, and North and South San Lorenzo 
Islands (on the latter, intergrades with C. t. martyris). 


CNEMIDOPHORUS TIGRIS CELERIPES Dickerson 


Cnemidophorus celeripes DickErson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, 
p. 472.—Van Denzoureu, Occ. Pap. California Acad. Sci., vol. 10, 1922, pp. 
549-551. 

Cnemidophorus tessellatus celeripes, Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 
202-205. 

Cnemidophorus tigris celeripes, SMitTH and BurcsEr, Bull. Chicago Acad. Sci., 
vol. 8, 1949, p. 282. 


Type.—Amer. Mus. Nat. Hist. No. 5514; C. H. Townsend collector. 
Type locality —San José Island, Baja California. 
Range.—Known only from the type locality. 


CHECKLIST OF REPTILES OF MEXICO 191 


CNEMIDOPHORUS TIGRIS MARTYRIS Siejneger 


Cnemidophorus martyris STEJNEGER, Proc. U.S. Nat. Mus., vol. 14, 1891, p. 407.— 
Van Densoures, Occ. Pap. California Acad. Sci., vol. 10, 1922, pp. 5388-540. 

Cnemidophorus tessellatus martyris, Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 
205-208. 

Cnemidophorus tigris martyris, SMira and Burcer, Bull. Chicago Acad. Sci.,. 
vol. 8, 1949, p. 282. 


Type.—U.S.N.M. No. 15620; E. Palmer collector. 

Type locality San Pedro Martir Island, Gulf of California, Sonora. 

Range.—The type locality (intergrades with C. t. canus on South 
San Lorenzo Island). 


CNEMIDOPHORUS TIGRIS RUBIDUS Cope 


Cnemidophorus tessellatus rubidus Corr, Trans. Amer. Philos. Soc., ser. 2, vol. 17, 
1892, p. 36, pl. 12, fig. 1—Burr, U.S. Nat. Mus. Bull. 154, 1931, pp. 199-202. 

Cnemidophorus rubidus, VAN DENBURGH, Occ. Pap. California Acad. Sci. vol. 10, 
1922, pp. 546-549. 

Cnemidophorus tigris rubidus, SMirH and Buresr, Bull. Chicago Acad. Sci., vol. 
8, 1949, p. 282. 

Cnemidophorus vandenburght Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 
1919, p. 477 (U.S.N.M. No. 64449; C. H. Townsend collector; Carmen 
Island, Baja California). 


Type.—U.S.N.M. Nos. 15149-15155, cotypes; U. S. Fish Commis- 
sion collector. 

Type locality.—Santa Margarita Island, Baja California. 

Range.—The southern third of Baja California, south of San 
Marcos Island, including adjacent islands in the Gulf and off the 
western edge of the peninsula (notably Carmen, Santa Margarita, 
and Magdalena Islands, Baja California). 


CNEMIDOPHORUS TIGRIS MULTISCUTATUS Cope 


Cnemidophorus tessellatus multiscutatus Corr, Trans. Amer. Philos. Soc., vol. 17, 
1892, p. 38. 

Cnemidophorus stejnegeri VAN DreNnBuRGH, Proc. California Acad. Sci., ser. 2, 
vol. 4, 1894, p. 300 (Stanford Univ. Mus. No. 1861; Ensenada, Baja Cali- 
fornia; J. M. Stovell collector). 

Cnemidophorus tessellatus stejnegeri, STEJNEGER and BarsBour, Check list North 
American Amphibia and Reptilia, ed. 1, 1917, p. 68.—Van Dernsurcu, 
Oce. Pap. California Acad. Sci., vol. 10, 1922, pp. 519-523, pl. 54.—Smrru, 
Handbook of lizards, 1946, pp. 426-428, pl. 120. 

Cnemidophorus bartolomas Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 
1919, p. 476 (U.S.N.M. No. 64448; C. H. Townsend collector, San Barto- 
lomé Bay, Baja Calfornia). 

Cnemidophorus dickersonae VAN DeNBURGH and SLEVIN, Proc. California Acad. 
Sci., ser. 4, vol. 11, 1921, p. 97 (California Acad. Sci. No. 49154; J. R. Slevin 
collector; Isla Partida near Angel de la Guarda Island, Baja California). 


Type.—U.S.N.M. Nos. 15160-3; U. S. Fish Commission donor. 
Type locality.—Cedros Island, Baja California. 


192 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Range.—Southern California west of the Sierra Nevada, south to 
central Baja California, exclusive of the extreme northeastern corner; 
islands in the Gulf of California near the center of the Peninsula 
(Isla Partida, Angel de la Guarda). 


Genus GYMNOPHTHALMUS Merrem 


Gymnophthalmus Mrrrem, Tentamen systematis Amphibiorum, 1820, p. 74.— 
Stuart, Occ. Pap. Mus. Zool. Univ. Michigan No. 409, 1939, pp. 1-10, pl. 1. 
Epaphelus Corr, Journ. Acad. Nat. Sci. Philadelphia, 1875, p. 115 (type, Epa- 
phelus sumichrasti Cope). 
Blepharactisis HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, 1860, p. 484 
(type, Blepharactisis speciosa Hallowell). 
Genotype.—Lacerta quadrilineata Linnaeus [=Gymnophthalmus lin- 
eata (Linnaeus)]. . 
Range.—Central Argentina north through South and Central 
America. In Mexico, Oaxaca and possibly Chiapas. 
Species.—Seven. Only one occurs in Mexico. 


GYMNOPHTHALMUS SUMICHRASTI (Cope) 


Epaphelus sumichrasti Corr, Journ. Acad. Nat. Sci. Philadelphia, 1876, p. 115. 

Gymnophthalmus sumichrasti,*. Bocourt, Mission scientifique au Mexique . . ., 
Etudes sur les reptiles, livr. 7, 1881, pp. 471-472, pl. 22H, fig. 2, 2a-2h.— 
BovuLEencER, Catalogue of the lizards in the British Museum, vol. 2, 1885, 
pp. 428-429.—Srvart, Occ. Pap. Mus. Zool. Univ. Michigan, No. 409, 1939, 
pp. 5-6: 

Type-—U.S.N.M. No. 30245-46; Francois E. Sumichrast collector. 

Type locality.—‘ Western part of the State of Tehuantepec’ (‘Near 
Ventose Bay” fide Dunn in Stuart, loc. cit.). 

Range.—Southern Oaxaca near the Isthmus of Tehuantepec east- 
ward on Pacific slopes possibly to Honduras. Recorded in Mexico 
only from Oaxaca: Tehuantepec, Bahia Ventosa, Mount Guengola, 
Cacoprieto. A specimen, presumably of this species, was seen near 
San Ricardo, Chiapas, by the junior author. 


Family HELODERMIDAE Gray 


Helodermidae Gray, Proc. Zool. Soe. London, 1837, p. 132. 


Genera.—A single genus, Heloderma Wiegmann, is known. 
Range.—Southern Utah to the Isthmus of Tehuantepec, on Pacific 


slopes. 
Genus HELODERMA Wiegmann 


Trachyderma WiEGMANN (nec Latreille), Isis von Oken, vol. 22, 1829, p. 421 (type, 
Trachyderma horridum Wiegmann). 
Heloderma WrEIaMANN, Isis von Oken, vol. 22, 1829, p. 624. 


% Gymnodactylus sumichrasti Gadow, Proc. Zool. Soc. London, 1905, p. 211, from the Isthmus of Tehuan- 
tepec, appears to be a lapsus for this species. 


CHECKLIST OF REPTILES OF MEXICO 193 


Genotype —Trachyderma horridum Wiegmann. 
Range.—Southern Utah to the Isthmus of Tehuantepec. 


Species—Two. 
KEY TO SPECIES OF HELODERMA 


1. Tail equal to or longer than body; head and neck about half of body length; 
predominating color black; pterygopalatine teeth present __horridum (p. 193) 

Tail about two-thirds of body length; head and neck about one-third of body 
length; colors pink, yellowish, and black; none or rarely 1 or 2 pterygo- 

p CYC sre eg 7 5 fo pS aU = Lg IL cepa suspectum (p. 193) 


HELODERMA SUSPECTUM Cope 


Heloderma suspectum Corxz, Proc. Acad. Nat. Sci. Philadelphia, 1869, pp. 5-6; 
Ann. Rep. U. S. Nat. Mus., 1898 (1900), pp. 476-483, fig. 87—Smrta, 
Handbook of lizards, 1946, pp. 472-475, pl. 134. 

Type-—U.S.N.M. No. 2971, three cotypes; Major Emory collector. 
Type locality.—Sierra de la Unidén, ‘‘Sonora’’(= Arizona). 
Range.—Sonora as far south as Guaymas; in the United States, 

Arizona, southern Nevada, and southwestern Utah. Recorded from 

Sonora: Guadelupe Cafion, San Pedro Bay, La Posa about 10 miles 

northwest of Guaymas, Monument 88, San Bernardino, Niggerhead 

Mountain, 10 miles south of Noria, ete. 


HELODERMA HORRIDUM (Wiegmann) 


Trachyderma horridum Wi1EGMANN, Isis von Oken, vol. 22, 1829, p. 421. 

Heloderma horridum, W1EGMANN, Isis von Oken, vol. 22, 1829, p. 628; Herpeto- 
logia Mexicana, pt. 1, 1834, pp. 24-25, pl. 1. —Ginruer, Biologia Centrali- 
Americana, Reptilia and Batrachia, 1885, pp. 43-44, pl. 26.—Smirx, Univ. 
Kansas Sci. Bull., vol. 22, 1935, pp. 145-146. 

Heloderma Hernandesii W1EGMANN, Herpetologia Mexicana, pt. 1, 1834, p. 25 
(for a theoretical form). 


Type.—Zool. Mus. Berlin; Ferdinand Deppe collector. 

Type locality.—‘‘ Vivit in ferventibus terrae Mexicanae regionibus,” 
here restricted to Huajintlan, Guerrero. 

Range.—Coastal areas from Oaxaca to Sonora, extending up 
the Rio Balsas Valley to Morelos.*® Recorded from Sonora: 
Alamos, Guirocoba; Sinaloa: Presidio near Mazatlan, San Blas; 
Jalisco: Autlan, Barranca de San Cristébal; Morelos: Cafion del 
Lobo near Cuernavaca; Oaxaca: Tapanatepec, Salina Cruz, Jimiltepec, 
Juchitan, Tehuantepec, Quiengola and Mixtequilla Mountains; 
Michoacén: Apatzingin, Parécuaro, Oropeo, Etticuaro, Puruaran; 
Colima: Paso del Rio, Colima; Guerrero: Huajintlan; Chiapas: Rancho 
San Bartolo. 


% A sight record for Zongolica, Veracruz (Martin del Campo, Anal. Inst. Biologfa, vol. 6, 1935, p. 297) 
is unacceptable and may be referable to Xenosaurus. Likewise unacceptable are records for Guanajuato 
and Yucatén. We are unable to find authoritative records for Nayarit, although Martin del Campo 
(loc. cit.) cites the state. 


194 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Family ANGUIDAE Cope 


Angulijdae Gray, Ann. Philos., vol. 26, 1825, p. 201. 
Anguidae Corn, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 228. 


Genera.—Eleven, of which nine are restricted to the Americas, 
one (Anguis) is restricted to the Old World, and one (Ophisaurus) 
occurs in both hemispheres. Five occur in Mexico.” 

Range.—Southern Canada to Argentina, West Indies, Europe, 
northern Africa, southern Asia. 


KEY TO MEXICAN GENERA OF ANGUIDAE 


No lateral goldhi ih et Ue ees A, ee Celestus (p. 194) 
A lateral ‘fold present. o-0U0 22 bees ba ST OOM arise Tee ee 2 

2. Lateral fold weakly developed, containing few or no granules; sides of neck 
with coarse granules; head much widened and depressed; pterygoid teeth 
absent; minimum scales in a transverse row across neck 4—6; arboreal 
SPECIES AELOe - Seeare (SAE We DAE LINER As ete Lee OR Ae oe O Abronia (p. 196) 
Lateral fold moderately to well developed, containing a moderate or large 
granular area; sides of neck with fine granules; head thicker, not depressed; 
pterygoid teeth various; minimum scales in a transverse row across neck 

no less, often more numerous; terrestrial species. ___.__.__._.---------- 3 

3. No anterior internasals, but instead a pair of enlarged supranasals, which 
much resemble internasals; no unpaired median internasals; nasal in contact 
with rostral; pterygoid teeth well developed_________----- Elgaria (p. 205) 
Anterior internasals, posterior internasals and supranasals ® all present; 
nasal seldom in contact with rostral; pterygoid teeth various_-__---_-- 4 

4. One or more median, unpaired postrostrals; frontals in contact with maxilla; 
pterygoid teeth well developed; tail extremely elongate_Gerrhonotus (p. 202) 

No postrostrals; frontals separated from maxilla; pterygoid teeth absent 
oF vestigial tail normal. 2008 bia Ae Bee. Brees Barisia (p. 198) 


Genus CELESTUS Gray 


Celestus Gray, Ann. Mag. Nat. Hist., vol. 2, 1839, p. 288.° 
Siderolamprus Corn, Proc. Acad. Nat. Sci. Philadelphia, 1860, p. 368 (type, S. 
enneagrammus Cope). 


Genotype.—Celestus striatus Gray. 
Range.—West Indies, Central America, and southern Mexico. 
Species.—About 18, two (possibly three) of which occur in Mexico. 


97 Ophisaurus ventralis, cited from Jalapa, Veracruz, as early as 1884 (Yarrow, U. S. Nat. Mus. Bull. 24, 
1884, p. 46), and retained in accounts of the species even up to the present time, has never been authoritatively 
recorded from Mexico and probably does not occur there. It almost certainly does not occur at Jalapa. 
It is possible that the record arose from a simple misunderstanding of Peale and Green’s Scincus ventralis 
(=Gerrhonotus liocephalus) which occurs in the same general area. Regardless of the nature of the error, 
we emphatically believe the record is erroneous. 

98 Supranasals absent in Gerrhonotus liocephalus austrinus, which has a postrostral. 

99 Dunn (Notulae Nat., No. 4, 1939, p. 3) expresses the opinion that forms here referred to Celestus, char- 
acterized by absence of sheaths on the claws, are congeneric with Diploglossus (Wiegmann, Herpetologia 
Mexicana, 1834, p. 36; type “‘Sc. fasciatus Gray’’= Diploglossus fasciatus), which possesses sheaths. 


CHECKLIST OF REPTILES OF MEXICO 195 


KEY TO MEXICAN SPECIES OF CELESTUS 


1. Median prefrontal in contact with 2 supraoculars, wider than long; no lateral 
prefrontals (fused with median loreal); 2 dorsolateral light stripes in adults 
separated by 4 and 2 half scale rows_-_------------- atitlanensis (p. 195) 

Median prefrontal in contact only with anterior supraocular, as long as wide; 
small lateral prefrontals usually present; adults not with 2 dorsolateral light 
Bimpess: 528 S22 3 OLE vO ie 8K asses SE ase. 2 

2. Snout normal, not especially elongate or flattened; lamellae under fourth toe, 
15-18; scales in 33-35 rows; first labial reaching to middle of naris; distance 
between anterior and posterior loreals less than the length of either; no ver- 
tical light bars on sides in young or adults_____--- enneagrammus (p. 195) 

Snout flattened, elongate; lamellae under fourth toe, 23-26; scales in 31-33 
rows; first labial reaching to anterior border of naris; distance between an- 
terior and posterior loreals as great or greater than length of either; vertical 
light bars present on sides in young and adults___-__----- rozellae (p. 195) 


CELESTUS ROZELLAE Smith 


Diploglossus steindachneri, GintTHER (nec Cope), Biologia Centrali-Americana, 
Reptilia and Batrachia, 1885, p. 34, pl. 22, fig. A. 
Celestus rozellae Smiru, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 372-374. 


Type.—U.S.N.M. No. 113526; Rozella Smith collector. 

Type locality—Palenque, Chiapas. 

Range.—Atlantic slopes, presumably from the Isthmus of Tehuan- 
tepec to British Honduras. Recorded in Mexico only from the type 


locality. 
CELESTUS ATITLANENSIS Smith, new species! 


Diploglossus (Celestus) stendachnert, Bocourt (nec Cope), Mission scientifique 
au Mexique . . . , Etudes sur les reptiles, livr. 6, 1879, pp. 383-384, pl. 22, 
fig. 3. 


Type.—Mus. Hist. Nat. Paris, specimen figured as above. 

Type locality —Atitlan, Guatemala. 

Range.—Known only from the type specimen from Atitlan, Guate- 
mala, but probably occurring along Pacific slopes from Chiapas to 


Nicaragua. 
CELESTUS ENNEAGRAMMUS (Cope) 


Siderolamprus enneagrammus Corn, Proc. Acad. Nat. Sci. Philadelphia, 1860, 
p. 368. 

Celestus enneagrammus, Corr, U. 8S. Nat. Mus. Bull. 32, 1887, p. 43; Ann. Rep. 
U.S. Nat. Mus., 1898 (1900), pp. 504-505.—Smiru, Proc. U. S. Nat. Mus., 
vol. 92, 1942, pp. 370-372, 374. 

Diploglossus steindachneri Corn, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 179 
(type locality, Orizaba, Veracruz; U.S.N.M. No. 6342).—BouLEnGER, Cata- 
logue of the lizards in the British Museum, vol. 2, 1885, p. 293. 


1 Diagnosis. — Allied to C. enneagrammus and C. rozellae, but with median prefrontal in contact with two 
(instead of one) supraoculars, no lateral prefrontals (presumably fused to median loreals), a very broad 
median prefrontal (broader than long), and a pattern in the single known adult (112 mm. snout to vent) 
consisting of two dorsolateral light stripes separating three broad dark stripes from each other. Similar to 
C. bivittatus Boulenger but with three loreals instead of two, seven supralabials to a point below middle of 
eye instead of six—H. M.S&. 


196 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Diploglossus chalybaeus Corn, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 321 
(type locality, Orizaba, Mexico, 4,000-6,000 feet; U.S.N.M. No. 6603). 


Type.—Mus. Comp. Zool. No. 2848; R. Montes de Oca collector. 

Type locality —Jalapa, Veracruz. 

Range.—Atlantic foothills of Veracruz and Oaxaca. Reported 
from Veracruz: Orizaba, Jalapa, Tequeyutepec, ‘‘one to three miles 
west of La Goya’; Oaxaca: Totontepec, “Tehuantepec,” Santa 


Efigenia. 
Genus ABRONIA Gray 


Abronia Gray, Ann. Mag. Nat. Hist., ser. 1, vol. 1, 1838, p. 3889.—TinEen, Amer. 
Midl. Nat., vol. 41, 1949, pp. 587-591. 


Genotype.—Gerrhonotus deppii Wiegmann [=Abronia deppii (Wieg- 
mann).]| 

Range.—Hidalgo, south to Guerrero and Chiapas. In Central 
America, Guatemala. 

Species.—Nine species, 10 forms; 6 species and 7 forms in Mexico. 


KEY TO MEXICAN FORMS OF ABRONIA 


1. Postmental unpaired: S22 VSS Owe iid OE A Le De Ee 2 

Postmental paired2_ 22.200) 2 eee) We tereern poh eek oben dea 3 

2. Seale rows 30-33; parietal separated from supraoculars or very narrowly in 

contact; dorsal markings usually indistinct, those on neck, if present, com- 

bining to form a large single mark_________-________ ochoterenai (p. 197) 

Scale rows 39; parietals broadly in contact with the supraocular; markings on 
back distinct, those on neck discrete, not forming a large single mark. 

matudai (p. 196) 

3. Suboculars missing or reduced to minute scales; one anterior temporal bordering 

OR DUbE = 2 Bake Se aes ou eRe eee ene ee A ee a ee deppii (p. 197) 

Suboculars present, well developed; 2 anterior temporals bordering orbit__ 4 

4. Area of granular scales on sides of neck very narrow; no granular zone in 

lateral fold..6 22.22.4222. 4... AR Be? OR eh oaxacae (p. 197) 

Area of granular scales covering entire sides of neck; a granular zone in lateral 

feolele 280) yee nee ¥en can bey sy Oh ml ee Noles Bee ee abt Ee ee 5 

5. Prominent light markings along the sides; dorsal osteoderms absent over 

posterior part of body; dorsal scales about 31-34 (average about 33); mini- 

mum number of scales in a single nuchal transverse row usually 6. 

taeniata taeniata (p. 197) 

No light markings on sides of body; dorsal osteoderms present over the whole 

bodys 240 ME Reinhart eb RT hel tS Le see nr PANN Ee Ah i tek GaN 6 

6. Dorsal scale rows 25 to 29; minimum number of scales in a transverse nuchal 

row, 4 or 5 (75 percent); chin and lower labials white with occasionally a few 

scattered dark blotchesss. 22.2222. -22b 2 22 taeniata graminea (p. 198) 

Dorsal scale rows 30-31; minimum number of scales in a transverse nuchal row, 

6; infralabials darker than chin and as dark as granular area of neck, with 

TN GUTS DENG Le Tae Tea 6 00 Gly eee eae fuscolabialis (p. 198) 


ABRONIA MATUDAI (Hartweg and Tihen) 
Gerrhonotus matudae Hartwec and T1HEN, Occ. Pap. Mus. Zool. Univ. Michigan, 


No. 497, 1946, pp. 3-5. 
Abronia matudai, Truen, Amer. Midl. Nat., vol. 41, 1949, p. 591. 


CHECKLIST OF REPTILES OF MEXICO 197 


Type.—Univ. Michigan Mus. Zool. No. 88331; Eizi Matuda 
collector. 

Type locality. —Volcan de Tacanaé, Chiapas, 2,000 meters. 

Range.—Known only from the type locality. 


ABRONIA OCHOTERENAI (Martin del Campo) 


Gerrhonotus vasconcelosti ochoterenati Martin DEL Campo, Anal. Inst. Biol. México, 
vol. 10, 1939, pp. 357-359, fig. 3 (cotypes). 

[Gerrhonotus] ochoterenai, SmitH, Proc. U. S. Nat. Mus., vol. 92, 1942, p. 368. 

Abronia ochoterenai, TrHEN, Amer. Midl. Nat., vol. 41, 1949, p. 591. 

?Gerrhonotus fimbriatus,2 Martin pet Campo, Anal. Inst. Biol. México, vol. 10, 
1939, p. 359. 


Type.—Instituto de Biologia, México, cotypes, male and female; 
Mario del Toro collector. 

Type locality.—Santa Rosa, Comit4n, Chiapas. 

Range.—Known only from the type locality. 


ABRONIA DEPPII (Wiegmann) 


Gerrhonotus deppii WIEGMANN, Isis von Oken, vol. 21, 1828, p. 379; Herpetologia 
Mexicana, pt. 1, 1834, pp. 31-32, pl. 9, figs. 3, 4. 

Abronia deppii, Gray, Ann. Nat. Hist., vol. 1, 1838, p. 389.—TinEen, Amer. 
Midl. Nat., vol. 41, 1949, p. 590. 

Gerrhonotus (Abronia) Deppeit, Bocourt, Mission Scientifique au Mexique... , 
Etudes sur les reptiles, livr. 5, 1878, pp. 325-327, pl. 214, figs. 3, 3a. 


Type.—Zool. Mus. Berlin; F. Deppe collector. 

Type locality —Mexico. Here restricted to Omilteme, Guerrero. 

Range.—Sierra Madre del Sur. Known only from the vicinity of 
Omilteme, Guerrero. 


ABRONIA OAXACAE (Giinther) 


Gerrhonotus oaxacae GitntTHER, Biologia Centrali-Americana, Reptilia and 
Batrachia, 1885, p. 36, pl. 24, figs. A, A’, A’’. 

Abronia oaxacae, T1HEN, Amer. Midl. Nat., vol. 41, 1949, p. 591. 

Type.—Brit. Mus. Nat. Hist., 2 adult males, 1 young; A. Boucard 
collector. 

Type locality —‘Oaxaca,’”’ Mexico. 

Range—Mountains in Oaxaca. Reported definitely only from 
“Luvina, Tehuantepec, Oaxaca.” 


ABRONIA TAENIATA TAENIATA (Wiegmann) 


Gerrhonotus taeniatus WIEGMANN, Isis von Oken, vol. 21, 1828, p. 379; Herpetologia 
Mexicana, 1834, p. 32, pl. 9, fig. 1, 2. 

Abronia taeniatus, Gray, Ann. Nat. Hist., vol. 1, 1838, p. 390.—Tisen, Amer. 
Midl. Nat., vol. 41, 1949, p. 590. 

Gerrhonotus (Abronia) taeniatus, Bocourt, Mission scientifique au Mexique . . ., 
Etudes sur les reptiles, livr. 5, 1878, pp. 8327-329, pl. 214A, figs. 4, 4a, and 5. 


2 This arrangement has been suggested verbally by Dr. Tihen, 


198 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Gerrhonotus deppit diguett Mocquarp, Bull. Mus. Hist. Nat. Paris, 1905, p. 79 
(Mus. Hist. Nat. Paris; Sierra de Zacapoaxtla, Puebla; Léon Diguet, collector). 
Type.—Zool. Mus. Berlin; F. Deppe collector. 
Type locality Mexico. Here restricted to El Chico, Hidalgo.* 
Range.—Southern Hidalgo, central and northern Puebla. Re- 
ported from Hidalgo: El Chico; Puebla: Zacapoaxtla, Ahuacatlan. 
Intergrades of taeniata and graminea are reported from La Joya, 


Veracruz. 
ABRONIA TAENIATA GRAMINEA (Cope) 


Gerrhonotus gramineus Corr, Proc. Acad. Nat. Sci. Philadelphia, Aug. 1864, p. 
179.—GintTHER, Biologia Centrali-Americana, Reptilia and Batrachia, 
1885, p. 36, pl. 24, fig. B. 

Abronia taeniata graminea, TrHEN, Amer. Midl. Nat., vol. 41, 1949, p. 591. 
Type.—U.S.N.M. (Field No. 50); Frangois E. Sumichrast collector. 
Type locality —Orizaba, Mexico. 

Range.—Central Veracruz and eastern Puebla. Reported from 

Puebla: P&ajaro Verde, Puente Colorado; Veracruz: Orizaba, 


Acultzingo. 
ABRONIA FUSCOLABIALIS (Tihen) 


Gerrhonotus fuscolabialis TrnEN, Copeia, 1944, pp. 112-115, figs. 1-3. 

Abronia fuscolabialis, TrneN, Amer. Midl. Nat., vol. 41, 1949, p. 591. 
Type.—R. T. Moore coll. No. 400 (California Inst. Tech.). 
Type locality—Mount Zempoaltepec, Oaxaca. 
Range.—Known only from the type locality. 


Genus BARISIA Gray 


Barisia Gray, Ann. Mag. Nat. Hist., ser. 1, vol. 1, 1838, p. 390.—Timmn, Amer. 
Midl. Nat., vol. 41, 1949, pp. 596-598, figs. 10-12; Univ. Kansas Sci. Bull., 
vol. 23, 1949, pp. 217-256, pls. 1-2, fig. 1. 

Mesaspis Corn, Proc. Amer. Philos. Soc., vol. 17, 1877, p. 96 (type, M. moreletii 
Bocourt). 


Genotype.—Gerrhonotus imbricatus Wiegmann [= Barisia imbricata 
imbricata (Wiegmann)]. 

Range.—Chibuahua and Coahuila south to Chiapas; Central 
America to Panama. 

Species. —Nine, with 16 forms; 8 species, 12 forms, occur in Mexico. 


KEY TO MEXICAN SPECIES OF BARISIA 


1. Postmental unpaired; prefrontal bones in contact______-_..------------ 2 
Postmental paired; prefrontal bones separated by frontonasal______-_-- 4 
2. Eighteen to 20 longitudinal dorsal scale rows; frontonasal normally present. 3 
Fourteen longitudinal dorsal scale rows; frontonasal normally absent. 
viridiflava (p. 200) 
3. Upper postnasal separated from lower by a loreonasal contact. 
moreleti rafaeli (p. 199) 


3 It is known that Deppe collected in this region. 


CHECKLIST OF REPTILES OF MEXICO 199 


Upper and lower postnasals in contact_______- moreleti temporalis (p. 199) 
4. Superciliary series complete; pterygoid teeth absent______.____________ 5 
Superciliary series incomplete; pterygoid teeth vestigial________________ 8 
5. Supranasals unexpanded; an anterior canthal present_________________-_ 6 
Supranasals expanded’ no anterior canthal 222 8a. a ee ee 7 


6. Scales of neck keeled; anterior loreal in contact with anterior canthal. 
gadovii gadovii (p. 200) 
Scales of neck smooth; anterior loreal separated from anterior canthal by a 
contact of posterior loreal with upper postnasal__gadovii levigata (p. 200) 


7. Frontonasal and postrostral present_.._._.__._.____________- modesta (p. 201) 
No frontonasal or postrostral present___._.-..._.__..------ antauges (p. 200) 

8. One loreal; more than 33 dorsal scale’ rows2__l-J.__........_..._______- 9 
Two loreals; fewer than 33 dorsal scale rows____-__------ rudicollis (p. 202) 

9. More than 1 superciliary (usually 3); transverse dorsal scale rows usually 
fewer (ham, 470 5-2 ee a ae fea ee SS i aes 2 oe pa 10 

A single (middle) superciliary element; transverse dorsal scale rows usually 
TNIOLO UAE a om eae ag ee te a st ae eae ae eke cape levicollis (p. 202) 


10. Fewer than 16 longitudinal dorsal scale rows; contact of anterior superciliary 
with loreal usually smaller than that of first medial supraocular with 
florea tt’ hater es = sag ES Beige | Sok. pala ee ie EM ic arnt peal 11 

Sixteen longitudinal dorsal scale rows; contact of anterior superciliary with 
loreal as great or greater than that of first medial supraocular with loreal. 
imbricata imbricata (p. 201) 

11. Lowest primary temporal in contact with penultimate as well as with ante- 

penultimate supralabial; 39-45 transverse dorsal scale rows. 
imbricata ciliaris (p. 202) 
Lowest primary temporal not in contact with penultimate supralabial; 35-39 
transverse dorsal scale rows._...--------- imbricata planifrons (p. 201) 


BARISIA MORELETI RAFAELI (Hartweg and Tihen) 


Gerrhonotus moreleti rafaeli Hartwre and Tien, Occ. Pap. Mus. Zool. Univ. 
Michigan, No. 497, 1946, pp. 8-10. 

Barisia moreleti rafaeli, TramN, Amer. Midl. Nat., vol. 41, 1949, pp. 223-224. 

Type.—Mus. Zool. Univ. Michigan No. 88228; Eizi Matuda 
collector. 

Type locality.—16 km. south of Siltepec, Chiapas, 2,300 meters 
elevation. 

Range.—High mountains of southern Chiapas. Reported from 
Cerro Paxtal, 1,500 meters; Chiquihuite, Volcan de Tacan4, 2,500 
meters; Cerro Malé. 


BARISIA MORELETI TEMPORALIS (Hartweg and Tihen) 


Gerrhonotus morelett temporalis HartwrG and TisEen, Occ. Pap. Mus. Zool. 
Univ. Michigan, No. 497, 1946, pp. 10-15. 
Barisia moreleti temporalis, TrupN, Amer. Midl. Nat., vol. 41, 1949, pp. 224-225. 
Type.—Mus. Zool. Univ. Michigan No. 94910; Norman Hartweg 
collector. 
Type locality.—Eleven km. southeast of Ciudad de las Casas, 
Chiapas, 2,300 meters elevation. 
Range.—Known only from the type locality. 


200 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 
BARISIA GADOVII GADOVII (Boulenger) 


Gerrhonotus gadoviti BouULENGER, Ann. Mag. Nat. Hist., ser. 8, vol. 12, 1913, pp. 
564-565. 
Barisia gadovii gadovii, THEN, Univ. Kansas Sci. Bull., vol. 33, 1949, pp. 230-231, 


Type.—Brit. Mus. Nat. Hist., two cotypes; Hans Gadow collector. 

Type locality —Omilteme, Guerrero. 

Range.—Sierra Madre del Sur, in Guerrero. Known only from 
mountains west of Chilpancingo and the type locality. 


BARISIA GADOVII LEVIGATA Tihen 


Barisia gadovii levigata TrHEN, Univ. Kansas Sci. Bull., vol. 33, 1949, pp. 231-233, 
pl. 2, figs. 3-5. 


Type.—U.S.N.M. No. 47212; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality —‘ Valley of Oaxaca,’’ Oaxaca. 

Range.—Mountains of central Oaxaca. Reported from ‘‘moun- 
tains west of Oaxaca city.” 


BARISIA VIRIDIFLAVA (Bocourt) 


Gerrhonotus viridiflavus Bocourt, Ann. Sci. Nat., vol. 17, 1873, art. 2 (unpaged). 

Barisia viridiflava, Trsen, Amer. Midl. Nat., vol. 41, 1949, pp. 227-229, fig. 1. 

Gerrhonotus bocourti Peters, Monatsb. Acad. Wiss. Berlin, 1876, p. 297 (type 
locality, ‘‘Mexico,”’ here restricted to Oaxaca, Oaxaca; Zool. Mus. Berl.). 

Gerrhonotus antauges (part), Bocourt, Mission scientifique au Mexique.. ., 
Etudes sur les reptiles, livr. 5, 1878, pp. 346-349, pl. 21B, figs. 7, 7a. 

Gerrhonotus obscurus GUnruer, Biologia Centrali-Americana, Reptilia and 
Batrachia, 1895, p. 40, pl. 25, figs. E, E’ (type locality, ‘‘Mexico,’”’ here 
restricted to Oaxaca, Oaxaca; Brit. Mus. Nat. Hist.). 


Type.—Mus. Hist. Nat. Paris; ‘‘un seul exemplaire qui a été cédé 
par M. Boucard, comme provenant des collections de M. F. Sumi- 
chrast.”’ 

Type locality—Mexico. Restricted by Tihen to the highlands of 
central Oaxaca, near the city of Oaxaca. 

Range.—Mountains north of Oaxaca (city). Reported from ‘‘sum- 
mit of Cerro San Felipe north of Oaxaca”’ (city). 


BARISIA ANTAUGES Cope 


Barissia antauges Corr, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 132. 
Barisia antauges, TinEN, Amer. Midl. Nat., vol. 41, 1949, pp. 235-236. 


Type.-—U.5S. N. M. No. 30221; Francois E. Sumichrast collector. 

Type locality —Orizaba, Veracruz [= Volcan Citlaltépetl]. 

Range.—Volcan Citlaltépetl (Mount Orizaba), Veracruz; known 
only from the type locality. 





CHECKLIST OF REPTILES OF MEXICO 201 
BARISIA MODESTA (Cope) 


Pterogasterus modestus Corr, Proc. Amer. Philos. Soc., 1877, pp. 96, 97. 
Gerrhonotus modestus, BOULENGER, Catalogue of the lizards in the British Museum, 
vol. 2, 1885, p. 276. 
Barisia modestus, TinEN, Amer. Midl. Nat., vol. 41, 1949, pp. 284-235. 
Type.—vU.S. Nat. Mus., three specimens; collector unknown. 
Type locality — ‘Probably Guatemaia,’”’ here restricted to Mount 


Orizaba. 
Range.—Region of Mount Orizaba (Volc4n Citlaltépetl), Veracruz 


(not known from Guatemala). 
BARISIA IMBRICATA IMBRICATA (Wiegmann) 


Gerrhonotus imbricatus WIEGMANN, Isis von Oken, vol. 21, 1828, p. 379; Herpetologia 
Mexicana, 1834, pt. 1, p. 34, pl. 10, figs. 2, 5. 

Barissia imbricata, Gray, Catalogue of the lizards in the British Museum, 1845, 
p. 55. 

Gerrhonotus (Barissia) imbricatus, Bocourt, Mission scientifique au Mexique . . ., 
Etudes sur les reptiles, livr. 5, 1878, pl. 21B, figs. 1, 1a, 2, 2a, and livr. 6, 
1879, pp. 363-365. 

Barisia imbricata imbricata, TrHEN, Amer. Midl. Nat., vol. 41, 1949, pp. 240-244. 

Gerrhonotus lichenigerus WaGuER, Descriptiones et icones amphibiorum, 1833, 
pl. 34, fig. 2 (type locality here restricted to Mexico, D. F.).—Wiramann, 
Herpetologia Mexicana, 1834, pl. 10, figs. 2-5. 

Gerrhonotus adspersus W1EGMANN, Herpetologia Mexicana, 1834, pl. 10 (type 
locality, Mexico by inference, here restricted to San Martin, México; Zool. 


Mus. Berl.; F. Deppe collector). 
Gerrhonotus olivaceus Barrp, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 255 
(type locality, “near San Diego;” U.S.N.M. No. 3096, two cotypes). 


Type.—Zool. Mus. Berlin, two cotypes; F. Deppe collector. 

Type locality—Mexico, here restricted to México, D. F. 

Range.—The periphery and southern part of the central Mexican 
Plateau, from Veracruz to Jalisco. Reported from numerous localities 
in the states of Jalisco, Michoacan, México, Morelos, Puebla, Oaxaca, 
Veracruz, Hidalgo, and Guanajuato, and from Distrito Federal. 


BARISIA IMBRICATA PLANIFRONS (Bocourt) 


Gerrhonotus (Barissia) planifrons Bocourt, Mission scientifique au Mexique . . ., 
Etudes sur les reptiles, livr. 5, 1878, pl. 21C, figs. 1, 1a; livr. 6, 1879, pp. 
361-363 (pl. 21C was published with the name, in livr. 5, 1878; this con- 
stitutes the “type description’’). 


Barisia imbricata planifrons, TinEN, Amer. Midl. Nat., vol. 41, 1949, pp. 246-247, 
Type.—Mus. Hist. Nat. Paris; Adolphe Boucard collector. 


Type locality.— “Oaxaca.” 
Range.—Oaxaca, mountainous areas; not known from a definite 


locality. 


202 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


BARISIA IMBRICATA CILIARIS (Smith) 


Gerrhonotus levicollis ciliaris SmirH, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 
365-367. 
Barisia imbricata ciliaris, TraEN, Amer. Midl. Nat., vol. 41, 1949, pp. 244-245. 

Type.—U.S.N.M. No. 47496; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality.—Sierra Guadelupe, Coahuila. 

Range.—The northern portion of the plateau from extreme southern 
Coahuila to Guanajuato, east to San Luis Potosi and west to Sinaloa. 
Reported from Coahuila: Sierra Guadelupe; Zacatecas: Sierra Madre; 
Durango: Coyotes, Inde; San Luis Potosi: mountains near San Luis 
Potosi (city), near Xilitla, near Jestis Maria; Guanajuato: San Felipe, 
Sierra de Santa Rosa; Hidalgo: Durango; Nuevo Ledn: Pablillo, 
Galeana; Sinaloa: Escuinapa. 


BARISIA LEVICOLLIS Stejneger 


Barissia levicollis SrrJNEGER, Proc. U.S. Nat. Mus., vol. 13, 1890, pp. 184-185. — 
Corz, Ann. Rep. U. 8. Nat. Mus., 1898 (1900), pp. 535-536, fig. 96. 

[Gerrhonotus] levicollis levicollis, Smitu, Proc. U. 8, Nat. Mus., vol. 92, 1942, p. 368. 

Barisia levicollis, Traen, Amer. Midl. Nat., vol. 41, 1949, pp. 247-248; Univ. 
Kansas Sci. Bull., vol. 33, 1949, pp. 247-248. 


Type.-—U.S.N.M. No. 9362; “evidently obtained by one of the 
surveying parties of the United States and Mexican Boundary 
Survey,” fide Stejneger. 

Type locality —‘‘Mexican boundary.’’ Probably Chihuahua, fide 
Tihen, loc. cit. 

Range.—Chihuahua and probably adjacent areas in Sonora. 
Reported from Chihuahua: north Chihuahua, Colonia Garcia, Meadow 
Valley, 65 miles east of Batopilas, Samachique. 


BARISIA RUDICOLLIS (Wiegmann) 


Gerrhonotus rudicollis W1EGMANN, Isis von Oken, vol. 21, 1828, p. 379; Herpeto- 
logia Mexicana, 1834, pp. 33-34, pl. 10, figs. 1, 4—Martfin peL Campo, 
Anal. Inst. Biol., vol. 10, No. 3-4, 1939, p. 360, fig. 4. 

Barissia rudicollis, Gray, Catalogue of the lizards in the collection of the British 
Museum, London, 1845, p. 55. 


Gerrhonolus (Barissia) rudicollis, Bocourt, Mission scientifique au Mexique..., 
Etudes sur les reptiles, livr. 5, 1878, pl. 21B, figs. 3, 3a; and livr. 6, 1879, 
pp. 367-369. 


Barisia rudicollis, Tiu=N, Amer. Midl. Nat., vol. 41, 1949, pp. 238-240, pl. 1. 


Type.—Zool. Mus. Berlin; F. Deppe collector. 

Type locality.—Mexico. Here restricted to Hacienda de le Gavia, 
México (state). 

Range.—Western México and perhaps adjoining parts of Mich- 
oacan. Reported from Mézico: Hacienda de la Gavia. 


CHECKLIST OF REPTILES OF MEXICO 203 


Genus GERRHONOTUS Wiegmann 


Gerrhonotus W1EGMANN, Isis von Oken, vol. 21, 1828, No. 3, p. 379.—TinEn, 
Amer. Midl. Nat., vol. 41, 1949, pp. 591-593. 

Pterogasienes PEALE and GREEN, Journ. Acad. Nat. Sci. Philadelphia, vol. 6, 
1830, p. 234 (type, Scincus ventralis Peale and Green). 


Genotype.—Gerrhonotus tessellatus Wiegmann [=Gerrhonotus lioce- 
phalus Wiegmann]. 

Range.—Central Texas southward through the plateaux of Mexico 
to southwestern Chiapas. 

Species.—One. Five forms recognized, all occurring in Mexico. 


KEY TO SPECIES OF GERRHONOTUS 


iee-khree loreals) (oreocanthals)) on eachisides- 25 ==. 2) Se 2 
Four’or more loreocanthals on each side! 2.200 ee SS nS 3 
2. Supranasals absent; azygous prefrontal much longer than broad; frontal 
widely separated from interparietal; all except lowermost anterior temporal 
touch fifth medial supraocular___._____.____ liocephalus austrinus (p. 204) 
Supranasals present; azygous prefrontal as broad as or broader than long; 
frontal in contact with or very narrowly separated from interparietal; only 

2 uppermost anterior temporals in contact with fifth supraocular. 
liocephalus liocephalus (p. 203) 
3. Dorsal bands obsolete; venter nearly without marking; 52-60 dorsal scales 
(average 55. 6); tail to body ratio, 2. 5-2. 6; caudals 157-163; second primary 

temporal touches the fifth medial supraocular. 

liocephalus loweryi (p. 204) 
Dorsal bands distinct; venter mottled and flecked______________________ 4 
4. Dorsal scales 45-54 (average 49); tail-body ratio, 1. 75-2. 1; caudals, 116-137. 
liocephalus infernalis (p. 204) 
Dorsal scales 49-52 (average 51); tail-body ratio 2. 3; caudals (approx.) 140. 
liocephalus ophiurus (p. 204) 


GERRHONOTUS LIOCEPHALUS LIOCEPHALUS Wiegmann 


G[errhonotus] liocephalus W1EGMANN, Isis von Oken, vol. 21, 1828, p. 381. 

Gerrhonotus liocephalus, Bocourt, Mission scientifique au Mexique ..., Etudes 
sur les reptiles, livr. 5, 1878, pp. 342-346, pl. 214A, figs. 1, 2, 2a. 

Gerrhonotus liocephalus liocephalus, Corr, Ann. Rep. U. S. Nat. Mus. 1898 (1900), 
pp. 516-517.—Smirtu, Proc. U. 8. Nat. Mus., vol. 92, 1942, p. 369. 

Glerrhonotus] tessellatus WIEGMANN, Herpetologia Mexicana, pt. 1, 1834, pp. 
32-33, pl. 10, fig. 3 (substitute name for preceding). 

Scincus ventralis PEALE and GREEN, Journ. Acad. Nat. Sci. Philadelphia, 1838, 
pt. 4, p. 233 (type locality, ‘‘Mining Districts of Mexico’ = ‘‘Valley of Mexico”’ 
fide Cope, here restricted to Magdalena, D. F.; type in Acad. Nat. Sci. 
Philadelphia, two cotypes). 


Type.—Zool. Mus. Berlin; F. Deppe collector. 

Type locality. —Mexico (Oaxaca, fide Bocourt loc. cit., p. 344, here 
restricted to Tlapancingo). 

Range.—Central plateau region, from Guanajuato to Guerrero and 
Oaxaca. Reported from Distrito Federal: Magdalena, Mixiuhca; 
Guerrero: Omilteme; Mézico: Temascaltepec; Oazaca: Llano Ocotal, 

861316—50——14 


204 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


La Concepcién, Tlapancingo, Tres Cruces; Puebla: Cacaloapam; 
Morelos: Cuernavaca; Guanajuato: Silao. 


GERRHONOTUS LIOCEPHALUS LOWERYI Tihen 


Gerrhonotus liocephalus loweryi T1rnEN, Trans. Kansas Acad. Sci., vol. 51, 1948, 
pp. 302-305. 
Type.—La. State Univ. Mus. No. 480; Marcella Newman collector. 
Type locality.—‘‘Xilitla region” (neighborhood of Xilitla, San Luis 
Potosi). 
Range.—Known only from the type locality and possibly also from 
Ciudad del Maiz, San Luis Potosi. 


GERRHONOTUS LIOCEPHALUS OPHIURUS Cope 


Gerrhonotus ophiurus Coreg, Proc. Acad. Nat. Sci. Philadelphia, 1866, pp. 321-322. 
Pterogasterus ophiurus, Corr, Proc. Amer. Philos. Soc., vol. 17, 1877, p. 96. 
Gerrhonotus liocephalus ophiurus, Corr, Ann. Rep. U. S. Nat. Mus., 1898 (1900), 
pp. 516-517. 
Gerrhonotus lemniscatus Bocourt, Nouv. Arch. Mus. Hist. Nat. Paris, vol. 7, 
1872, pp. 105-107 (Mus. Hist. Nat. Paris; Veracruz). 
Type.—U.S.N.M. No. 30206; Francois E. Sumichrast collector. 
Type locality.—Orizaba, Veracruz, Mexico. 
Range.—Eastern foothills of the plateau. Known only from Vera- 
cruz: Orizaba, Jicaltepec, Cérdoba, Cerro Gordo; Puebla: Hueyta- 
malco (Teziutlan). 


GERRHONOTUS LIOCEPHALUS INFERNALIS Baird 


Gerrhonotus infernalis Barrp, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 255. 

Gerrhonotus liocephalus infernalis, Corn, Ann. Rept. U. S. Nat. Mus., 1898 (1900), 
pp. 517-519, fig. 9.—Smiru, Handbook of lizards, 1946, pp. 463-464, pl. 
131.—TineEn, Amer. Midl. Nat., vol. 41, 1949, p. 593. 

Type.—? U.S.N.M. No. 3090; C. B. R. Kennerly collector. 

Type locality —Devils River, Tex. 

Range.—San Luis Potosi, Coahuila and probably adjoining states 
of Chihuahua and Nuevo Leon, northward to central Texas across 
the Rio Grande valley from Rio Grande City to mouth of Devil’s 
River. Reported from Coahuila: Carmen Mountains, Cerro Encarna- 
cién, Sierra Guadelupe south of La Cuchilla, Monclova; San Luis 
Potosi: Alvarez. 


GERRHONOTUS LIOCEPHALUS AUSTRINUS Hartweg and Tihen 
Gerrhonotus liocephalus austrinus Harrwea and TinEn, Occ. Pap. Mus. Zool. 
Univ. Michigan, No. 497, 1946, pp. 6, 7. 


Type.—Univ. Michigan Mus. Zool. No. 94921; Eizi Matuda collector. 

Type locality—3,200 meters elevation on Cerro Malé, Porvenir, 
Chiapas. 

Range.—Known only from the type locality. 


CHECKLIST OF REPTILES OF MEXICO 205 


Genus ELGARIA Gray 


Elgaria Gray, Ann. Mag. Nat. Hist., ser. 1, vol. 1, 1838, p. 390.—Tinen, Amer. 
Midl. Nat., vol. 41, 1949, pp. 593-596, figs. 8-9. 

Tropidolepis SKILTON (nec Cuvier), Amer. Journ. Sci., ser. 2, vol. 7, 1849, p. 202 
(type, T. scincicauda Skilton). 


Genotype.—Cordylus (Gerrhonotus) multicarinatus Blainville= Elgaria 
multicarinata multicarinata. 

Range—Chibuahua and Baja California, northward through 
California to British Columbia, Utah, and Montana. 

Species.—Six species and 13 forms. Six forms occur in Mexco. 


KEY TO MEXICAN SPECIES OF ELGARIA 


1. Alternate black and white marks on Jabials (white spots somewhat ocellate). 
kingii group 4 
No alternate black and white marks on labials__-_---_ multicarinata group 2 
2. \Pemporalisealestall’or partly. keeled2o3— Se. Lb? _ ooh Or Sek AES a Ot 3 
Temporal scales smooth; granular area of lateral fold white with gray or black 
reticulations; scales on arm smooth; 2 rows of scales keeled on thigh; lateral 
body scalesismooth=2{= 22 2. t.2-2 28) See ere LEE cedrosensis (p. 206) 
3. Large (180 mm. maximum); dorsals all heavily keeled; two upper temporal 
rows sharply keeled, others weakly keeled or nearly smooth; three rows of 
keeled scales on arm; dark transverse bands indented with white markings 
5 scale rows above lateral fold and on back; ventral longitudinal lines faint 
OF ODSOletete. = eee ee eo eee multicarinata webbii (p. 207) 
Smaller (114 mm. maximum); dorsals less strongly keeled; two upper temp- 
orals strongly keeled; 1 to 3 keeled rows on arm; dark ventral longitudinal 

lines; wide brown band on temporal region from eye to ear. 

multicarinata nana (p. 207) 
4, Granular area of lateral fold white crossed by black bands usually continuous 
with dark bodv bands; eight dorsal scale rows keeled moderately, lateral rows 
faintly keeled; 4 internasals of about same size; arm scales smooth; thigh 
Seales; withs2irows keeledsst eases oe ee ee paucicarinatus (p. 206) 
Granular area of lateral fold dull gray; heavy broad dorsal bands on neck, body, 
tail, the latter almost encircling tail; usually 6 or 8 weakly keeled dorsal 
scale rows; scales on arm and thigh smooth._.........-....-.------- 5 
5. Dorsal longitudinal scale rows usually 16; scales from occiput to posterior 
part of thigh in 51 to 56 transverse rows (average, 52.9); transverse row of 
four anterior temporals, uppermost always in contact with uppermost 
secondary temporal; granular area in fold usually black_ kingii kingii (p. 205) 
Dorsal longitudinal scale rows usually 14; scales from occiput to posterior part 
of thigh 55 to 60 (average 57.3); four anterior temporals, uppermost sep- 
arated from uppermost secondary temporal (60 percent); granular area in 
lel agama perigee Ge etc aks SE kingii nobilis (p. 206) 


ELGARIA KINGII KINGII Gray 


Elgaria kingiit Gray, Ann. Mag. Nat. Hist., vol. 1, 1838, p. 390. 

Elgaria kingii kingii, Trmen, Trans. Kansas Acad. Sci., vol. 51, 1948, pp. 299-300, 

Gerrhonotus kingii, Bocourt, Nouv. Arch. Mus. Hist. Nat. Paris, vol. 7, p. 106; 
Mission scientifique au Mexique . . ., Etudes sur les reptiles, livr. 5, 1878, 
pp. 339-342, pl. 21C, figs. 2, 2a (name attributed to Bell, his manuscript 
published by Gray). 


206 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Gerrhonotus multifasciatus DumMérit and Bocourt, Erpétologie générale, vol. 5, 
1839, p. 401 (type locality, “‘Mexico’’, here restricted to Mojirachic, Chi- 
huahua; Mus. Hist. Nat. Paris). 

Type.—Brit. Mus. Nat. Hist., adult (in bad state); from the T. 
Bell collection, collector unknown. 

Type locality—Mexico [so stated by Boulenger, Catalogue of 
Lizards ..., vol. 2, 1885, p. 275]. Tihen suggests ‘presumably 
Chihuahua.”’ Here restricted to Mojarachic. 

Range.—Eastern Sonora and western Chihuahua, probably south- 
ward into Durango and Sinaloa. Reported from Chihuahua: Madera, 
Mojarachic, Chiricahui, Colonia Garcia (possibly intergrades with 
with nobilis); Sonora: (no specific locality). 


ELGARIA KINGIT NOBILIS Baird and Girard 


Elgaria nobilis Batrp and Grrarp, Proc. Acad. Nat. Sci. Philadelphia, vol. 6, 
1852, p. 129. 

Gerrhonotus nobilis, Batrp, United States and Mexican boundary survey, vol. 2, 
Reptiles, 1859, p. 11, pl. 25, fig. 108. 

Elgaria kingit nobilis, Truen, Amer. Midl. Nat., vol. 41, 1949, p. 596. 

Gerrhonotus kingit, Smita, Handbook of lizards, 1946, pp. 452-454, pl. 127. 

Type.—Possibly in Acad. Nat. Sci. Philadelphia. 

Type localityx—Fort Webster Copper Mines of the Gila [Santa 
Rita del Cobre], N. Mex. 

Range.—Southwestern New Mexico and southeastern Arizona. 
While no records are available from Mexico, specimens have been 
taken within a few hundred yards of the border, and the form must. 
occur in Mexico (Sonora). 


ELGARIA PAUCICARINATUS (Fitch) 


Gerrhonoltus paucicarinatus Fircu, Copeia, 1934, No. 4, 1935, pp. 172-173. 
Elgaria paucicarinata, TirHEN, Amer. Midl. Nat., vol. 41, 1949, p. 595. 

Type—Mus. Vert. Zool. Univ. California No. 11768, male; C. C. 
Lamb collector. 

Type locality —Todos Santos, Baja California. 

Range.—Southern end of Baja California in the vicinity of the type 


locality. 
ELGARIA CEDROSENSIS (Fitch) 


Gerrhonotus cedrosensis Firc#, Copeia, 1934, No. 1, pp. 6-7. 
Elgaria cedrosensis, TrHeN, Amer. Midl. Nat., vol. 41, 1949, p. 595. 


Type.—California Acad. Sci. No. 56187; J. R. Slevin collector. 
Type locality—‘Cafion on southeast side of Cedros Island,” Baja. 


California. 
Range.—Cedros Island, Baja California. 


CHECKLIST OF REPTILES OF MEXICO 207 


ELGARIA MULTICARINATA NANA (Fitch) 


Gerrhonotus scincicauda nanus Fircu, Copeia, 1934, p. 7. 
Gerrhonotus multi-carinatus nanus, Fircu, Amer. Midl. Nat., vol. 20, 1938, pp. 
397-399, fig. 2. 
Elgaria multicarinata nana, TrHEN, Amer. Midl. Nat., vol. 41, 1949, p. 596. 
Type.—Mus. Vert. Zool. Univ. California No. 5402; A. B. Howell 
collector. 
Type locality—South Island, Los Coronados Islands, Baja 
California. 
Range.—Los Coronados Islands, Baja California. 


ELGARIA MULTICARINATA WEBBII (Baird) 


Gerrhonotus webbit Barrp, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 225. 

Gerrhonotus scincicauda webbit, GRINNELL and Camp, Univ. California Publ. Zool., 
vol. 17, 1917, p. 168. 

Gerrhonotus multi-carinatus webbii, Fircu, Amer. Midl. Nat., vol. 20, 1938, pp. 
395-397.—Smiru, Handbook of lizards, 1946, pp. 460-462, pl. 130. 

Elgaria multicarinatus webbii, Tru=N, Amer. Midl. Nat., vol. 41, 1949, p. 596. 

Gerrhonotus scincicauda ignavus VAN DENBURGH, Proc. California Acad. Sci., ser. 
3, vol. 4, 1905, p. 19, pl. 7, figs. 1-2 (type locality, San Martin Island, Baja 
California). 

Type. —U.S.N.M. No. 3078; T. H. Webb collector. 

Type locality—‘“‘San Diego to El Paso,” ‘probably from near 
vicinity of San Diego, San Diego Co. California [?]’’ (Fitch, loc. cit.). 
Here restricted to San Diego, Calif. 

Range.—Southern California and extreme northern Baja California. 
Reported in Baja California from San Pedro MaArtir Mountains, 


Alcatraz. 
Family XENOSAURIDAE Cope 


Xenosauridae Core, Proc. Acad. Nat. Sci. Philadelphia, vol. 18, 1866 (1867), 
p. 322. 
Genera.—A single genus, Xenosaurus Peters, is known. 
Range.—San Luis Potosi to Guatemala. 


Genus XENOSAURUS Peters 
Xenosaurus Peters, Monatsb. Acad. Wiss. Berlin, 1861, p. 453. 


Genotype.—Xenosaurus fasciatus Peters=Xenosaurus grandis 
(Gray). 
Range.——San Luis Potosi to Guatemala. 


Species.— Three. 
KEY TO SPECIES OF XENOSAURUS 


1. Arm with widely spaced tubercles on its dorsal surface; a row of supraoculars 
each 1% to 2 times as long as wide; cream, black spotted on venter____- 2 
Arm regularly covered with tubercles on dorsal surface; supraoculars not 
forming a series of more or less regular enlarged scales; venter uniformly gray 

or gray-white without spots or flecks_____._________ newmanorum (p. 208) 


208 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


2. Scales bordering posterior gular fold small, rather widely separated from each 
other, considerably smaller than median chest scales______ grandis (p. 208) 
Scales bordering posterior gular fold larger, in contact with each other or nar- 
rowly separated, about as large as median chest scales._rackhami (p. 208) 


XENOSAURUS NEWMANORUM Taylor 


Xenosaurus newmanorum Taytor, Univ. Kansas Sci. Bull., vol. 33, 1949, pp. 
183-187. 
Type.—Louisiana State Univ. No. 85; Marcella Newman collector. 
Type locality —‘‘Xilitla Region’”’ (near Xilitla, San Luis Potosf). 
Range.—Known only from the type locality. 


XENOSAURUS GRANDIS (Gray) 


Cubina grandis Gray, Ann. Mag. Nat. Hist., ser. 2, vol. 18, 1856, p. 270. 
Xenosaurus grandis, Corr, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 322.— 
Bocourr, Mission scientifique au Mexique . . . , Etudes sur les reptiles, 
livr. 5, 1878, pp. 303-306, pl. 20F, figs. la-d, 1f, 1g—Cope, Ann. Rep. U.S. 
Nat. Mus., 1898 (1900), pp. 538-539.—Barrows and Smiru, Univ. Kansas 
Sci. Bull., vol. 31, 1947, pp. 227-281, pls. 11-16. 
Xenosaurus fasciatus Peters, Monatsb. Acad. Wiss. Berlin, 1861, p. 453 (type 
locality, Huanusco [=Huatusco], Veracruz; Berl. Mus.). 
Type.—Brit. Mus. Nat. Hist.; A. Sallé collector. 
Type locality —Cérdoba, Veracruz (probably from nearby moun- 
tains). 
Range.—Foothills of central Veracruz southward to the Isthmus. 
Reported from Veracruz: Huanusco [=Huatusco], Cérdoba, Orizaba; 
Oaxaca: “Oaxaca,” Tehuantepec.‘ 


XENOSAURUS RACKHAMI Stuart 


Xenosaurus rackhami Stuart, Proc. Biol. Soe. Washington, vol. 54, 1941, pp. 
47-48.—SmirTH, Journ. Washington Acad. Sci., vol. 39, 1949, p. 43. 

Type.—Mus. Zool. Univ. Michigan No. 89072. 

Type locality —Finca Volcan, 49 km. east of Coban, Alta Verapaz, 
Guatemala, 4,000 feet. 

Range.—The highlands of central eastern Chiapas and adjacent 
Guatemala. Recorded in Mexico only from Chiapas: Santa Rosa 
(near Comit4n). 


Family ANNIELLIDAE Boulenger 


Aniellidae Corn, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 230. 
Anniellidae BoutenarEr, Catalogue of the lizards in the British Museum, vol. 2, 
1885, p. 299. 


Genera.—Only a single genus, Anniella Gray, is known. 
Range.—Southwestern California and northwestern Baja California. 
4 Fide Cope, U. 8. Nat. Mus. Bull. 32, 1887, p. 39. A sight record cited by Martin del Campo, Folleto 


Div. Cient. Inst. Biol., No. 2, 1934, p. 7, of ‘‘Heloderma horridum”’ from Sierra de Zangolica, Veracruz, may 
belong with this species; it is very unlikely that Heloderma actually occurs on the Atlantic coast. 


CHECKLIST OF REPTILES OF MEXICO 209 


Genus ANNIELLA Gray 


Anniella Gray, Ann. Mag. Nat. Hist., ser. 2, vol. 10, 1852, p. 440. 


Genotype-—Anmiella pulchra Gray. 
Range.—Southern California and northern Baja California. 
Species.—Two, one with two subspecies. 


KEY TO MEXICAN SPECIES OF ANNIELLA 


1. Rostral sharply pointed in profile; fourth supralabial largest. 
geronimensis (p. 209) 
Rostral rounded in profile; second supralabial largest. 
pulchra pulchra (p. 209) 


ANNIELLA GERONIMENSIS Shaw 


Anniella geronimensis SHaw, Trans. San Diego Soc. Nat. Hist., vol. 9, 1940, 
pp. 225-228, 2 figs. 
Type.—lL. M. Klauber No. 7543. 
Type locality.—Isla San Gerénimo, Baja California. 
Range.—Known only from the type locality. 


ANNIELLA PULCHRA PULCHRA Gray 


Anniella pulchra Gray, Ann. Mag. Nat. Hist., ser. 2, vol. 10, 1852, p. 440.— 

Van Densoregs, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 465-467, 
1. 42. 

dala pulchra pulchra, GRINNELL and Camp, Univ. California Publ. Zool., 
vol. 17, 1917, p. 170.—MI.LuER, Copeia, 1943, p. 2. 

Anniella tecana BouLENGER, Ann. Mag. Nat. Hist., ser. 5, vol. 20, 1887, p. 50 
(Brit. Mus. Nat. Hist.; El Paso, Tex., in error, here restricted to San Diego, 
California). 

Type— Brit. Mus. Nat. Hist.; Sir John Richardson collector. 

Type locality.—California, here restricted to San Diego. 

Range.—Pacific slopes of central and southern California, and 
extreme northern Baja California, west of the desert and the Sierra 

San Pedro MArtir. Recorded in Baja California: San Salado Cafion, 

San José, San Quintin, Los Coronados Islands. 


Subclass ARCHOSAURIA 


Archosauria RomER, Vertebrate paleontology, 1945, p. 597. 
Orders.—A single living order, the Loricata, exists. 


Order LORICATA 


Loricata Mrrrem, Tentamen systematis amphibiorum, 1820, p. 34. 


Families —Three families exist at the present time, two of which 
occur in Mexico. 


210 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


KEY TO LIVING FAMILIES OF LORICATA 


1. Dentary teeth 25-26, none received by sockets in upper jaw; maxillary bones 
broadly in contact with each other on dorsal surface of skull; mandibular 
symphysis extraordinarily long, only 2 or 3 teeth at rear of mandible not 
opposite tite symp yas aac F be Cote te I ae Gavialidae ° 

Dentary teeth 20 or fewer, all or nearly all received by sockets in upper jaw; 
maxillary bones never in contact with each other on dorsal surface of skull; 
mandibular symphysis shorter, 4 or more teeth at rear of mandible not 


opposite the symphysis 9 © 25 rie ea eI el 2 

2. Fourth mandibular tooth fitting into a pit in upper jaw, and 17 to 22 dentary 
LOC ke wae sess ape apres 2 oy Ea ee ye Alligatoridae (p. 211) 
Fourth mandibular tooth fitting into a notch in upper jaw or, if into a pit, 
dentarysteethy 1 6.or: fewer: 222-5. 524352602520. 265 Crocodylidae (p. 210) 


Family CROCODYLIDAE Gray 


Crocodilidae Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 195.—Mzrrens, Sencken- 
bergiana, vol. 26, 1948, pp. 252-312. 
Genera.—Four genera and 17 species and subspecies are recognized 
at the present time; one genus occurs in the Americas. 
Range.— World wide in tropical lands. 


Genus CROCODYLUS Laurenti 


Crocodylus LauRENTI, Specimen medicum exhibens synopsin reptilium, 1768, p. 53. 

Molinia Gray, Catalogue of the tortoises, crocodiles and amphisbaenians in the 
collection of the British Museum, 1844, pp. 60, 272 (type, Crocodylus ameri- 
canus Laurenti=? Crocodylus acutus Cuvier). 

Genotype.—Crocodylus niloticus Laurenti. 

Species.— Twelve species, two with two subspecies, are recognized; 
only four species, one with two subspecies, occur in the New World, 
and two in Mexico. 

Range.—Northern Australia, New Guinea, East Indies, Siam, 
Ceylon, tropical Africa and India, Madagascar; southern Florida, 
Greater Antilles, northern Mexico south to Ecuador and Venezuela. 


KEY TO MEXICAN SPECIES OF CROCODYLUS 


1. Premaxillo-maxillary suture transverse__--_-------------- moreletii (p. 211) 
Premaxillo-maxillary suture extending posteriorly in an arc to level of posterior 
border of seventh toothy 2202 52 2 ee ea acutus (p. 210) 


CROCODYLUS ACUTUS ACUTUS Cuvier * 


Crocodilus acutus Cuvier, Ann. Mus. Hist. Nat. Paris, vol. 10, 1807, p. 55, pl. 
1, figs, pli 2, fe. o. 
Crocodylus acutus acutus, Miuter and Hewumicu, Ibero-Amerik. Stud., vol. 
13, 1940, pp. 128-130. 


— 


5 Not in Mexico; restricted to India. 

6 Allocation of the Mexican and, for that matter, Central American specimens is at present very uncertain. 
South American (Colombia, Ecuador) specimens are referrable to the race C. a. lewyanus, and Santo Do 
mingo specimens to C. a. acutus, but no study has been made of material from elsewhere. 


CHECKLIST OF REPTILES OF MEXICO 211 


? Crocodilus americanus LAURENTI, Specimen medicum exhibens synopsin rep- 
tilium, 1768, p. 54 (type, pl. 106 in Seba’s “Locupletissimi Rerum Naturalium 
Thesauri Occurata Descriptio,’ vol. 1, 1734; type locality, America, here 
restricted to Veracruz, Veracruz; unidentifiable).—Bocourt, Mission scien- 
tifique au Mexique ..., Etudes sur les reptiles, livr. 1, 1870, pp. 
30-31, pl. 8, fig. 1, pl. 9, fig. 1. 

Crocodilus biscutatus Cuvier, Ann. Mus. Hist. Nat. Paris, vol. 10, 1807, p. 53, 
pl. 2, fig. 6 (type presumably in Mus. Hist. Nat. Paris; type locality, 
‘Senegal’, in error; here restricted to Tampico, Tamaulipas). 

Crocodilus mezicanus Bocourt, Nouv. Arch. Mus. Hist. Nat. Paris, 1869, pp. 
20-21 (type presumably in Mus. Hist. Nat. Paris; type locality, Tampico, 
Tamaulipas); Mission scientifique au Mexique . .. , Etudes sur les reptiles, 
livr. 1, 1870, pp. 34-35, pl. 8, fig. 3. 

Crocodilus pacificus Dumériu and Bocourt, Mission scientifique au Mexique .. ., 
Etudes sur les reptiles, livr. 1, 1870, pp. 31-33, pl. 9, fig. 5 (type presumably 
in Mus. Hist. Nat. Paris; type locality, Rio Nagualate, Guatemala). 


Type.—Mus. Hist. Nat. Paris, presumably. 

Type locality —Santo Domingo (Republic). 

Range.—Greater Antilles, Central America north to Tamaulipas 
and Sinaloa. Reported from various localities in the states of 
Quintana Roo (Isla de Mujeres), Campeche, Colima, Guerrero, 
Michoacan, Oaxaca, Tabasco, Tamaulipas, Veracruz, and Maria 
Magdalena Island, Nayarit. 


CROCODYLUS MORELETI Duméril and Duméril 


Crocodilus moreletii DuMéRIL and DumErRIL, Catalogue méthodique de la collection 
des reptiles, 1851, p. 28.—Bocourt, Mission scientifique au Mexique . . ., 
Etudes sur les reptiles, livr. 1, 1870, pp. 37-38, pl. 9, fig. 2.—Scumip7, Publ. 
Field Mus. Nat. Hist., zool. ser., vol. 12, 1924, pp. 79-84, pl. 6, figs. 2, 7, 8. 

Alligator lacordairei PREUDHOMME DE Bork, Bull. Acad. Roy. Sci. Lett. Beaux- 
Arts Belgique, ser. 2, vol. 28, 1869, p. 110, pl. (type presumably in Brussels; 
M. Levy collector; type locality, Belize, British Honduras). 


Type.—Mus. Hist. Nat. Paris, presumably. 

Type locality —Lake Petén, Guatemala. 

Range.—Atlantic slopes from Guatemala to Tamaulipas. Reported 
in Mexico from the states of Campeche, Chiapas (Palenque), Colima, 
Tabasco, Tamaulipas, Veracruz; records from Colima (by Dugés) 
and the Pacific side of Chiapas (Tapachula, by Sumichrast) are to be 
considered erroneous until corroborated. 


Family ALLIGATORIDAE Gray 


Alligatoridae Gray, Catalogue of tortoises, crocodiles and amphisbaenians in the 
British Museum, 1844, p. 56.—Mertens, Senckenbergiana, vol. 26, 1943, 
pp. 252-312. 


Genera.—F our, of which only one occurs in Mexico.’ 
17 Alligator mississippiensis is recorded from ‘‘Mexico” by Garman (Bull. Essex Inst., vol. 16, 1884, p. 11) 


and Boettger (Kat. Rept. Senckenb. Mus., pt. 1, 1893, p. 19), but we regard the records as erroneous 
until verified; 


212 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Genus CAIMAN Spix 


Caiman Srix, Animalia nova site species lacertarum ... Brasiliam . . ., 1825, p. 3. 
Perosuchus Corn, Proc. Acad. Nat. Sci. Philadelphia, 1868, p. 203 (genotype, 
Perosuchus fuscus Cope). 
Genotype.—Caiman fissipes Spix (=Crocodilus latirostris Daudin). 
Species.—Two species, one with three subspecies, are recognized; 
one occurs in Mexico. 
Range.—The Isthmus of Tehuantepec to Argentina. 


CAIMAN CROCODILUS FUSCUS (Cope) 


Perosuchus fuscus Cops, Proc. Acad. Nat. Sci. Philadelphia, 1868, p. 203. 

Caiman crocodilus fuscus, MERTENS, Senckenbergiana, vol. 26, 1943, p. 275. 

Allifigator] (Jacare) Chiaparius Bocourt, Ann. Sci. Nat., ser. 6, vol. 3, No. 12, 
1876, pp. 1-2 (type probably in Mus. Hist. Nat. Paris; type locality, Tonald4, 
Chiapas). 

Allfigator] Chiapasius Bocourt, Journ. Zool., vol. 5, 1876, pp. 400-401 (equivalent 
to preceding). 

Type.—Unknown. 

Type locality.— Rio Magdalena, Colombia. 

Range.—Oaxaca and perhaps Michoacan to Colombia. Reported 
from the states of Oazraca: Tapanatepec, Agua Fria; and Chiapas: 
Tonalé, Belén, Colonia Soconusco. A record from the Balsas 
Valley in Michoac4n (by Gadow) may be correct, but one from 
Yucatan (Tozzer) is, we believe, certainly incorrect. 


SPECIES INQUIRENDAE 


A few species whose natural ranges remain unknown have been 
included in their appropriate systematic place in the preceding pages. 
A few others, described from ‘‘Mexico,”’ have since been found to occur 
elsewhere. Among these are the following: 


HERPETOCHALCIS HETEROPUS Boettger 
Herpetochalcis heteropus BorrtcEr, Bericht. Offenb. Ver. Nat., vols. 22-23, 1883, 
pp. 150-151. 


The type locality was stated to be perhaps California, or Mexico, 
or Central America. The name is apparently a synonym of Chalcides 
heteropus Lichtenstein, Nomencl. Mus. Zool. Berol., 1856, p. 17 
(=Bachia heteropa) described from La Guaira. Venezuela, and re- 
stricted in range to that country. 


LEIOSAURUS BELLI Duméril and Bibron 


Leiosaurus belli DumMi&RIL and Brpron, Erpétologie générale, vol. 4, 1837, p. 242. 


The type locality was stated to be ‘“Mexico,” but the species (and 
genus) has since been discovered to be restricted to Argentina. 


CHECKLIST OF REPTILES OF MEXICO 213 


ANEUPORUS OCCIPITALIS Bocourt 
Aneuporus occipitalis Bocourt, Mission scientifique au Mexique .. ., Etudes sur 
les reptiles, livr. 4, 1874, pp. 215-217, pl. 18, fig. 1. 
This has since been renamed Tropidurus bocourti by Boulenger, since 
Bocourt’s name is suppressed as a homonym of Tropidurus occipitalis 
(Peters). The species occurs in Peru. 


SPHAERODACTYLUS ANTHRACINUS Cope 
Sphaerodactylus anthracinus Corn, Proc. Acad. Nat. Sci. Philadelphia, 1861, p. 
500 (type locality, ‘‘Mexico” [Jalapa]). 

Barbour (Mem. Mus. Comp. Zool. Harvard, vol. 47, 1921, pp. 
258-259) associates this name with a species from Andros Island, but 
Boulenger (Catalogue of the lizards in the British Museum, vol. 1, 
1885, p. 225) records a specimen from ‘San Domingo,” and Peters 
(Monatsb. Akad. Wiss. Berlin, 1869, p. 874) records one from the 
state of Puebla (possibly in the vicinity of Izticar de Matamoros). 
The final word has not yet been said. For a discussion see Taylor, 
Univ. Kansas Sci. Bull., vol. 31, 1947, pp. 300-301. 


Three other species recorded from ‘Mexico,’ but with a type 
locality elsewhere, are known to be restricted to extralimital areas: 


BACHIA DORBIGNYI (Duméril and Bibron) 


Bachia dorbignyi (Duméril and Bibron), Mit.umr, Reisen in den Vereinigten 
Staaten, Canada und Mexico, vol. 3, pt. 3, 1865, p. 604. 


This species, still recognized by this name, occurs in Bolivia and 
Chile. 
ECPHYMOTES OBTUSIROSTRIS (Wiegmann) 
Ecphymotes obtusirostris (Wiegmann), MUuuEr, op. cit., p. 600. 
This name is a synonym of Anisolepis undulatus (Wiegmann), 
which species occurs in Brasil. 
TROPIDURUS PTYCHOPLEURUS Lichtenstein 


Tropidurus ptychopleurus Lichtenstein, MULLER, op. cit., p. 602. 


This name is a synonym of Liolaemus tenuis (Duméril and Bibron), 
which species occurs in Chile. 


One other name, supposedly based upon a Mexican specimen, has 
been impossible to place: 


214 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


DASIA MICROCEPHALUS (Hallowell) 


Euprepis microcephalus HALLOWELL, Proc. Acad. Nat. Sci. Philadelphia, vol. 8, 
1856, p. 155; Trans. Amer. Philos. Soc., ser. 2, vol. 11, 1857, pp. 79-80. 
Diploglossus microcephalus, BoULENGER, Catalogue of the lizards in the British 

Museum, vol. 3, 1887, p. 504. 

[Dasia] microcephalus, Smitu, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 369-370. 
Type.—Acad. Nat. Sci. Philadelphia; W. H. Keating collector. 
Type locality.—Mexico. 

Remarks.—The first words in Hallowell’s original description are 
“Syn. Scincus ventralis, Peale and Green.’”’ Then follows the descrip- 
tion. We believe the citation of Peale and Green’s name as a synonym 
(an earlier one at that) of Hallowell’s own name could be construed 
to make his name unavailable, that is, actually a synonym of Scincus 
ventralis and thus a part of the synonymy of Gerrhonotus l. liocephalus. 
The fact remains that the animal described by Hallowell, and pre- 
sumably the type of his name, is far different from Gerrhonotus. We 
recommend association of the name with the species represented by 
the animal described and not assignment to the synonomy of Scincus 
ventralis. 

The type is not in good condition (portions of the head mutilated), 
but appears to belong to a non-American genus, probably Dasia.® 
It may possibly be one of the original series of three specimens of 
Peale and Green’s Scincus ventralis, since only two of them are now 
present in the series labeled as cotypes, and thus would be explained 
Hallowell’s citation of Scincus ventralis as a synonym of his species. 
It is also possible that a confusion of specimens occurred, the original 
Gerrhonotus being exchanged for the present type of microcephalus, 
which Hallowell erroneously thought was one of Peale and Green’s 
cotypes. One of these alternatives must be true: Either a peculiar 
skink, unknown except by the type of microcephalus, occurs in Mexico, 
or else some shift of specimens occurred in the Philadelphia Academy 
collections between 1830 and 1856. We favor the latter alternative. 


Still another species is represented by a specimen questionably 
secured in Mexico, while the range of the species is suspected of being 
entirely extralimital. 


DIPLOGLOSSUS MONOTROPIS (Kuhl) 


Diploglossus monotropis (KuHL), Beitrige zur Zoologie und vergleichende Ana- 
tomie, 1820, p. 128. 


The species is known to occur from Costa Rica to Ecuador. The 
U.S. National Museum has one specimen questionably from ‘‘Colima.”’ 
Occurrence in Mexico is highly questionable. 


8 We are indebted to Dr. E. R. Dunn for this information. 


CHECKLIST OF REPTILES OF MEXICO 215 


STATE LISTS 


The accompanying lists have been constructed in the same way as 
those for the snakes (U. S. Nat. Mus. Bull. 187, 1945, pp. 202-203) 
and amphibians (U. S. Nat. Mus. Bull. 194, 1948, pp. 100-101). 
Table 2 shows for each state and each of the indicated groups, first, 
the number of forms recorded; second, the number of genera repre- 
sented (these two numbers separated by a diagonal line); and third, 
the relative position of the state (so far as number of forms is con- 
cerned) as compared with other states (this number in parentheses). 
The lists of snakes and amphibians have been brought more or less 
up to date by inclusion of the additional forms defined and recorded 
by various authors such as Blair, Bogert, Burger, Conant, Klauber, 
Smith, Tanner, and Taylor (see Univ. Kansas Sci. Bull., vol. 33, 1950, 
pp. 313-380, for references). 


BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


216 





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CHECKLIST OF REPTILES OF MEXICO 


217 


AGUASCALIENTES 


Kinosternon integrum 
Cnemidophorus sackii communis 


Sceloporus spinosus spinosus 
Sceloporus torquatus melanogaster 


BAJA CALIFORNIA 


Amyda emoryi 

Caretta caretta gigas 

Clemmys marmorata pallida 
Dermochelys coriacea 

Eretmochelys imbricata 

Gopherus agassizii 

Pseudemys scripta nebulosa 

Bipes biporus 

Anniella geronimensis 

Anniella pulchra pulchra 

Callisaurus draconoides draconoides 
Callisaurus draconoides carmenensis 
Callisaurus draconoides crinitus 
Callisaurus draconoides gabbii 
Callisaurus draconoides splendidus 
Cnemidophorus catalinensis 
Cnemidophorus ceralbensis 
Cnemidophorus hyperythrus beldingi 
Cnemidophorus hyperythrus caeruleus 
Cnemidophorus hyperythrus danheimae 
Cnemidophorus hyperythrus hyperythrus 
Cnemidophorus hyperythrus pictus 
Cnemidophorus hyperythrus schmidti 
Cnemidophorus labialis 
Cnemidophorus maximus 
Cnemidophorus tigris canus 
Cnemidophorus tigris celeripes 
Cnemidophorus tigris multiscutatus 
Cnemidophorus tigris rubidus 
Cnemidophorus tigris tigris 
Coleonyx variegatus variegatus 
Coleonyx variegatus abbotti 
Coleonyx variegatus peninsularis 
Coleonyx variegatus slevini 
Crotaphytus collaris baileyi 
Crotaphytus insularis 

Ctenosaura hemilopha 

Dipsosaurus carmenensis 
Dipsosaurus catalinensis 

Dipsosaurus dorsalis dorsalis 
Dipsosaurus dorsalis lucasensis 
Elgaria cedrosensis 

Elgaria multicarinata nana 

Elgaria multicarinata webbii 

Elgaria paucicarinata 

Eumeces gilberti rubricaudatus 
Eumeces lagunensis 


Eumeces skiltonianus skiltonianus 
Gambelia wislizenii wislizenii 
Petrosaurus repens 

Petrosaurus thalassinus 
Phrynosoma cerroense 
Phrynosoma coronatum coronatum 
Phrynosoma coronatum blainvillii 
Phrynosoma coronatum frontale 
Phrynosoma coronatum jamesi 
Phrynosoma m’callii 

Phrynosoma platyrhinos platyrhinos 
Phrynosoma solare 
?Phyllodactylus tuberculosus 
Phyllodactylus unctus 

Sator angustus 

Sator grandaevus 

Sauromalus ater 

Sauromalus australis 

Sauromalus hispidus 

Sauromalus klauberi 

Sauromalus slevini 

Sceloporus graciosus vandenburgianus 
Sceloporus magister lineatulus 
Sceloporus magister magister 
Sceloporus magister monserratensis 
Sceloporus magister rufidorsum 
Sceloporus magister zosteromus 
Sceloporus occidentalis biseriatus 
Sceloporus orecutti licki 
Sceloporus orcutti orcutti 
Streptosaurus mearnsi 
Streptosaurus slevini 

Uma notata notata 

Urosaurus graciosus 

Urosaurus microscutatus 
Urosaurus nigricaudus 

Urosaurus ornatus symmetricus 
Uta concinna 

Uta mannophorus 

Uta martinensis 

Uta squamata 

Uta stansburiana elegans 

Uta stansburiana hesperis 

Uta stansburiana stejnegeri 

Uta stellata 

Xantusia gilberti 

Xantusia henshawi 

Xantusia vigilis 


218 


BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


CAMPECHE 


Chelydra serpentina 

Claudius angustatus 
Dermatemys mawii 
Eretmochelys imbricata 
Geoemyda areolata 

Kinosternon acutum 
Kinosternon cruentatum cruentatum 
Kinosternon leucostomum 
Pseudemys scripta ornata 
Ameiva undulata gaigeae 
Ameiva undulata stuarti 

Anolis kidderi 

Anolis lemurinus bourgeaei 
Anolis limifrons rodriguezii 
Anolis sagrei mayensis 

Anolis tropidonotus 

Anolis ustus 

Basiliscus vittatus 
Cnemidophorus deppii oligoporus 


Cnemidophorus sackii angusticeps 
Coleonyx elegans elegans 
Corythophanes cristatus 
Ctenosaura similis similis 
Enyaliosaurus erythromelas 
Eumeces schwartzei 
Hemidactylus turcicus turcicus 
Tguana iguana rhinolopha 
Laemanctus serratus 

Scincella cherriei ixbaac 
Mabuya mabouya mabouya 
Sceloporus chrysostictus 
Sceloporus lundelli lundelli 
Seeloporus serrifer serrifer 
Sceloporus serrifer plioporus 
Sceloporus teapensis 
Sphaerodactylus glaucus glaucus 
Crocodylus acutus acutus 
Crocodylus moreletii 


CHIAPAS 


Geoemyda pulcherrima incisa 
Kinosternon abaxillare 
Kinosternon cruentatum cruentatum 
Kinosternon leucostomum 
Lepidochelys olivacea 
Pseudemys scripta ornata 
Abronia matudai 

Abronia ochoterenai 

Ameiva festiva edwardsi 
Ameiva undulata hartwegi 
Ameiva undulata parva 
Ameiva undulata stuarti 
Ameiva undulata thomasi 
Anolis biporeatus 

Anolis humilis uniformis 
Anolis limifrons rodriguezii 
Anolis pentaprion 

Anolis sericeus 

Barisia moreleti rafaeli 

Barisia moreleti temporalis 
Basiliscus vittatus 

Celestus rozellae 
Cnemidophorus deppii deppii 
Cnemidophorus guttatus immutabilis 
Cnemidophorus sackii bocourti 
Coleonyx elegans elegans 
Corythophanes cristatus 
Corythophanes hernandezii 
Corythophanes percarinatus 


Ctenosaura similis similis 

Eumeces sumichrasti 

Gerrhonotus liocephalus austrinus 

Gonatodes fuscus 

Heloderma horridum 

Iguana iguana rhinolopha 

Scincella assata assata 

Scincella assata taylori 

Scincella cherriei cherriei 

Lepidophyma flavimaculata flavimacu- 
lata 

Lepidophyma smithii smithii 

?Lepidophyma smithii tehuanae 

Mabuya mabouya mabouya 

Phrynosoma asio 

Phyllodactylus magnus 

Sceloporus carinatus 

Sceloporus malachiticus acanthinus 

Seeloporus malachiticus taeniocnemis 

Sceloporus melanorhinus stuarti 

Sceloporus prezygus 

Sceloporus siniferus siniferus 

Sceloporus squamosus 

Sceloporus teapensis 

Sceloporus variabilis variabilis 

Urosaurus bicarinatus anonymorphus 

Xenosaurus rackhami 

Caiman crocodilus fuscus 

Crocodylus moreletii 


CHECKLIST OF REPTILES OF MEXICO 


219 


CHIHUAHUA 


Chrysemys picta bellii 
?Gopherus berlandieri 
Kinosternon hirtipes 
Kinosternon sonoriense 
Terrapene ornata 

Anolis nebuloides 

Barisia levicollis 

Cnemidophorus inornatus 
Cnemidophorus sackii communis 
Cnemidophorus sackii stictogrammus 
Cnemidophorus sackii scalaris 
Cnemidophorus tigris marmoratus 
Crotaphytus collaris baileyi 
Ctenosaura hemilopha 

Elgaria kingii kingii 

Eumeces callicephalus 

Eumeces multivirgatus 

Eumeces obsoletus 

Eumeces parviauriculatus 
Gambelia wislizenii wislizenii 
Holbrookia bunkeri 


Holbrookia maculata approximans 
Holbrookia texana 

Phrynosoma cornutum 
Phrynosoma douglassii hernandesi 
Phrynosoma modestum 
Phrynosoma orbiculare orbiculare 
Phyllodactylus homolepidurus 
Sceloporus clarkii clarkii 
Sceloporus grammicus disparilis 
Sceloporus jarrovii jarrovii 
Sceloporus magister magister 
Sceloporus nelsoni 

“eloporus poinsettii 

Sceloporus sealaris slevini 
Sceloporus undulatus consobrinus 
Sceloporus undulatus virgatus 
Urosaurus ornatus caeruleus 
Urosaurus ornatus linearis 
Urosaurus ornatus schmidti 
Urosaurus unicus 

Uta stansburiana stejnegeri 


COAHUILA 


Amyda emoryi 

Gopherus berlandieri 

Kinosternon flavescens flavescens 
Kinosternon flavescens stejnegeri 
Pseudemys scripta elegans 
Pseudemys scripta gaigeae 
Terrapene coahuila 

Barisia imbricata ciliaris 
Cnemidophorus inornatus 
Cnemidophorus sackii communis 
Cnemidophorus sackii semifasciatus 
Cnemidophorus sackii scalaris 
Cnemidophorus tigris marmoratus 
Coleonyx brevis 

Crotaphytus collaris baileyi 
Crotaphytus reticulatus 

Eumeces obsoletus 

Gambelia wislizenii wislizenii 
Gerrhonotus liocephalus infernalis 
Holbrookia maculata approximans 
Holbrookia maculata dickersonae 


861316—50——-15 


Holbrookia maculata lacerata 
Holbrookia texana 

Scincella laterale 

Phrynosoma cornutum 
Phrynosoma modestum 
Sceloporus cautus 

Sceloporus couchii 

Sceloporus goldmani 
Sceloporus grammicus disparilis 
Sceloporus jarrovii oberon 
Sceloporus magister magister 
Sceloporus merriami annulatus 
Sceloporus olivaceus 

Sceloporus ornatus caeruleus 
Sceloporus ornatus ornatus 
Sceloporus parvus parvus 
Sceloporus poinsettii 

Sceloporus sealaris slevini 
Sceloporus undulatus consobrinus 
Sceloporus variabilis marmoratus 
Uma exsul 

Uta stansburiana stejnegeri 


220 


BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


COLIMA 


Geoemyda rubida 

Geoemyda pulcherrima pulcherrima 
Kinosternon hirtipes 

Kinosternon integrum 

Lepidochelys olivacea 

Ameiva undulata sinistra 

Anolis nebuloides 

Anolis nebulosus 

Anolis schmidti 

Basiliscus vittatus 

Cnemidophorus deppii lineatissimus 
Cnemidophorus guttatus immutabilis 
Cnemidophorus sackii communis 
Coleonyx elegans nemoralis 
Ctenosaura pectinata 

Eumeces colimensis 

Eumeces parvulus 

Heloderma horridum 


DISTRITO 


Kinosternon hirtipes 

Barisia imbricata imbricata 
Cnemidophorus sackii communis 
Eumeces copei 

Gerrhonotus liocephalus liocephalus 
Phrynosoma orbiculare orbiculare 


Iguana iguana rhinolopha 
?Laemanctus longipes 

Scincella assata taylori 

Mabuya mabouya mabouya 
Phrynosoma asio 

Phrynosoma orbiculare dugesii 
Phyllodactylus lanei 

Sceloporus dugesii dugesii 
Sceloporus grammicus microlepidotus 
Sceloporus horridus oligoporus 
Sceloporus melanorhinus calligaster 
Sceloporus pyrocephalus 
Sceloporus utiformis 

Urosaurus auriculatus 

Urosaurus bicarinatus tuberculatus 
Urosaurus clarionensis 

Crocodylus acutus acutus 
Crocodylus moreletii 


FEDERAL 


Sceloporus aeneus aeneus 

Sceloporus grammicus microlepidotus 
Sceloporus scalaris scalaris 
Sceloporus spinosus spinosus 
Sceloporus torquatus torquatus 


DURANGO 


Kinosternon fiavescens stejnegeri 
Kinosternon sonoriense 
Pseudemys scripta gaigeae 

Anolis nebulosus 

Barisia imbricata ciliaris 
Cnemidophorus sackii communis 
Cnemidophorus sackii scalaris 
Cnemidophorus tigris marmoratus 
Coleonyx brevis 

Coleonyx fasciatus 

Crotaphytus collaris baileyi 
Ctenosaura pectinata 

EKumeces brevirostris 

Eumeces callicephalus 

Holbrookia maculata approximans 
Holbrookia maculata dickersonae 
Holbrookia texana 


Phrynosoma cornutum 
Phrynosoma douglassii brachycercum 
Phrynosoma modestum 
Phrynosoma orbiculare orbiculare 
?Phyllodactylus lanei 

Sceloporus grammicus disparilis 
Sceloporus horridus oligoporus 
Sceloporus jarrovii jarrovii 
Sceloporus lineolateralis 
Seceloporus maculosus 

Sceloporus magister magister 
Sceloporus poinsettii 

Sceloporus scalaris scalaris 
Sceloporus scalaris slevini 
Sceloporus spinosus spinosus 
Sceloporus undulatus consobrinus 
Uta stansburiana stejnegeri 


CHECKLIST OF REPTILES OF MEXICO 


221 


GUANAJUATO 


Kinosternon hirtipes 

Kinosternon integrum 

Barisia imbricata ciliaris 

Barisia imbricata imbricata 
Cnemidophorus sackii communis 
Eumeces callicephalus 

Eumeces dugesii 

Gerrhonotus liocephalus liocephalus 
Holbrookia maculata dickersonae 
Laemanctus serratus 
Phrynosoma boucardi 


Phrynosoma orbiculare orbiculare 
Sceloporus aeneus aeneus 
Sceloporus dugesii intermedius 
Sceloporus grammicus disparilis 
Sceloporus grammicus microlepidotus 
Sceloporus jarrovii minor 
Sceloporus scalaris scalaris 
Sceloporus spinosus spinosus 
Sceloporus torquatus melanogaster 
Sceloporus torquatus torquatus 
Sceloporus variabilis variabilis 


GUERRERO 


Chelonia mydas 

Geoemyda pulcherrima pulcherrima 
Geoemyda rubida 

Kinosternon integrum 

Lepidochelys olivacea 

Abronia deppii 

Ameiva undulata dextra 

Anolis dunni 

Anolis gadovii 

Anolis liogaster 

Anolis megapholidotus 

Anolis nebuloides 

Anolis nebulosus 

Anolis taylori 

Barisia gadovii gadovii 

Basiliscus vittatus 

Bipes canaliculatus 

Bipes tridactylus 

Cnemidophorus deppii deppii 
Cnemidophorus deppii lineatissimus 
Cnemidophorus guttatus immutabilis 
Cnemidophorus sackii sackii 
Coleonyx elegans nemoralis 
Ctenosaura pectinata 

Eumeces brevirostris 

Eumeces ochoterenai 

Gerrhonotus liocephalus liocephalus 
Heloderma horridum 


861316—50 16 





Hemidactylus frenatus 

Iguana iguana rhinolopha 

Scincella assata taylori 

Mabuya mabouya mabouya 
Phrynosoma asio 

Phrynosoma taurus 

Phyllodactylus bordai 
Phyllodactylus deleampi 
Phyllodactylus lanei 
Phyllodactylus magnatuberculatus 
Phyllodactylus magnus 

?Sceloporus asper 

Sceloporus formosus scitulus 
Sceloporus gadoviae 

Sceloporus grammicus grammicus 
Sceloporus horridus horridus 
Sceloporus horridus oligoporus 
Sceloporus melanorhinus calligaster 
Sceloporus mucronatus omiltemanus 
Sceloporus ochoterenai 

Sceloporus pyrocephalus 

Sceloporus siniferus siniferus 
Sceloporus stejnegeri 

Sceloporus utiformis 

Urosaurus bicarinatus bicarinatus 
Urosaurus bicarinatus anonymorphus 
Crocodylus acutus acutus 


222 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


HIDALGO 


Kinosternon hirtipes 

Abronia taeniata taeniata 
Barisia imbricata imbricata 
Barisia imbricata ciliaris 
Cnemidophorus sackii communis 
Eumeces lynxe furcirostris 
Eumeces lynxe lynxe 

Gaigeia gaigeae 

Gaigeia sylvatica 

Laemanctus serratus 

Scincella gemmingeri gemmingeri 
Scincella gemmingeri forbesorum 


Kinosternon integrum 
Pseudemys scripta ornata 
Ameiva undulata sinistra 
Anolis nebulosus 

Barisia imbricata imbricata 
Basiliscus vittatus 


Cnemidophorus deppii lineatissimus 


Cnemidophorus sackii communis 
Ctenosaura pectinata 

Eumeces brevirostris 

Eumeces callicephalus 

Eumeces dugesii 

Heloderma horridum 

Holbrookia maculata approximans 
Iguana iguana rhinolopha 
Scincella assata taylori 
Phrynosoma orbiculare orbiculare 
?Phyllodactylus lanei 

?Sceloporus aeneus aeneus 


Kinosternon hirtipes 

Barisia imbricata imbricata 
Barisia rudicollis 
Cnemidophorus sackii communis 
Eumeces copei 

Eumeces indubitus 


Gerrhonotus liocephalus liocephalus 


Phrynosoma orbiculare orbiculare 
Sceloporus aeneus aeneus 


Phrynosoma boucardi 

Phrynosoma orbiculare orbiculare 
Sceloporus aeneus bicanthalis 
Sceloporus grammicus disparilis 
Sceloporus grammicus microlepidotus 
Sceloporus jarrovii immucronatus 
Sceloporus mucronatus mucronatus 
Sceloporus parvus scutulatus 
Sceloporus scalaris scalaris 
Sceloporus spinosus spinosus 
Sceloporus torquatus torquatus 
Sceloporus variabilis variabilis 


JALISCO 


Sceloporus asper 

Sceloporus bulleri 

Sceloporus clarkii boulengeri 
Sceloporus dugesii dugesii 
Sceloporus grammicus microlepidotus 
Sceloporus heterolepis 

Sceloporus horridus albiventris 
Sceloporus horridus oligoporus 
Sceloporus melanorhinus calligaster 
Sceloporus nelsoni 

Sceloporus pyrocephalus 
Sceloporus scalaris scalaris 
Sceloporus scalaris unicanthalis 
Sceloporus spinosus spinosus 
Sceloporus torquatus melanogaster 
Sceloporus utiformis 

Urosaurus bicarinatus tuberculatus 
Urosaurus gadovi 


MEXICO 


Sceloporus aeneus bicanthalis 
Sceloporus grammicus microlepidotus 
Sceloporus jarrovii sugillatus 
Sceloporus jarrovii minor 

Sceloporus mucronatus mucronatus 
Sceloporus scalaris scalaris 
Sceloporus spinosus spinosus 
Sceloporus torquatus torquatus 


CHECKLIST OF REPTILES OF MEXICO 


223 


MICHOACAN 


Geoemyda rubida 

Kinosternon hirtipes 
Kinosternon integrum 

Ameiva undulata sinistra 

Anolis nebulosis 

Barisia imbricata imbricata 
Basiliscus vittatus 
Cnemidophorus deppii lineatissimus 
Cnemidophorus guttatus immutabilis 
Cnemidophorus sackii communis 
Cnemidophorus sackii sackii 
?Coleonyx elegans nemoralis 
Ctenosaura pectinata 
Enyaliosaurus clarki 

Eumeces altamirani 

Eumeces brevirostris 

Eumeces callicephalus 

Eumeces copei 

Eumeces dugesii 

Eumeces indubitus 

Heloderma horridum 


Iguana iguana rhinolopha 

Mabuya mabouya mabouya 
Phrynosoma asio 

Phrynosoma orbiculare orbiculare 
Phyllodactylus lanei 

Sceloporus aeneus aeneus 
Sceloporus asper 

Sceloporus dugesii intermedius 
Sceloporus gadoviae 

Sceloporus grammicus microlepidotus 
Sceloporus horridus oligoporus 
Sceloporus melanorhinus calligaster 
Sceloporus pyrocephalus 
Sceloporus scalaris scalaris 
Sceloporus spinosus spinosus 
Sceloporus torquatus torquatus 
Sceloporus torquatus melanogaster 
Sceloporus utiformis 

Urosaurus bicarinatus bicarinatus 
Urosaurus gadovi 

Crocodylus acutus acutus 


MORELOS 


Kinosternon integrum 

Ameiva undulata sinistra 

Anolis nebulosus 

Barisia imbricata imbricata 
Cnemidophorus guttatus immutabilis 
Cnemidophorus sackii sackii 
Ctenosaura pectinata 
?Enyaliosaurus clarki 

Eumeces copei 

Eumeces indubitus 

Gerrhonotus liocephalus liocephalus 


Heloderma horridum 

Phrynosoma orbiculare orbiculare 
Sceloporus aeneus aeneus 

Sceloporus gadoviae 

Sceloporus grammicus microlepidotus 
Sceloporus horridus horridus 
Sceloporus ochoterenai 

Sceloporus siniferus siniferus 
Sceloporus torquatus torquatus 
Urosaurus bicarinatus bicarinatus 


NAYARIT 


Chelonia mydas 

Geoemyda pulcherrima pulcherrima 
Kinosternon hirtipes 

Kinosternon integrum 

Terrapene nelsoni 

Anolis nebulosus 

Cnemidophorus deppii lineatissimus 
Cnemidophorus sackii communis 
Cnemidophorus sackii mariarum 
Ctenosaura pectinata 

Eumeces callicephalus 

Eumeces parvulus 


Iguana iguana rhinolopha 
Peropus mutilatus 
Phyllodactylus lanei 
Sceloporus asper 

Sceloporus clarkii boulengeri 
Sceloporus dugesii dugesii 
Sceloporus horridus albiventris 
Sceloporus melanorhinus calligaster 
Sceloporus nelsoni 

Sceloporus utiformis 
Urosaurus ornatus schottii 
Crocodylus acutus acutus 


224. BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


NUEVO LEON 


Amyda emoryi 

Gopherus berlandieri 
Pseudemys floridana texana 
Pseudemys scripta elegans 
Barisia imbricata ciliaris 
Cnemidophorus inornatus 
Cnemidophorus sackii gularis 
Coleonyx brevis 
Crotaphytus collaris baileyi 
Crotaphytus reticulatus 
Eumeces brevilineatus 
Eumeces dicei 

Eumeces obsoletus 
Holbrookia texana 

Scincella caudaequinae 


Phrynosoma cornutum 
Phrynosoma modestum 
Phrynosoma orbiculare orbiculare 
Sceloporus couchii 

Sceloporus cyanogenys 
Sceloporus grammicus disparilis 
Sceloporus jarrovii minor 
Sceloporus olivaceus 

Sceloporus parvus parvus 
Sceloporus poinsettii 

Sceloporus scalaris slevini 
Sceloporus torquatus binocularis 
Sceloporus undulatus consobrinus 
Sceloporus variabilis marmoratus 


OAXACA 


Chelonia mydas 

Dermatemys mawii 
Eretmochelys imbricata 
Geoemyda pulcherrima incisa 
Geoemyda rubida 

Kinosternon cruentatum cruentatum 
Kinosternon integrum 
Lepidochelys olivacea 
Pseudemys scripta ornata 
Pseudemys umbra 

Staurotypus salvinii 

Abronia fuscolabialis 

Abronia oaxacae 

Ameiva undulata amphigramma 
Ameiva undulata dextra 
Ameiva undulata parva 

Ameiva undulata stuarti 
Ameiva undulata undulata 
Anolis heliactin 

Anolis limifrons rodriguezii 
Anolis milleri 

Anolis nebuloides 

Anolis nebulosus 

Anolis sericeus 

Anolis tropidonotus 

Barisia gadovii levigata 

Barisia imbricata imbricata 
Barisia imbricata planifrons 
Barisia viridiflava 

Basiliscus vittatus 

Celestus enneagrammus 
Cnemidophorus deppii deppii 
Cnemidophorus guttatus guttatus 
Cnemidophorus guttatus immutabilis 


Cnemidophorus sackii australis 
Cnemidophorus sackii bocourti 
Coleonyx elegans elegans 
Coleonyx elegans nemoralis 
Corythophanes hernandezii 
Ctenosaura acanthura 
Ctenosaura pectinata 
Ctenosaura similis similis 
Enyaliosaurus quinquecarinatus 
Eumeces brevirostris 

Gaigeia dontomasi 

Gaigeia radula 

Gerrhonotus liocephalus liocephalus 
Gymnophthalmus sumichrasti 

| Heloderma horridum 

Iguana iguana rhinolopha 
Laemanctus deborrei 
Laemanctus longipes 
Laemanctus serratus 

Scincella assata taylori 
Scincella cherriei stuarti 
Scincella gemmingeri gemmingeri 
Scincella silvicola 
Lepidophyma smithii tehuanae 
Mabuya mabouya mabouya 
Phrynosoma asio 

Phrynosoma braconnieri 
Phyllodactylus magnus 
Phyllodactylus muralis 
Sceloporus aeneus bicanthalis 
Sceloporus edwardtaylori 
Sceloporus formosus formosus 
Sceloporus gadoviae 

Sceloporus grammicus grammicus 





CHECKLIST OF REPTILES OF MEXICO 


Sceloporus grammicus microlepidotus 
Sceloporus horridus horridus 
Sceloporus jalapae 

Sceloporus malachiticus salvini 
Sceloporus melanorhinus melanorhinus 
Sceloporus mucronatus omiltemanus 
Sceloporus siniferus cupreus 

Sceloporus siniferus siniferus 

Sceloporus spinosus caeruleopunctatus 
Sceloporus teapensis 


225 


Sceloporus variabilis smithi 
Sceloporus variabilis variabilis 
Sphaerodactylus glaucus glaucus 
Sphaerodactylus glaucus torquatus 
Urosaurus bicarinatus anonymorphus 
Urosaurus bicarinatus nelsoni 
Xenosaurus grandis 

Caiman crocodilus fuscus 

Crocodylus acutus acutus 


PUEBLA 


Kinosternon integrum 

Kinosternon leucostomum 

Abronia taeniata taeniata 

Abronia taeniata graminea 

Ameiva undulata amphigramma 
Ameiva undulata sinistra 

?Anolis cumingii 

Anolis nebuloides 

Anolis nebulosus 

Barisia imbricata imbricata 
Cnemidophorus deppii lineatissimus 
Cnemidophorus sackii sackii 
Cnemidophorus sackii communis 
Ctenosaura pectinata 

Eumeces brevirostris 

Eumeces copei 

Eumeces lynxe furcirostris 
Eumeces lynxe lynxe 

Gerrhonotus liocephalus liocephalus 
Gerrhonotus liocephalus ophiurus 


| Iguana iguana rhinolopha 
Phrynosoma braconnieri 
Phrynosoma orbiculare orbiculare 
Phrynosoma orbiculare cortezii 
Phrynosoma taurus 
Sceloporus aeneus aeneus 
Sceloporus aeneus bicanthalis 
Sceloporus formosus formosus 
Sceloporus gadoviae 
Sceloporus grammicus microlepidotus 
Sceloporus jalapae 
Sceloporus megalepidurus 
Sceloporus mucronatus aureolus 
Sceloporus mucronatus mucronatus 
Sceloporus pictus 
Sceloporus scalaris scalaris 
Sceloporus spinosus spinosus 
Sceloporus torquatus torquatus 
Sceloporus variabilis variabilis 
Urosaurus bicarinatus bicarinatus 


QUERETARO 


Cnemidophorus sackii communis 
Eumeces callicephalus 
Lepidophyma smithii occulor 
Phrynosoma orbiculare orbiculare 


Sceloporus jarrovii immucronatus 
Sceloporus jarrovii minor 
Sceloporus variabilis variabilis 


QUINTANA ROO® 


Chelonia mydas 

Eretmochelys imbricata 
Geoemyda areolata 

Kinosternon creaseri 
Kinosternon cruentatum consors 
Lepidochelys kempii 

Pseudemys scripta ornata 


Terrapene mexicana yucatana 
Ameiva undulata gaigeae 
Anolis cozumelae 

Anolis limifrons rodriguezii 
Anolis tropidonotus 
Aristelliger georgeensis 
Basiliscus vittatus 


° Morfin (Informe rendido por la comision geogréfico-exploradora de Quintana Roo al C. Secretario de 


Fomento, 1918, pp. 1-57, pls. 1-10) records, in a previously overlooked paper, a number of snakes from 
Quintana Roo. They are Boa imperator (= Constrictor constrictor imperator), Bothrops atriz (sic,=B. atror 
asper), Crotalus basiliscus (=C. d. durissus), and Elaps fulvius (=Micrurus affinis mayensis), all from 
“between Payo Obispo and Champoton.”’ 


226 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Cnemidophorus deppii cozumelus 
Cnemidophorus sackii angusticeps 
Coleonyx elegans elegans 
Corythophanes hernandezii 
Ctenosaura similis similis 


Iguana iguana rhinolopha 
Mabuya mabouya mabouya 
Sceloporus chrysostictus 
Sceloporus cozumelae 
Crocodylus acutus acutus 


SAN LUIS POTOSI 


Kinosternon hirtipes 

Kinosternon integrum 

Terrapene mexicana mexicana 
Ameiva undulata podarga 

Anolis petersi 

Anolis sericeus 

Anelytropsis papillosus 

Barisia imbricata, ciliaris 
Corythophanes hernandezii 
Cnemidophorus sackii communis 
Cnemidophorus sackii gularis 
Crotaphytus collaris baileyi 
Ctenosaura acanthura 

Eumeces lynxe lynxe 

Eumeces tetragrammus 
Gerrhonotus liocephalus infernalis 
Gerrhonotus liocephalus loweryi 
Holbrookia maculata approximans 
Holbrookia maculata dickersonae 
Holbrookia texana 


Laemanctus serratus 

Scincella caudaequinae 
Lepidophyma smithii occulor 
Phrynosoma cornutum 
Phrynosoma modestum 
Phrynosoma orbiculare orbiculare 
Sceloporus cautus 

Sceloporus goldmani 

Sceloporus grammicus disparilis 
Sceloporus jarrovii minor 
Sceloporus jarrovii immucronatus 
Sceloporus olivaceus 

Sceloporus parvus parvus 
Sceloporus parvus scutulatus 
Sceloporus serrifer plioporus 
Sceloporus spinosus spinosus 
Sceloporus torquatus melanogaster 
Sceloporus variabilis variabilis 
Xenosaurus newmanorum 


SINALOA 


Caretta caretta gigas 

Geoemyda pulcherrima pulcherrima 
Kinosternon hirtipes 
Kinosternon integrum 
Pseudemys scripta ornata 
Anolis nebulosus 

Anolis utowanae 

Barisia imbricata, ciliaris 
Callisaurus draconoides bogerti 
Callisaurus draconoides brevipes 
Cnemidophorus sackii communis 
Coleonyx fasciatus 

Ctenosaura hemilopha 
Ctenosaura pectinata 
Dipsosaurus dorsalis sonoriensis 
Kumeces humilis 


Eumeces parvulus 

Heloderma horridum 

Holbrookia maculata elegans 
Holbrookia maculata thermophila 
Iguana iguana rhinolopha 

Peropus mutilatus 

Phrynosoma solare 

Phyllodactylus lanei 

Sceloporus elarkii boulengeri 
Sceloporus horridus albiventris 
Sceloporus nelsoni 

Sceloporus utiformis 
?Sphaerodactylus glaucus torquatus 
Urosaurus bicarinatus tuberculatus 
Urosaurus ornatus schottii 


SONORA 


Chelonia mydas 

Dermochelys coriacea 
Eretmochelys imbricata 

Geoemyda pulcherrima pulcherrima 
Gopherus agassizii 


Kinosternon flavescens stejnegeri 
Kinosternon integrum 
Kinosternon sonoriense 
Lepidochelys olivacea 
Pseudemys scripta hiltoni 


CHECKLIST OF REPTILES OF MEXICO 


Terrapene klauberi 

Terrapene ornata 

Anolis nebuloides 

Callisaurus draconoides brevipes 
Callisaurus draconoides gabbii 
Callisaurus draconoides inusitatus 
Callisaurus draconoides ventralis 
Cnemidophorus bacatus 
Cnemidophorus burti 
Cnemidophorus gadovi 
Cnemidophorus sackii communis 
Cnemidophorus sackii stictogrammus 
Cnemidophorus tigris aethiops 
Cnemidophorus tigris gracilis 
Cnemidophorus tigris martyris 
Cnemidophorus tigris tigris 
Coleonyx variegatus variegatus 
Coleonyx variegatus sonoriensis 
Crotaphytus dickersonae 
Crotaphytus collaris baileyi 
Ctenosaura hemilopha 
Dipsosaurus dorsalis dorsalis 
Dipsosaurus dorsalis sonoriensis 
Elgaria kingii kingii 

Eumeces callicephalus 

Eumeces parviauriculatus 
Gambelia wislizenii wislizenii 
Heloderma horridum 

Heloderma suspectum 


227 


Holbrookia maculata approximans 
Holbrookia maculata thermophila 
Holbrookia texana 

Phrynosoma cornutum 
Phrynosoma ditmarsi 
Phrynosoma douglassii hernandesi 
Phrynosoma m’callii 

Phrynosoma modestum 
Phrynosoma platyrhinos goodei 
Phrynosoma solare 
Phyllodactylus homolepidurus 
Sauromalus obesus townsendi 
Sauromalus varius 

Sceloporus clarkii clarkii 
Sceloporus clarkii boulengeri 
Sceloporus jarrovii jarrovii 
Sceloporus magister magister 
Sceloporus nelsoni 

Sceloporus scalaris slevini 
Sceloporus undulatus virgatus 
Uma notata cowlesi 

Urosaurus bicarinatus tuberculatus 
Urosaurus ornatus linearis 
Urosaurus ornatus schottii 
Urosaurus ornatus symmetricus 
Uta nolascensis 

Uta palmeri 

Uta stansburiana stejnegeri 

Uta taylori 


TABASCO 


Claudius angustatus 
Dermatemys mawii 

Geoemyda areolata 

Kinosternon acutum 
Kinosternon cruentatum cruentatum 
Kinosternon leucostomum 
Pseudemys scripta ornata 
Staurotypus triporcatus 

Ameiva undulata amphigramma 
Ameiva undulata stuarti 

Anolis capito 

Anolis humilis uniformis 

Anolis laeviventris 

Anolis lemurinus bourgeaei 
Anolis sagrei mayensis 

Anolis sericeus 

Anolis tropidonotus 


Basiliscus vittatus 
Coleonyx elegans elegans 
Ctenosaura similis similis 
Eumeces schwarizei 
Hemidactylus turcicus turcicus 
Iguana iguana rhinolopha 
Laemanctus deborrei 
Scincella cherriei cherriei 
Lepidophyma flavimaculata flavimacu- 
lata 
Mabuya mabouya mabouya 
Sceloporus serrifer plioporus 
Sceloporus teapensis 
Sphaerodactylus glaucus glaucus 
Crocodylus acutus acutus 
Crocodvlus moreletii 


228 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


TAMAULIPAS 


Amyda emoryi 

Gopherus berlandieri 
Pseudemys scripta cataspila 
Pseudemys scripta elegans 
Terrapene mexicana mexicana 
Ameiva undulata podarga 
Anolis sericeus 

Basiliscus vittatus 
Cnemidophorus sackii communis 
Cnemidophorus sackii gularis 
Coleonyx brevis 

Crotaphytus reticulatus 
Ctenosaura acanthura 
Eumeces brevilineatus 
Eumeces dicei 


Eumeces obsoletus 

Eumeces tetragrammus 
Hemidactylus turcicus turcicus 
Holbrookia texana 

Laemanctus serratus 
Phrynosoma cornutum 
Sceloporus cyanogenys 
Sceloporus grammicus disparilis 
Sceloporus olivaceus 

Sceloporus serrifer piioporus 
Sceloporus spinosus spinosus 
Sceleporus variabilis marmoratus 
Sceloporus variabilis variabilis 
Crocodylus acutus acutus 
Crocodylus moreletii 


TLAXCALA 


Phrynosoma orbiculare orbiculare Sceloporus megalepidurus 


Sceloporus grammicus microlepidotus 


VERACRUZ 


Chelonia mydas 

Chelydra serpentina 

Claudius angustatus 
Dermatemys mawii 

Geomyda areolata 
Kinosternon acutum 
Kinosternon herrerai 
Kinosternon integrum 
Kinosternon leucostomum 
Pseudemys scripta cataspila 
Staurotypus triporcatus 
Terrapene mexicana mexicana 
Abronia taeniata graminea 
Abronia taeniata taeniata 
Ameiva undulata amphigramma 
Anelytropsis papillosus 

Anolis barkeri 

Anolis cymbops 

Anolis laeviventris 

Anolis lemurinus bourgeaei 
Anolis petersii 

Anolis sericeus 

Anolis tropidonotus 

Barisia antauges 

Barisia imbricata imbricata 
Barisia modesta 

Basiliscus vittatus 

Celestus enneagrammus 
Cnemidophorus deppii oligoporus 


Cnemidophorus guttatus guttatus 
Cnemidophorus sackii communis 
Cnemidophorus sackii gularis 
Coleonyx elegans elegans 
Corythophanes cristatus 
Corythophanes hernandezii 
Ctenosaura acanthura 
Ctenosaura similis similis 
Eumeces brevirostris 
Eumeces lynxe furcirostris 
Eumeces lynxe lynxe 
Eumeces sumichrasti 
Eumeces tetragrammus 
Gerrhonotus liocephalus ophiurus 
Hemidactylus mabouia 
Hemidactylus turcicus turcicus 
Holbrookia propinqua piperata 
Iguana iguana rhinolopha 
Laemanctus longipes 
Laemanctus serratus 
Scincella cherriei stuarti 
Scincella gemmingeri gemmingeri 
Scincella silvicola 
Lepidophyma flavimaculata flavimacu- 
lata 
Mabuya mabouya mabouya 
Phrynosoma orbiculare cortezii 
Sceloporus aeneus bicanthalis 
Sceloporus formosus formosus 


CHECKLIST OF REPTILES OF MEXICO 


Sceloporus grammicus microlepidotus 
Sceloporus jalapae 

Sceloporus jarrovii immucronatus 
Sceloporus malachiticus salvini 
Sceloporus megalepidurus 

Sceloporus mucronatus mucronatus 
Sceloporus mucronatus aureolus 
Sceloporus pictus 

Sceloporus serrifer plioporus 


229 


Sceloporus spinosus spinosus 
Sceloporus teapensis 

Sceloporus torquatus torquatus 
Sceloporus variabilis variabilis 
Sphaerodactylus glaucus glaucus 
Xenosaurus grandis 

Crocodylus acutus acutus 
Crocodylus moreletii 


YUCATAN 


Caretta caretta caretta 
Chelonia mydas 
Dermatemys mawii 
Geoemyda areolata 

Kinosternon creaseri 

Kinosternon cruentatum consors 
Pseudemys scripta ornata 
Terrapene mexicana yucatana 
Ameiva undulata gaigeae 

Anolis beckeri 

Anolis kidderi 

Anolis lemurinus bourgeaei 

Anolis limifrons rodriguezii 

Anolis sagrei mayensis 

Anolis tropidonotus 

Anolis ustus 

Basiliscus vittatus 
Cnemidophorus sackii angusticeps 


Coleonyx elegans elegans 
Corythophanes cristatus 
Corythophanes hernandezii 
Ctenosaura similis similis 
Enyaliosaurus defensor 
Kumeces schwartzei 
Hemidactylus turcicus turcicus 
Laemanctus alticoronatus 
Laemanctus serratus 

Scincella cherriei ixbaac 
Mabuya mabouya mabouya 
Sceloporus chrysostictus 
Sceloporus cozumelae 
Sceloporus lundelli gaigeae 
Sceloporus serrifer serrifer 
Sphaerodactylus glaucus glaucus 
Thecadactylus rapicaudus 


ZACATECAS 


Barisia imbricata ciliaris 
Cnemidophorus sackii communis 
Eumeces callicephalus 
Holbrookia maculata dickersonae 
Phrynosoma modestum 
Phrynosoma orbiculare orbiculare 
Sceloporus cautus 


Sceloporus grammicus disparilis 
Sceloporus jarrovii minor 
Sceloporus scalaris scalaris 
Sceloporus spinosus spinosus 
Sceloporus torquatus melanogaster 
Sceloporus undulatus consobrinus 






















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INDEX 


abaxillare, Kinosternon, 22, 218. 
about, Coleonyx variegatus, 42, 44, 
abnormis, Dermatemys, 19. 
Abronia, 194, 196. 
deppii, 196, 197, 221. 
fuscolabialis, 196, 198, 224. 
matudai, 196, 218. 
oaxacae, 196, 197, 224. 


ochoterenai, 196, 197, 218. 

taeniata graminea, 196, 198, 225, 
228. 

taeniata taeniata, 196, 197, 222, 
225, 228 


taeniatus, 197. 
acanthinus, Sceloporus, 108. 
Sceloporus malachiticus, 107, 108, 
218. 
acanthura, Ctenosaura, 73, 74, 224, 226, 
228. 


Lacerta, 74, 
acuta, Kinosternon scorpioides, 23. 
acutirostris, Anolis, 64. 
Testudo, 20. 
acutum, Kinosternon, 22, 23, 227, 228. 
acutus, Crocodylus, 910. 
Crocodylus acutus, 210, 218, 220, 
221, 223, 225, 226, 227, 228, 229. 
adspersus, Gerrhonotus, "201." 
aeneus, Sceloporus, 137. 
Seeloporus aeneus, 186, 137, 220, 
221, 222, 223, 225. 
ok Cnemidophorus _ tessellatus, 
189. 
Sento Beer tigris, 177, 189, 
227. 
Agama, collaris, 91. 
cornuta, 99. 
(Phrynosoma) coronata, 102. 
cristata, 68, 69. 
orbiculaire, 97. 
torquata, 126. 
undulata, 105. 
agassizii, Chelonia, 17. 
Gopherus, 28, 217, 226. 
Testudo, 28. 
Xerobates, 28. 
agilis, Scincus, 156. 
Agkistrodon, 3. 
Akleistops, 151. 
guatemalensis, 151, 152. 
albiventris, Sceloporus horridus, 
116; 222, 226. 
albogularis, Gonatodes, 45. 
Gymnodactylus, 45. 


110, 


alfaronis, Cnemidophorus, 178. 
alliacea, Mabuia, 156. 
Alligator chiapasius, 212. 

lacordairei, 211. 

mississippiensis, 211. 
Alligatoridae, 210, 211. 
alpha, Sceloporus grammicus, 120. 
altamirani, Humeces, 160, 161, 163, 223. 
alticoronatus, Laemanctus, 70, 229, 
amblygrammus, Eumeces’ skiltonianus, 

16 


Ambystomidae, 2. 
Ameiva, 170. 
ameiva, 170. 
edwardsii, 174. 
festiva edwardsii, 171, 174, 218. 
tesselata, 188. 
undulata amphigramma, 171, 172, 
224, 225, 227, 228. 
undulata dextra, 171, 173, 221, 224. 
undulata gaigeae, 171, 172, 218, 
225, 228. 
undulata hartwegi, 171, 218. 
undulata parva, 171, 173, 218, 224. 
undulata podarga, pia 172, 226, 


228. 

undulata sinistra, 171, 174, 220, 
222, 223, 225. 

undulata stuarti, 171, 173, 218, 224, 
227. 


undulata thomasi, 171, 173, 218. 
undulata undulata, 171, 174, 224. 
ameiva, Ameiva, 170. 
americana, Lacerta, 170. 
americanus, Basiliscus, ales 
Crocodylus, 210, 211. 
Amoebopsis, 154. 
amphigramma, Ameiva undulata, 171, 
172, 224, 225, 227, 228. 
Amphisbaenia, 2, 12. 37. 
Amyda, 18. 
cartilaginea, 18. 
emoryi, 18, 217, 219, 224, 228. 
javanica, 18. 
spinifera, 18. 
Anapsida, 12. 
Ancistrodon, 3. 
Anelytropsidae, 39, 170. 
Anelytropsis, 170. 
papillosus, 170, 226, 228. 
Aneuporus occipitalis, 213. 
Anguidae, 39, 40, 194. 
Anguis, 194. 
angustatus, Claudius, 26, 218, 227, 228. 
angusticeps, Cnemidophorus, ‘183. 


231 


232 


BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


angusticeps, Cnemidophorus sackii, 178, | Anolis sagraei, 55. 


183, 218, 226, 229. 
angustus, Sator, 139, 217. 
Aniellidae, 208. 
annectens, Ctenosaura (Cachryx), 77. 
Anniella, 208, 209. 
geronimensis, 209, 217. 
pulchra, 209. 
pulchra pulchra, 209, 217. 
texana, 209. 
Anniellidae, 39, 208. 
annulata, Geoclemmys, 29. 
annulatus, Sceloporus merriami, 
133, 219. 
Anolis, 4, 54, 55. 
acutirostris, 64. 
aureolus, 64. 
baccatus, 57, 62. 
barkeri, 56, 58, 228. 
beckeri, 57, 62, 229. 
biporcatus, 58, 65, 67, 218. 
boulengerianus, 65. 
bourgeaei, 66, 67. 
bullaris, 56. 
capito, 58, 65, 227. 
carneus, 65. 
carolinensis, 55, 56. 
cobanensis, 63. 
copei, 65. 
cozumelae, 56, 59, 225. 
cumingii, 58, 67, 225. 
cymbops, 57, 58, 62, 228. 
damulus, 57, 61. 
dunni, 56, 61, 221. 
fuscoauratus, 55. 
gadovii, 56, 61, 221. 
guentherii, 64. 
guntherii, 57, 63. 
heliactin, 58, 67, 224. 
humilis uniformis, 56, 60, 218, 227. 
impetigosus, 56, 60. 
intermedius, 63. 
jacobi, 68. 
kidderi, 58, 66, 218, 229. 
laeviventris, 9, 57, 58, 62, 63, 227, 228. 
lemurinus bourgeaei, 58, 66, 218, 
227, 228, 229. 
limifrons rodriguezii, 56, 57, 64, 
218, 224, 225, 229. 
liogaster, 57, 62, 221. 
megapholidotus, 56, 59, 221. 
metallicus, 56, 59. 
milleri, 56, 57, 64, 224. 
nannodes, 63. 
nebuloides, 58, 66, 219, 220, 221, 
224, 225, 227. 
nebulosus, 58, 65, 220, 221, 222, 
2238, 224, 225, 226. 
panamensis, 61. 
pentaprion 55, 57, 61, 218. 
(Coecoéssus) pentaprion, 61. 
petersii, 58, 65, 226, 228. 
petersii bivittatus, 65. 
rodriguezii, 64. 
rubigenosus, 64. 
ruthveni, 60. 


132 


? 


sagrei mayensis, 56, 59, 218, 227, 
229. 
sallaei, 67. 
schiedii, 56, 57, 58, 63, 64. 
schmidti, 56, 58, 60, 220. 
pericels, 58, 67, 218, 224, 226, 227, 
8. 


stuarti, 63. 
sulcifrons, 61. 
taylori, 57, 61, 221. 
tropidonotus, 56, 59, 60, 218, 224, 
225, 227, 228, 229. 
uniformis, 59, 60. 
ustus, 58, 66, 218, 229. 
ustus veraepacis, 67. 
utowanae, 57, 62, 226. 
viridis, 55. 
wiegmanni, 63. 
anonymorpha, Uta, 146. 
Uta bicarinata, 146. 
anonymorphus, Urosaurus bicarinatus, 
141, 146, 218, 221, 225. 
Anota, 95. 
calidiarum, 101. 
goodei, 102. 
m’callii, 95, 99. 
modesta, 101. 
platyrhina, 101. 
antauges, Barisia, 199, 200, 228. 
Gerrhonotus, 200. 
anthracinus, Sphaerodactylus, 52, 213. 
approximans, Holbrookia, 83. 
Holbrookia maculata, 82, 83, 219, 
220, 222, 226, 227. 
Archosauria, 12, 209. 
areolata, Emys, 30. 
Geoemyda, 29, 30, 218, 225, 227 
228, 229. 
Aristelliger, 40, 41, 51. 
georgeensis, 51, 225. 
irregularis, 51. 
larioll: 
praesignis, 51. 
arizonae, Cnemidophorus, 184. 
armata, Iguana (Ctenosaura), 74. 
articulata, Cyclura, 74. 
asio, Batrachosoma, 102. 
Phrynosoma, 95, 102, 218, 220, 
221, 2238, 224. 
asper, Sceloporus, 107, 109, 221, 222, 
223. 


, 


Aspidonectes emoryi, 18. 
Aspidoscelis, 174. 
assata, Scincella assata, 157, 160, 218. 
assatum, Leiolopisma assatum, 160. 
Lygosoma assatum, 160. 
assatus, Lampropholis, 160. 
ater, Sauromalus, 79, 80, 217. 
Atheca, 12, 13. 
atitlanensis, Celestus, 195. 
atrix, Bothrops, 225. 
aureolus, Anolis, 64. 
Sceloporus mucronatus, 123, 124, 
225, 229. 
auriculata, Uta, 146. 


INDEX 


auriculatus, Urosaurus, 140, 146, 220. 
australis, Cnemidophorus communis, 
181 


Cnemidophorus sackii, 178, 181, 224. 
Sauromalus, 79, 80, 217. 
austrinus, Gerrhonotus liocephalus, 203, 
204, 218. 


bacatus, Cnemidophorus, 176, 187, 
baccatus, Anolis, 57, 62. 
Bachia dorbignyi, 213. 
heteropa, 212. 
baileyi, Crotaphytus, 92. 
Crotaphytus collaris, 92, 217, 
220, 224, 226, 227. 
bakeri, Ctenosaura, 73. 
balsas, Cnemidophorus mexicanus, 
Barisia, 194, 198. 
antauges, 199, 200, 228. 
gadovii gadovii, 199, 200, 221. 
gadovii levigata, 199, 200, 224. 
imbricata ciliaris, 199, 202, 
220, 221, 222, 224, 226, 229. 
imbricata imbricata, 198, 199, 201, 
220, 221, 222, 223, 224, 225, 228. 
imbricata planifrons, 199, 201, 224. 
levicollis, 199, 202, 219. 
modesta, 199, 201, 228. 
moreleti rafaeli, 198, 199, 218. 
moreleti temporalis, 199, 218. 
rudicollis, 199, 202, 222. 
viridiflava, 198, 200, 224. 
barkeri, Anolis, 56, 58, 228. 
bartolomas, Cnemidophorus, 191. 
Basiliscus, 54, 71. 
americanus, 71. 
basiliscus, 71. 
(Cristasaura) nuchalis, 72. 
vittatus, 71, 218, 220, 221, 222, 223, 
224, 225, 227, 228, 229. 
basiliscus, Basiliscus, 71. 
Crotalus, 225. 
Batrachosoma, 94. 
asio, 102. 
beckeri, Anolis, 57, 62, 229. 
beldingi, Cnemidophorus hyperythrus, 
175, 186, 217. 
Verticaria, 186. 
Verticaria hyperythra, 186. 
belli, Leiosaurus, 212. 
bellii, Chrysemys picta, 34, 219. 
Emys, 34. 
Iguana (Ctenosaura), 74. 
Plestiodon, 163. 
berardii, Emys, 19. 
berendtianum, Cinosternum, 23. 
berlandieri, Gopherus, 28, 219, 224. 
Testudo, 28. 
Xerobates, 28. 
bicanthalis, Sceloporus aeneus, 136, 137, 
222, 224, 225, 228. 
bicarinata, Uta, 146. 
bi-carinata, Uta bi-carinata, 146. 
bicarinatus, Phymatolepis, 140. 
Urosaurus bicarinatus, 141, 
221, 223, 225. 


227, 


219, 


181. 


219, 


146, 


233 


bi-carinatus, Phymatolepis, 146. 
Bimanus, 38. 
binocularis, Sceloporus, 126. 
Sceloporus ferrariperezi, 126. 
Sceloporus torquatus, 122, 126, 224. 
Bipedidae, 37. 
Bipes, 37, 38. 
biporus, 38, 217. 
canaliculatus, 38, 221. 
tridactylus, 38, 221. 
biporeata, Dactyloa, 55, 65. 
biporeatus, Anolis, 58, 65, 67, 218. 
Staurotypus, 27. 
biporus, Bipes, 38, 217. 
Euchirotes, 38. 
bi-seriatus, Sceloporus, 118. 
biseriatus, Sceloporus occidentalis, 117, 
TISP2I7: 
bischoffi, Holbrookia, 84. 
biscutatus, Crocodilus, 211. 
bissa, Caretta, 16. 
bivittata, Daconura, 71, 72. 
bivittatus, Anolis petersii, 65. 
blainvillii, Phrynosoma, 103. 
Phrynosoma coronatum, 97, 103, 217. 
Blepharactisis, 192. 
speciosa, 192. 
Boa imperator, 225. 
bocourti, Cnemidophorous, 181. 
Cnemidophorus communis, 181. 


Cnemidophorus sackii, 178, 181, 
218, 224. 
Gerrhonotus, 200. 
Tropidurus, 213. 
bocourtii, Eumeces, 166. 
Callisaurus draconoides, 86, 


bogerti, 
89, 
Coleonyx variegatus, 42, 44. 
bordai, Phyllodactylus, 46, 47, 221. 
Bothrops atrix, 225. 
boucardi, Phrynosoma, 96, 98, 222. 
boucardii, Phrynosoma, 98. 
Tapaya, 98. 
boulengeri, Sceloporus, 113. 
Sceloporus clarkii, 111, 113, 222, 
223, 226, 227. 
boulengerianus, Anolis, 65. 
bourgeaei, Anolis, 66, 67. 
Anolis lemurinus, 58, 66, 218, 227, 
228, 229. 
brachycercum, Phrynosoma douglassii, 
96, 100, 220. 
Brachydactylus, 41. 
mitratus, 41. 
brachylopha, Ctenosaura teres, 75. 
braconnieri, Goniodactylus, 45. 
Phrynosoma, 95, 100, 224, 225. 
brevigulare, Cinosternum, 25. 
brevilineatus, Eumeces, 162, 164, 224, 228. 
brevipes, Callisaurus draconoides, 86, 
88, 226, 227. 
brevirostris, Ctenosaura, 75. 
Eumeces, 162, 168, 220, 221, 222, 
223, 224, 225, 228. 
Mabouia, 168. 
brevis, Coleonyx, 42, 43, 219, 220, 224, 228, 


234 


bufonium, Phrynosoma, 99. 
bullaris, Anolis, 56. 

Dactyloa, 55. 
bulleri, Sceloporus, 122, 125, 222. 
bunkeri, Holbrookia, 82, 84, 219. 
burti, Cnemidophorus, 177, 185, 227. 


Cachryx, 76, 77. 
defensor, 77. 
erythromelas, 77. 
caerulea, Uta, 143. 
Verticaria, 186. 
caeruleopunctatus, Sceloporus spinosus, 
110, 116, 225. 
caeruleus, Cnemidophorus hyperythrus, 
175, 186, 217. 
Sceloporus ornatus, 122, 127, 219. 
Urosaurus ornatus, 141, 148, 219. 
Caiman, 212. 
crocodilus fuscus, 212, 218. 
fissipes, 212. 
calidiarum, Anota, 101. 
Phrynosoma, 101. 
callicephalus, Eumeces, 161, 164, 219, 
220, 221, 222, 223, 225, 227, 229. 
calligaster, Sceloporus melanorhinus, 
110, 113, 220, 221, 222, 223. 
eallirostris, Emys, 32. 
Callisaurus, 55, 85. 
carmenensis, 87. 
crinitus, 87. 
draconoides, 85, 86. 
draconoides bogerti, 86, 89, 226. 
dreeonvldes brevipes, 86, 88, 226, 
ceopotien carmenensis, 86, 87, 
draconoides crinitus, 86, 87, 217. 
draconoides draconoides, 86, 217. 
draconoides gabbii, 86, 87, 217, 227. 
draconoides inusitatus, 86, 88, 227. 
draconoides splendidus, 86, 87, 217. 
draconoides ventralis, 86, 88, 227. 
inusitatus, 88. 
rhodostictus, 87. 
splendidus, 87. 
ventralis, 88. 
ventralis gabbii, 87. 
ventralis inusitatus, 88. 
ventralis ventralis, 88. 
canaliculatus, Bipes, 38, 221. 
Chirotes, 39. 
canus, Cnemidophorus, 190. 
Cnemidophorus tessellatus, 190. 
Cnemidophorus tigris, 176, 190, 217. 
Caouana, 14, 15. 
caouana, Testudo, 15, 16. 
capito, Anolis, 58, 65, 227. 
Caretta, 14, 15. 
bissa, 16. 
caretta, 16. 
caretta caretta, 15, 16, 229. 
caretta gigas, 15, 16, 217, 226. 
gigas, 16. 


BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Caretta nasuta, 15. 
remivaga, 15. 
caretta, Caretta, 16. 
Caretta caretta, 15, 16, 229. 
Testudo, 16. 
Carettochelyidae, 18. 
Carettoidea, 12, 14. 
carinatus, Sceloporus, 134, 135, 218. 
carmenensis, Callisaurus, 87. 
Callisaurus draconoides, 86, 87, 217. 
Dipsosaurus, 78, 217. 
carneus, Anolis, 65. 
carolina, Cistudo, 34. 
Terrapene, 34. 
carolinensis, Anolis, 55, 56. 
cartilaginea, Amyda, 18. 
caspica, Clemmys caspica, 33. 
amas Cnemidophorus, 176, 188, 
Di 
Dipsosaurus, 78, 217. 
cataspila, Emys, 32. 
Pseudemys scripta, 31, 32, 228. 
caudaequinae, Leiolopisma, 158. 
Scincella, 157, 158, 224, 226. 
Caudata, 2. 
cautus, Sceloporus, 117, 219, 226, 229. 
cedrosensis, Elgaria, 205, 206, 217. 
Gerrhonotus, 206. 
celeripes, Cnemidophorus, 190. 
Cnemidophorus tessellatus, 190. 
Cnemidophorus tigris, 176, 190, 217. 
Celestus, 194. 
atitlanensis, 195. 
enneagrammus, 195, 224, 228. 
rozellae, 195, 218. 
striatus, 194. 
cephalo, Testudo, 16. 
ceralbensis, Verticaria, 188. 
Cnemidophorus, 175, 188, 217. 
cerroense, Phrynosoma, 97, 103, 217. 
Chalcides heteropus, 212. 
suleata, 39. 
Chalinocnemis, 51. 
chalybaeus, Diploglossus, 196. 
Chamaeleo mexicanus, 68. 
Chamaeleopsis, 68. 
hernandezii, 68. 
chamaeleopsis, Corythophanes, 68. 
Chamaesaura propus, 39. 
Chelidridae, 20. 
Cheliurus, 20. 
Chelona, 17. 
Chelone, 17. 
Chelonia, 14, 17. 
agassizii, 17. 
dussumieri, 15. 
lachrymata, 16. 
maculosa, 17. 
mydas, 17, 221, 228, 224, 225, 226, 
228, 229. 
olivacea, 14, 15. 
pseudo-caretta, 16. 
pseudo-mydas, 16. 
virgata, 17. 


Chelonias, 17. 

Cheloniidae, 14. 

Chelonura, 20. 

Chelopus rubida, 30. 

Chelydra, 20. 
rossignonii, 20. 
serpentina, 20, 21, 218, 228. 

Chelydridae 19, 20. 

cherriei, Leiolopisma, 158. 
Leiolopisma cherriei, 157. 
Lygosoma assatum, 157, 
Lygosoma cherriei, 157. 
Mocoa, 157. 


INDEX 2335 


Clemmys, 29, 37. 
caspica caspica, 33. 
guttata, 37. 
marmorata pallida, 37, 217. 
Cnemaspis, 45. 
Cnemidophorus, 1, 4, 170, 174. 
alfaronis, 178. 
angusticeps, 183. 
arizonae, 184. 
bacatus, 176, 187, 227. 
bartolomas, 191. 
bocourti, 181. 
burti, 177, 185, 227. 


Scincella cherriei, 157, 218, 227. canus, 190. 


chiapasius, Alligator, 212. 
ee Urosaurus ornatus, 
i143. 
Uta ornata, 143. 
Chirotes canaliculatus, 39. 
Chloremys, 19. 
Chrysemys, 29, 33. 
nebulosa, 32. 
picta bellii, 34, 219. 
scripta elegans, 32. 


catalinensis, 176, 188, 217. 
142, celeripes, 190. 

ceralbensis, 175, 188, 217. 

communis, 182. 

communis australis, 181. 

communis bocourti, 181. 

communis copei, 181, 182. 

communis occidentalis, 182. 

costatus, 181. 

decemlineatus, 178. 


chrysostictus, Sceloporus, 105, 133, 218, deppei cozumela, 179. 
226, 229. deppii, 178. 
ciliaris, Barisia imbricata, 199, 202, 219, deppii cozumelus, 176, 179, 226. 


220, 221, 222, 224, 226, 229. 

Gerrhonotus levicollis, 202. 
cinereus, Sphaerodactylus, 52. 
Cinosternidae, 21. 
Cinosternon, 3. 

cruentatum, 23. 

effeldtii, 23. 

hirtipes, 25. 

leucostomum, 25. 

punctatum, 26. 

shawianum, 23. 
Cinosternum berendtianum, 23, 

brevigulare, 25. 

cobanum, 25. 

henrici, 25. 

hirtipes, 25. 

integrum, 25. 

rostellum, 25. 

scorpioides integrum, 25. 

sonoriense, 26. 

triliratum, 23. 
Cistudo carolina, 34. 

mexicana, 34. 

(Onychotria) mexicana, 34. 

ornata, 36. 

yucatana, 35. 


clarionensis, Urosaurus, 141, 142, 220. 


Uta, 142. 
clarki, Ctenosaura, 76. 
Enyaliosaurus, 76, 223. 
clarkii, Sceloporus, 113. 


Sceloporus clarkii, 111, 113, 219, 


227. 
Claudius, 21, 26. 
angustatus, 26, 218, 227, 228. 
megalocephalus, 26. 
pictus, 27. 
severus, 27. 
clausa, Testudo, 34. 


deppii deppii, 176, 178, 218, 221, 
294, 


deppii lineatissimus 176, 179, 220, 
221, 222, 223, 225. 

deppii oligoporus, 176, 179, 218, 
228. 


dickersonae, 191. 

disparilis, 189. 

estebanensis, 189. 

gadovi, 178, 185, 227. 

gracilis, 189. 

grahamii, 188. 

gularis, 183. 

gularis gularis, 183. 

gularis gularis obsoletus, 182. 

gularis gularis verus, 182. 

gularis meeki, 184 

gularis octolineatus, 183. 

gularis scalaris, 182. 

gularis sealous, 183. 

gularis semifasciatus, 184. 

gularis sericeus, 183. 

gularis velox, 184. 

guttatus, 183. 

guttatus guttatus, 175, 179, 224, 
228. 

guttatus immutabilis, 175, 180, 218, 
220, 221, 223, 224. 

guttatus striatus, 180. 

hyperythrus, 174. 

hyperythrus beldingi, 175, 186, 217. 

hyperythrus caeruleus, 175, 186, 
217. 

hypersebaus danheimae, 175, 186, 
217. 


hyperythrus hyperythrus, 175, 187, 
217. 
hyperythrus pictus, 175, 186, 217. 


236 


Cnemidophorus hyperythrus schmidti, 


175, 187, 217. 
immutabilis, 180. 
inornatus, 177 184, 219, 224. 
labialis, 178, 185, 217, 
lativittis, 178. 
lineatissimus, 179. 
mariarum, 183. 
marmoratus, 190. 
martyris, 191. 
maximus, 176, 188, 217. 
melanostethus, 189, 190. 
mexicanus, 181. 
mexicanus balsas, 181. 
mexicanus typica, 181. 
microlepidopus, 180. 
motaguae, 181. 
octolineatus, 184. 
perplexus, 184, 185. 
punctilineatus, 189. 
rubidus, 191. 
sackii, 181. 
sackii angusticeps, 178, 183, 218, 

226, 229. 
sackii australis, 178, 181, 224. 
sackii bocourti, 


228) 229. 


sackii gularis, 178, 183, 224, 226, 


228 
sackii mariarum, 178, 183, 223. 
sackii sackii, 178, 180, 221, 223, 225. 
sackii scalaris, 178, 182, 219, 220. 
sackii semifasciatus, 178, 184, 219. 


nae stictogrammus, 178, 183, 219, 
227. 


septemvittatus, 184. 
sexlineatus, 177, 185. 
sexlineatus sackii, 180, 181. 
sexlineatus tigris, 182. 
stejnegeri, 191. 
tesselatus, 3, 176, 188. 
tessellatus, 3. 
tessellatus aethiops, 189. 
tessellatus canus, 190. 
tessellatus celeripes, 190. 
tessellatus martyris, 191. 
tessellatus multiscutatus, 191. 
tessellatus rubidus, 191. 
tessellatus stejnegeri, 191. 
tessellatus tessellatus, 189. 
tigris aethiops, 177, 189, 227. 
tigris canus, 176, 190, 217. 
tigris celeripes, 176, 190, 217. 
tigris gracilis, 777 189, 227, 
tigris marmoratus, 
220. 


tigris martyris, 176, 191, 227. 
tigris multiscutatus, 177, 191, 217. 
tigris rubidus, 176, 191. 
tigris tigris, 177, 189, 217, 227. 
undulatus, 174. 
unicolor, 180. 

Cnemidophorus vandenburghi, 191. 
variolosus, 190. 


178, 181, 218, 224, 
sackii communis, 178, 182, 217, 219, 
220, 221, 222, 223, 225, 226, 227, 


177, 190, 219, 


BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


coahuila, Terrapene, 34, 36, 219. 
cobanensis, Anolis, 63. 
cobanum, Cinosternum, 25. 
Coccoéssus, 55, 61. 
cochranae, ‘Seeloporus, 134. 
Coleonyx, "41, 42. 
brevis, 42, 43, 219, 220, 224, 228. 
elegans, Al, 42, 43. 
elegans elegans, 42, 218, 224, 226, 
227, 228, 229. 
elegans nemoralis, 42, 48, 220, 221, 
223, 224. 
fasciatus, 42, 43, 220, 226. 
variegatus, 44. 
variegatus abbotti, 42, 44, 217. 
variegatus bogerti, 42, 44. 
variegatus peninsularis, 42, 44, 217. 
variegatus slevini, 42, 45, 217. 
variegatus sonoriensis, 42, 45, 227. 
varoeue variegatus, 42, 44, 217, 
22 


coleonyx, Gymnodactylus, 42. 
colimensis, Eumeces, 162, 169, 220. 
collaris, Agama, 91. 
Crotaphytus, 92. 
Crotaphytus collaris, 92. 
Colpochelys, 15. 
communis, Cnemidophorus, 182. 
Cnemidophorus sackii, 178, 182, 
QUT 2192) 2205 e220 222.0293. 
225, 226, 227, 228, 229: 
completa, Ctenosaura, 73. 
concentrica, Testudo, 36. 
concinna, Testudo, 31. 
Uta, 148, 150, 217. 
consobrinus, Sceloporus, 118. 
Sceloporus undulatis, 117, 118, 219, 
220, 224, 229. 
consors, Kinosternon cruentatum, 22, 
24, 225, 229. 
conspicuosa, Ctenosaura, 75. 
Constrictor constrictor imperator, 225. 
durissus durissus, 225. 
copei, Anolis, 65. 
Cnemidophorus communis, 181, 182. 
Eumeces, 162, 166, 220, 222, 223, 
225. 
copeii, Crotaphytus, 94. 
Cophosaurus, 81. 
texana, 81. 
texanus, 85. 
Cordylus (Gerrhonotus) multicarinatus, 
205. 
coriacea, Dermochelys, 13, 217, 226. 
Testudo, 13. 
cornuta, Agama, 99. 
Tapaya, 99. 
cornutum, Phrynosoma, 94, 95, 99, 219, 
220, 224, 226, 227. 
coronata, Agama (Phrynosoma), 102. 
coronatum, Phrynosoma, 94, 96. 
Phrynosoma coronatum, 97, 102, 
217. 
cortezii, Phrynosoma orbiculare, 96, 98, 
225, 228. 
Tapaya, 98. 


INDEX 


Corythaeolus, 71. 
Corythophanes, 3, 54, 68. 
chamaeleopsis, 68. 
cristatus, 68, 69, 218, 228, 229. 
ee odeet 68, 218, 224, 226, 228, 
229. 


mexicanus, 68. 
percarinatus, 68, 69, 218. 
Corytophanes, 3, 68, 69. 
costatus, Cnemidophorus, 181. 
couchii, Sceloporus, 105, 129, 132, 219, 
224 


cowlesi, Uma notata, 89, 90, 227. 
cozumela, Cnemidophorus deppei, 179. 
cozumelae, Anolis, 56, 59, 225. 
Sceloporus, 129, 130, 226, 229. 
cozumelus, Cnemidophorus deppii, 176, 
179, 226. 
creaseri, Kinosternon, 22, 23, 225, 229. 
Cricosaura, 151. 
crinitus, Callisaurus, 87. 
Callisaurus draconoides, 86, 87, 217. 
Cristasaura, 71, 72. 
mitrella, 71. 
cristata, Agama, 68, 69. 
cristatus, Corythophanes, 68, 69, 218, 
228, 229. 


Crocodilidae, 210. 
Crocodilus biscutatus, 211. 
latirostris, 212. 
mexicanus, 211. 
pacificus, 211. 
Crocodylidae, 210. 
Crocodylus, 210. 
acutus, 210. 
acutus acutus, 210, 218, 220, 221, 
223, 225, 226, 227, 228, 229. 
americanus, 210, 211. 
moreletii, 210, 211, 218, 220, 227, 
228, 229. 
niloticus, 210. 


Crotalus basiliscus, 225. 
polystictus, 6. 
Crotaphytus, 55, 91. 
baileyi, 92. 
collaris, 92. 
collaris baileyi, 92, 217, 219, 220, 
224, 226, 227. 
collaris collaris, 92. 
collaris dickersonae, 93. 
copeii, 94. 
dickersonae, 92, 93, 227. 
dorsalis, 77, 78. 
fasciatus, 94. 
fasciolatus, 94. 
gambelii, 94. 
insularis, 92, 93, 217. 
reticulatus, 92, 93, 219, 224, 228. 
wislizenii, 94. 
cruentatum, Cinosternon, 23. 
Kinosternon cruentatum, 22, 23, 
218, 224, 227. 


Cryptoblepharus, 155. 
Cryptodira, 12, 18, 19. 
Ctenocercus, 55. 


861316—s0 17 





237 


Ctenosaura, 54, 73, 74. 
acanthura, 73, 74, 224, 226, 228. 
(Cachryx) annectens, 77. 
bakeri, 73. 
brevirostris, 75. 
clarki, 76. 
completa, 73. 
conspicuosa, 75. 
eycluroides, 73, 74. 
defensor, 77. 
erythromelas, 76, 77. 
hemilopha, 73, 75, 217, 219, 226, 
227. 
insulana, 75. 
interrupta, 75. 
multispinis, 74. 
parkeri, 75. 
pectinata, 73, 75, 220, 221, 
223, 224, 225, 226. 
similis, 73, 218. 
similis similis, 218, 224, 226, 227, 
228, 229. 
teres brachylopha, 75. 
Cubina grandis, 208. 
cumingii, Anolis, 58, 67, 225. 
cupreus, Sceloporus, 134. 
Sceloporus siniferus, 184, 225. 
pyeMee eure! Sceloporus, 1238, 125, 224, 
8. 


222, 


Sceloporus torquatus, 125. 
Cyclura articulata, 74. 
denticulata, 74. 
palearis, 76. 
pectinata, 75. 
quinquecarinata, 76. 
semicristata, 74. 
shawii, 74. 
(Ctenosaura) similis, 73. 
teres, 74. 
cycluroides, Ctenosaura, 73, 74. 


cymbops, Anolis, 56, 57, 58, 62, 228. 


Daconura bivittata, 71, 72. 


Dactyloa, 55. 
biporcata, 55, 65. 
bullaris, 55. 
(Anolis) laeviventris, 62. 
nebulosa, 55, 65. 
schiedii, 63. 
Dactyloperus, 51. 
damulus, Anolis, 57, 61. 


danheimae, Cnemidophorus hyperyth- 
rus, 175, 186, 217. 


Dasia microcephalus, 214. 
deborrei, Laemanctus, 70, 71, 224, 227. 
decemlineatus, Cnemidophorus, 178. 


defensor, Cachryx, 77. 
Ctenosaura, 77. 
Enyaliosaurus, 76, 77, 229. 


deleampi, Phyllodactylus, 46, 47, 221. 
delicatissima, Iguana, 72. 
delicatissimus, Sceloporus, 131. 


238 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


denticulata, Cyclura, 74. dugesii, Eumeces, 161, 169, 221, 222, 
deppii, Abronia, 196, 197, 221. 223. 
Cnemidophorus, 178. Phrynosoma orbiculare, 97, 98, 220. 
Cnemidophorus deppii, 176, 178, See 127. 
218, 221, 224. celoporus dugesii, 122, 127, 220 
Gerrhonotus, 196, 197. if 222, 223. ghtned : 
Dermatemyidae, 19. apaya, 98. 
Dermatemys, 19. dunni, Anolis, 56, 61, 221. 
abnormis, 19. durissus, Constrictor duri 225 
mawii, 19, 218, 224, 227, 228, 229. Ur geen RO eee 
Dermochelyidae, 13. dussumieri, Chelonia, 15. 
Dermochelys, 13. 
coriacea, 13, 217, 226. Eephymotes obtusirostris, 213. 
* ego 13. edwardsii, Ameiva, 174. 
evisia, 20. Ameiva festiva, 171, 174, 218. 
or Ameiva undulata, 171, 173, 221, | edwardtaylori, Sceloporus, 110, 112, 224. 
dicei, Eumeces, 162, 169, 224, 228. effeldtii, Cinosternon, 23. 
dickersonae, Cnemidophorus, 191. Elaps fulvius, 225. , 
Crotaphytus, 92, 93, 227. elegans, Chrysemys scripta, 32. 
Crotaphytus collaris, 93. Coleonyx, 41, 42, 43. 
Holbrookia, 83. Coleonyx elegans, 42, 218, 224, 226, 
Holbrookia maculata, 82, 83, 219, 227, 228, 229. 
220, 221, 226, 229. Emys, 32. 
digueti, Gerrhonotus deppii, 198. Holbrookia, 84. 
Sceloporus, 114. Holbrookia elegans, 84. 
Diplodactylus, 47. oan maculata, 82, 84, 226. 
unctus, 47. seudemys, 32. 


. Pseudemys scripta, 31, 32, 219, 228. 
Diploglossus, 39. Pseudemys troostii, 32. 
chalybaeus, 196. Uta, 148, 149. 
naler cep alae Uta stansburiana, 148, 150, 217. 
MCT OSEODIS aa Elgaria, 194, 205. 


steindachneri, 195. : 
Dipsesnurussaur7: cedrosensis, 205, 206, 217. 


: kingii, 205. 
carmenensis, 78, 217. Be es 
catalinensis, 78, 217. kingii kingii, 205, 219, 227. 
dorsalis, 78. kingii nobilis, 205, 206. 
dorsalis dorsalis, 77, 78, 217, 227. multicarinata multicarinata, 205. 
dorsalis lucasensis, Ceoweenlide multicarinata nana, 205, 207, 217. 
dorsalis sonoriensis, 77, 79, 226, 227. multicarinata webbii, 205, 207, 217. 
Discodactylus, 46. nobilis, 206. 
dispar, Sceloporus, 120. paucicarinatus, 205, 206, 217. 
Ee ean a ea emoryi, Amyda, 18; 219, 224, 228, 
Sceloporus grammicus, 119, 120, 3 ee See e een Se 
219, 220, 221, 222, 224° 296, | Emyidae, 19, 28, 29. 
228, 229. Emys areolata, 30. 
Sceloporus microlepidotus, 120. bellii, 34. 
ditmarsi, Phrynosoma, 95, 104, 227. pertain ae 
Doliosaurus, 95. cataapila, (2 ; 
m’calli, 99. elegans 39 : 
modestus, 101. eS Fe 
platyrhinos, 101. grayl, 33. 
dontomasi, Gaigeia, 153, 224. incisa, 30. 
dorbignyi, Bachia, 213. ornata, 31. 
dorsalis, Crotaphytus, 77, 78. pulcherrima, 30. 
Dipsosaurus, 78. umbra, 33. 
Dipsosaurus dorsalis, 77, 78, 217, venusta, 31. 
a Emysaurus, 20. 
douglassii, Phrynosoma, 96, 100. rossignonii, 20. 
draconoides, Callisaurus, 85, 86. enneagrammus, Celestus, 195, 224, 228. 


Callisaurus draconoides 86, 217. Siderolamprus, 194. 


INDEX 


Enysliosaurus, 54, 75. 

clarki, 76, 223. 

defensor, 76, 77, 229. 

erythromelas, 76, 77, 218. 

quinquecarinatus, 76, 224. 
Epaphelus, 192. 

sumichrasti, 192. 
Eremonia, 15. 
Eretmochelys, 14, 16. 

imbricata, 16, 217, 218, 224, 225, 

226. 

squamata, 17. 
erythromelas, Cachryx, 77. 

Ctenosaura, 76, 77. 

Enyaliosaurus, 76, 77, 218. 


espiritensis, Verticaria, 187. 
estebanensis, Cnemidophorus, 189. 


Eublepharis fasciatus, 43. 
variegatus, 44. 


Euchelonia, 17. 


Euchirotes, 38. 
biporus, 38. 
Eumeces, 155, 160. 
altamirani, 160, 161, 163, 223. 
bocourtii, 166. 
brevilineatus, 162, 164, 224, 228. 
brevirostris, 162, 168, 220, 221, 222, 
223, 224, 225, 228. 
callicephalus, 161, 164, 219, 220, 
221, 222, 223, 225, 227, 229. 


colimensis, 162, 169, 220. 

copei, 162, 166, 220, 222, 223, 225. 

dicei, 162, 169, 224, 228. 

dugesii, 161, 169, 221, 222, 223. 

eee rubricaudatus, 161, 167, 
Ai 

humilis, 162, 166, 226. 

indubitus, 162, 168, 222, 223. 

lagunensis, 161, 167, 217. 

lynxe furcirostris, 161, 163, 222, 
225, 228. 

lynxe lynxe, 161, 163, 222, 225, 226, 
228. 


multivirgatus, 161, 165, 219. 
obsoletus, 161, 165, 219, 224, 228. 
ochoterenai, 162, 169, 221. 
parviauriculatus, 162, 166, 219, 227. 
parvulus, 162, 166, 220, 223, 226. 
pavimentatus, 160. 
quadrilineatus, 167. 
rovirosae, 164. 
schmidti, 164. 
schwartzei, 160, 162, 218, 227, 229. 
skiltonianus amblygrammus, 167. 
skiltonianus lagunensis, 168. 
Sn eaNe skiltonianus, 161, 167, 
217. 
sumichrasti, 161, 164, 218, 228. 
tetragrammus, 161, 165, 226, 
Euphryne, 79. 
obesus, 79, 81. 
Euprepes lynxe, 163. 
Euprepis microcephalus, 214. 


gaigeae, 
228. 3 


239 


exsul, Hemidactylus, 51. 
Uma, 89, 219. 
fantasticus, Sphaerodactylus, 52. 
fasciatus, Coleonyx, 42, 43, 220, 226. 
Crotaphytus, 94. 
Eublepharis, 43. 
Xenosaurus, 207, 208. 
fasciolatus, Crotaphytus, 94. 
ferrariperezi, Sceloporus, 126. 
fimbriatus, Gerrhonotus, 197. 
fissipes, Caiman, 212. 
flavescens, Kinosternon flavescens, 22, 
24, 219. 
Platythyra, 21, 24. 
flavilenta, Holbrookia maculata, 83. 
flavimaculata, Lepidophyma flavima- 
culata, 152, 218, 227, 228. 
flavimaculatus, Lepidophyma, 151, 152. 
flavomaculatum, Lepidophyma flavo- 
maculatum, 152. 
floridanus, Sceloporus spinosus, 117. 
forbesorum, Leiolopisma, 159. 
Leiolopisma gemmingeri, 159. 
Scincella gemmingeri, 157, 159, 222. 
formosus, Sceloporus, 107, 108. 
Sceloporus formosus, 107, 108, 224, 
225, 228. 
Tropidolepis, 108. 
franciscensis, Verticaria, 187. 
frenatus, Hemidactylus, 40, 50, 221. 
frontale, Phrynosoma blainvillii, 103. 
ce coronatum, 97, 103, 
217. 
frontalis, Phrynosoma, 103. 
fulvius, Elaps, 225. 
fulvus, Sceloporus, 135. 
furcirostris, Eumeces lynxe, 161, 168, 
222, 225, 228. 
fuscoauratus, Anolis, 55. 
fuscolabialis, Abronia, 196, 198, 224. 
Gerrhonotus, 198. 
fuscus, Caiman crocodilus, 212, 218. 
Gonatodes, 45, 218. 
Gonatodes albogularis, 45. 
Gymnodactylus, 45. 
Perosuchus, 212. 
Stenodactylus, 45. 


gabbii, Callisaurus draconoides, 86, 87, 
DUT, 22 


Callisaurus ventralis, 87. 

gadovi, Cnemidophorus, 178, 185, 227. 
Urosaurus, 140, 145, 222, 223. 
Uta, 145. 

gadowine, Sceloporus, 138, 221, 223, 224, 


gadovii, Anolis, 56, 61, 221. 
Barisia gadovii, 199, 200, 221. 
Gerrhonotus, 200. 

Ameiva undulata, 171, 

18, 225, 228. 

Gaigeia, 153, 154, 222. 

Lepidophyma, 153, 154. 

Pseudemys scripta, 31, 33. 

Sceloporus lundelli, 111, 112, 229. 


172, 


240 


Gaigeia, 151, 153. 
dontomasi, 153, 224. 
gaigeae, 153, 154, 222. 
radula, 153, 224. 
sylvatica, 153, 154, 222. 
Gambelia, 55, 98. 
wislizenii, 93, 94. 
wislizenii wislizenii, 94, 217, 219, 
227. 
gambelii, Crotaphytus, 94. 
Gastrotropis, 55. 
Gavialidae, 210. 
Gecko mabouia, 50. 
Geckonidae, 40. 
Gehyra, 51. 
Gekko rapicauda, 49. 
tuberculosus, 50. 
Gekkonidae, 39, 40, 41. 
gemmingeri, Leiolopisma, 159. 
Leiolopisma gemmingeri, 159. 
Lygosoma (Mocoa), 159. 
Oligosoma, 159. 
Scinecella gemmingeri, 157, 159, 222, 
224, 228. 
Geoclemmys annulata, 29. 
Geoemyda, 29. 
areolata, 29, 30, 218, 225, 227, 228, 
229. 
pulcherrima incisa, 29, 30, 218, 224. 
pulcherrima pulcherrima, 29, 30, 
220, 221, 228, 226. 
rubida, 29, 30, 220, 221, 223, 224. 
georgeensis, Aristelliger, 51, 225. 
Idiodactylus, 51. 
geronimensis, Anniella, 209, 217. 
Gerrhonotus, 194, 203. 
adspersus, 201. 
antauges, 200. 
bocourti, 200. 
cedrosensis, 206. 
deppii, 196, 197. 
deppii digueti, 198. 
fimbriatus, 197. 
fuscolabialis, 198. 
gadovil, 200. 
gramineus, 198. 
imbricatus, 198, 201. 
infernalis, 204. 
kingii, 205, 206. 
lemniscatus, 204. 
levicollis ciliaris, 202. 
levicollis levicollis, 202. 
lichenigerus, 201. 
liocephalus, 203. 
liocephalus austrinus, 203, 204, 218. 
liocephalus infernalis, 203, 204, 219, 
226. 
liocephalus liocephalus, 203, 214, 
220, 221, 222, 223, 224, 225. 
liocephalus loweryi, 203, 204, 226. 
liocephalus ophiurus, 203, 204, 225, 
228. 


matudae, 196. 
microcephalus, 214. 
modestus, 201. 
moreleti rafaeli, 199. 


BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Gerrhonotus moreleti temporalis, 199. 
multi-carinatus nanus, 207. 
multifasciatus, 206. 
nobilis, 206, 
oaxacae, 197. 
obscurus, 200. 
olivaceus, 201. 
ophiurus, 204. 
paucicarinatus, 206. 
(Barissia) planifrons, 201. 
rudicollis, 202. 
scincicauda ignavus, 207. 
scincicauda nanus, 207. 
scincicauda webbii, 207. 
taeniatus, 197. 
tessellatus, 203. 


vasconcelosii ochoterenai, 197. 
viridiflavus, 200. 
webbii, 207. 


gigas, Caretta, 16. 

Caretta caretta, 15, 16, 217, 226. 
gilberti, Xantusia, 154, 155, 217. 
glaucus, Sphaerodactylus, 53. 

Sphaerodactylus glaucus, 52, 53, 

218, 225, 227, 229. 
goldmani, Sceloporus, 136, 187, 219, 226. 
Terrapene, 35. 
Gonatodes, 40, 41, 45. 

albogularis, 45. 

albogularis fuscus, 45. 

fuscus, 45, 218. 

Goniodactylus braconnieri, 45. 
goodei, Anota, 102. 

Phrynosoma, 102. 

oer platyrhinos, 96, 101, 
Gopherus, 28. 

agassizii, 28, 217, 226. 

berlandieri, 28, 219, 224. 
gracilis, Cnemidophorus, 189. 

NOT bigris, led. , Looe 
graciosa, Uta, 144. 
graciosus, Sceloporus, 106. 

Urosaurus, 140, 141, 144, 217. 

Urosaurus ornatus, 145. 
grahamii, Cnemidophorus, 188. 
graminea, Abronia taeniata, 196, 198, 

225, 228. 
gramineus, Gerrhonotus, 198. 
grammicus, Sceloporus, 105, 119. 

Sceloporus grammicus, 119, 221, 224. 
grandaevus, Sator, 139, 140, 217. 
grandis, Cubina, 208. 

Xenosaurus, 207, 208, 225, 229. 
erayi, Emys, 33. 
grayii, Poriodogaster, 151. 
guatemalensis, Akleistops, 151, 152. 
guentheri, Sceloporus, 108. 
guentherii, Anolis, 64. 
gularis, Cnemidophorus, 183. 

Cnemidophorus gularis, 183. 

Cnemidophorus sackii, 178, 

224, 226, 228. 

Uta, 144, 

guntherii, Anolis, 57, 63. 


188, 


INDEX 


guttata, Clemmys, 37. 
guttatus, Cnemidophorus, 183. 
Cnemidophorus guttatus, 175, 179, 
224, 228. 
guttulatus, Lamprosaurus, 160, 165. 
Gymnodactylus albogularis, 45. 
coleonyx, 42. 
fuscus, 45. 
scapularis, 42. 
Gymnophiona, 2. 
Gymnophthalmus, 39, 170, 192. 
lineata, 192. 
sumichrasti, 192, 224. 


harlanii, Phrynosoma, 99. 
hartwegi, Ameiva undulata, 171, 218. 
heliactin, Anolis, 58, 67, 224. 
Heloderma, 192. 
hernandesii, 193. 
horridum, 193, 208, 218, 220, 221, 
222, 223, 224, 226, 227. 
suspectum, 193, 227. 
Helodermidae, 40, 192. 
Hemichirotes, 38. 
tridactylus, 38. 
Hemidactylus, 40, 41, 49. 
exsul, 51. 
frenatus, 40, 50, 221. 
javanicus, 49. 
mabouia, 40, 50, 228. 
mabuya, 49. 
mutilatus, 51, 52. 
navarri, 51, 52. 
tureicus, 40, 50. 
mueeus turcicus, 50, 218, 227, 228, 
29. 
hemilopha, Ctenosaura, 73, 75, 217, 219, 
226, 227. 
henrici, Cinosternum, 25. 
henshawi, Xantusia, 154, 155, 217. 
hernandesi, Phrynosoma, 100. 
Phrynosoma douglassii, 96, 100, 219, 
227. 
Phrynosoma orbiculare, 96, 97, 98. 
Tapaya, 100. 
hernandesii, Heloderma, 193. 
Iguana, 72. 
hernandezii, Chamaeleopsis, 68. 
Corythophanes, 68, 218, 224, 226, 
228, 229. 
Tapaya, 100. 
Herpetochalcis heteropus, 212. 
herrerai, Kinosternon, 22, 24, 228. 
hesperis, Uta stansburiana, 148, 149, 217. 
heterolepis, Sceloporus, 119, 121, 222. 
heteropa, Bachia, 212. 
heteropus, Chalcides, 212. 
Herpetochalcis, 212. 
heterurus, Sceloporus, 120. 
hiltoni, Pseudemys scripta, 31, 32, 226. 
hirtipes, Cinosternon, 25. 
Cinosternum, 25 


241 


Holbrookia, 54, 81. 
approximans, 83. 
bischoffi, 84. 
bunkeri, 82, 84, 219. 
dickersonae, 83. 
elegans, 84. 
elegans elegans, 84. 
elegans thermophila, 84. 
lacerata, 83. 
maculata, 82. 
maculata approximans, 82, 83, 219, 
220, 222, 226, 227. 
maculata dickersonae, 82, 83, 219, 
220, 221, 226, 229. 
maculata elegans, 82, 84, 226. 
maculata flavilenta, 83. 
maculata lacerata, 82, 83, 219. 
maculata pulchra, 82, 84. 
maculata thermophila, 82, 84, 226, 
227. 
propinqua, 85. 
propinqua piperata, 82, 85, 228. 
propinqua propinqua, 82, 83, 85. 
pulchra, 84. 
texana, 82, 85, 219, 220, 224, 226, 
227, 228. 
thermophila, 84. 
homolepidurus, Phyllodactylus, 46, 48, 
219, 227. 
Phyllodactylus tuberculosus, 48. 
Homalosaurus, 86. 
ventralis, 86, 88. 


horridum, Heloderma, 193, 208, 218, 
220, 221, 222, 223, 224, 226, 227. 
Trachyderma, 192, 193. 


horridus, Sceloporus horridus, 110, 116, 
221, 223, 225. 

humeralis, Sceloporus, 134. 

humilis, Eumeces, 162, 166, 226. 

hyperythra, Verticaria hyperythra, 187. 


hyperythrus, Cnemidophorus, 174. 
Cnemidophorus hyperythrus, 175, 
187, 217. 


Idiodactylus, 51. 
georgeensis, 51. 
ignavus, Gerrhonotus scincicauda, 207. 
Iguana, 54, 72. 
(Ctenosaura) armata, 74. 
(Ctenosaura) bellii, 74. 
delicatissima, 72. 
hernandesii, 72. 
iguana iguana, 72. 
iguana rhinolopha, 72, 218, 220, 
221, 222, 223, 224, 225, 226, 227, 
228. 
(Ctenosaura) lanceolata, 74. 
H[ypsilophus] rhinolophus, 72. 
(Ctenosaura) similis, 73. 
tuberculata, 72. 


Kinosternon, 22, 25, 219, 220, 221, | iguana, Iguana iguana, 72. 


222, 223, 226. 
hispidus, Sauromalus, 79, 80, 217. 


Lacerta, 72. 
Iguanidae, 40, 53. 


242 


imbricata, Barisia imbricata, 198, 199, 
201, 220, 221, 222, 223, 224, 225 
228. 
Eretmochelys, 
225, 226. 
Testudo, 16. 
imbricatus, Gerrhonotus, 198, 201. 
immucronatus, Sceloporus jarrovii, 123, 
128, 222, 225, 226, 229. 
immutabilis, Cnemidophorus, 180. 
Cnemidophorus guttatus, 175, 180, 
218, 220, 221, 223, 224. 
imperator, Boa, 225. 
Constrictor constrictor, 225. 
impetigosus, Anolis, 56, 60. 
incerta, Scincella, 156. 
incisa, Emys, 30. 
Geoemyda pulcherrima, 29, 30, 218, 
224, 


indubitus, Eumeces, 162, 168, 222, 223. 
infernalis, Gerrhonotus, 204. 
Gerrhonotus liocephalus, 203, 204, 
219, 226. 
inornatus, Cnemidophorus, 177, 184, 
219, 224. 
Sphaerodactylus, 53. 
insulana, Ctenosaura, 75. 
insularis, Crotaphytus, 92, 93, 217. 
integrum, Cinosternum, 25. 
“Cinosternum scorpioides, 2D 
Kinosternon, 22, 25, 217, 220, 221, 
222223. 924, 295, 226, 228. 
Kinosternum, 21, 25. 
intermedius, Anolis, 63. 
Sceloporus, 127. 
Sceloporus dugesii, 
223 


, 


16, 217, 218, 224, 


122, 127, 221, 


Tropidolepis, 127. 
interrupta, Ctenosaura, 75. 
inusitatus, Callisaurus, 88. 
Callisaurus draconoides, 86, 88, 227. 
Callisaurus ventralis, 88. 
irregularis, Aristelliger, 51. 
Phymatolepis (Uta), 146. 
Urosaurus, 140, 146. 
Uta, 146. 
ixbaac, Leiolopisma cherriei, 158. 
Lygosoma assatum, 158. 
Scincella cherriei, 157, 158, 218, 229. 


jacobi, Anolis, 68. 
jalapae, Sceloporus, 136, 225, 229. 
jamesi, Phrynosoma, 102. 
Phrynosoma coronatum, 97, 102, 
217. 
jarrovii, Sceloporus, 127. 
Sceloporus jarrovii, 
219, 220, 227. 
javanica, Amyda, 18. 
javanicus, Hemidactylus, 49. 


kempii, Lepidochelys, 14, 15, 225. 
Thalassochelys (Colpochelys), ily 
kidderi, Anolis, 58, 66, 218, 229. 
kingii, Elgaria, 205. 
Elgaria kingii, 205, 219. 
Gerrhonotus, 205, ‘206. 


122, 123, 128, 


BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Kinosternidae, 19, 21. 

Kinosternon, 3, 21. 
abaxillare, 22, 218. 
acutum, 22, 23, 218, 227, 228. 
creaseri, 22, 23, 225, 229. 
cruentatum consors, 22, 24, 225, 


229. 
cruentatum cruentatum, 22, 23, 
218, 224, 227: 


flavescens flavescens, 22, 24, 219. 

flavescens stejnegeri, 22, 24, 219, 
220, 226. 

herrerai, 22, 24, 228. 

hirtipes, 22, 25, 219, 220, 221, 222, 
223, 226. 

integrum, 22, 25, 217, 220, 221, 
222, 223, 224, 225, 226, 228. 

leucostomum, 22, 25, 218, 225, 227, 
228. 

longicaudatum, 21. 

scorpioides, 21. 

scorpioides acuta, 23. 

sonoriense, 22, 26, 219, 220, 226. 


Kinosternum integrum, 21. 
mexicanum, 23. 
sonoriense, 26. 


klauberi, Terrapene, 34, 35, 227. 
Sauromalus, 79, 80, 217. 


labialis, Cnemidophorus, 178, 185, 217. 
lacerata, Holbrookia, 83. 
Holbrookia maculata, 82, 83, 219. 


Lacerta acanthura, 74. 
americana, 170. 
iguana, 72. 
lumbricoides, 39. 
mexicana, 39. 
orbicularis, 94, 95, 97. 
quadrilineata, 192. 
sexlineata, 174, 185. 


turcica, 50. 
Lacertus mabouya, 155, 156. 
orbicularis, 97. 


lachrymata, Chelonia, 16. 
lacordairei, Alligator, 211. 
Laemanctus, 54, 69. 
alticoronatus, 70, 229. 
deborrei, 70, 71, 224, 227. 
longipes, 69, 70, 71, 220, 224, 228. 
serratus, 70, 218, 221, 222, 224, 
226, 228, 229. 


laeviventris, Anolis, 57, 58, 62, 63, 227, 
228 


Dactyloa (Anolis), 62. 
lagunensis, Eumeces, 161, 167, 217. 
Eumeces skiltonianus, 168. 

Plestiodon, 168. 

Plestiodon skiltonianus, 168. 
Lampropholis assatus, 160. 
Lamprosaurus, 160. 

guttulatus, 160, 165. 
lanceolata, Iguana (Cisngesurayl 74, 
lanei, Phyllodactylus, 47, 48, 220, 221, 

222, 223, 226. 


INDEX 


lar, Aristelliger, 51. 
laterale, Leiolopisma, 159. 
Lygosoma, 159. 
Scincella, 157, 159, 219. 
lateralis, Lysoptychus, 105, 182. 
Sceloporus, 132. 
Scincus, 159. 
Uta, 144. 
Uta ornata, 144. 
Uta (Phymatolepis), 144. 
latirostris, Crocodilus, 212. 
lativittis, ‘Cnemidophorus, 178. 
Leiolopisma, 156. 
assatum assatum, 160. 
assatum taylori, 160. 
caudaequinae, 158. 
cherriei, 158. 
cherriei cherriei, 157. 
cherriei ixbaac, 158. 
cherriei stuarti, 158. 
forbesorum, 159. 
gemmingeri, 159. 
gemmingeri forbesorum, 159. 
gemmingeri gemmingeri, 159. 
laterale, 159. 
silvicolum, 158. 
telfairii, 156. 
Leiosaurus belli, 212. 
lemniscatus, Gerrhonotus, 204. 
Lepidochelys, 14. 
kempii, 14, 15, 225. 
olivacea, 14, 15, 218, 220, 221, 
224, 226. 
Lepidophyma, 151. 
flavimaculata flavimaculata, 
218, 227, 228. 
flavimaculata obscura, 151. 
flavimaculatus, 151, 152. 
ona flavomaculatum, 
52. 
gaigeae, 153, 154. 
smithii, 152. 
smithii occulor, 152, 153, 225, 226. 
smithii smithii, 152, 218. 
smithii tehuanae, 152, 218, 224. 
sylvatica, 154. 
Lepidosauria, 12, 37. 
leucostoma, Swanka, 25. 
leucostomum, Cinosternon, 25. 
Kinosternon, 22, 25, 218, 225, 227, 
228. 
levicollis, Barisia, 199, 202, 219. 
Gerrhonotus levicollis, 202. 
levigata, Barisia gadovii, 199, 200, 224. 
lichenigerus, Gerrhonotus, 201. 
licki, Seeloporus, 114. 
‘Seeloporus orcutti, 111, 114, 217. 
Limnochelone, 19 
micrura, 19. 
linearis, Urosaurus ornatus, 


152, 


Uta ornata, 143. 
lineata, Gymnophthalmus, 192. 
lineatissimus, Cnemidophorus, 179. 
Cnemidophorus deppii, 176, 179, 
220, 221, 222, 223, 225. 


142, 143, 


243 


lineatulus, Sceloporus, 115. 

Sceloporus magister, 110, 115, 217. 
lineolateralis, Sceloporus, 122, 126, 220. 
lineolatus, Sphaerodactylus, 52. 
liocephalus, Gerrhonotus, 203. 

Gerrhonotus liocephalus, 208, 214, 

220, 221, 222, 223, 224, 225. 
liogaster, Anolis, 57, 62, 221. 
Liolaemus tenuis, 213. 
littoralis, Malaclemmys, 36. 

Malaclemmys terrapin, 36. 

Malaclemys terrapin, 36. 
longicaudatum, Kinosternon, 21. 
longicaudatus, Tapaya orbicularis, 97. 
longipes, Laemanctus, 69, 70, 71, 220, 

224, 228. 
Loricata, 2, 12, 209. 
loweryi, Gerrhonotus liocephalus, 203, 
204, 226. 
lucasensis, Dipsosaurus dorsalis, 78, 
17. 
lumbricoides, Lacerta, 39. 
lunaei, Sceloporus, 107. 
lundelli, Sceloporus lundelli, 
218. 
Lygosoma assatum assatum, 160. 

assatum cherriei, 157. 

assatum ixbaac, 158. 

cherriei cherriei, 157. 

cherriei stuarti, 158. 

(Mocoa) gemmingeri, 159. 

laterale, 159. 
lynxe, Eumeces lynxe, 161, 163, 222, 

225, 226, 228. 

Euprepes, 163. 
Lysoptychus, 105. 

lateralis, 105, 132. 


ALL w012 


mabouia, Gecko, 50. 
Hemidactylus, 40, 50, 228. 
Mabouia brevirostris, 168. 
mabouya, Lacertus, 155, 156. 
Mabuia alliacea, 156. 
Mabuya, 155. 
mabouya mabouya, 156, 218, 220, 
221, 222, 224; 226, 227, 228, 229. 
mabuya, Hemidactylus, 49. 
maccalli, Anota, 99. 
Macrochelys, 20. 
maculata, Holbrookia, 82. 
Swanka, 238, 25. 
maculosa, Chelonia, fe 
maculosus, Sceloporus, 105, 133, 220. 
magister, Seeloporus magister, 111, 112, 
114, 217, 219, 220, 227. 
magnatuberculatus, ‘Phyllodactylus, 47, 


48, 221. 

magnus, Phyllodactylus, 47, 48, 218, 
221, 224 

malachiticus, Sceloporus malachiticus, 
107. 


Malaclemmys littoralis, 36. 
terrapin littoralis, 36. 
Malaclemys, 29, 36. 
terrapin littoralis, 36. 
mannophorus, Uta, 148, 149, 217. 


244 


mariarum, Cnemidophorus, 183. 


Cnemidophorus sackii, 178, 183, 
223. 

marmoratus, Cnemidophorus, 190. 
Cnemidophorus tigris, 177, 190, 


219, 220. 
Sceloporus, 131. 
Sceloporus variabilis, 130, 131, 219, 
224, 228. 
Staurotypus, 27. 
martinensis, Uta, 148, 149, 217. 
martyris, Cnemidophorus, "191. 
Cnemidophorus tessellatus, 191. 
Cnemidophorus tigris, 176, 191, 227. 
matudae, Gerrhonotus, 196. 
matudai, Abronia, 196, 218. 
mawii, Dermatemys, 19, 218, 224, 227, 
228, 229. 
maximus, Cnemidophorus, 176, 188, 
Qt: 
mayensis, Anolis sagrei, 56, 59, 218, 227, 
229 


Hee affinis, 225. 
m’ealli, Doliosaurus, 99. 
m ’eallii, Anota, 95, 99. 
Phrynosoma, 96, 99,217, 227. 
mearnsi, Streptosaurus, 91, 217. 
Uta, 91. 
meeki, Cnemidophorus gularis, 184. 
Megadactylus, 85. 
megalepidurus, Sceloporus, 106, 
225, 228, 229. 
megalocephalus, Claudius, 26. 
megapholidotus, Anolis, 56, 59, 221. 
Megemys, 17. 
melanogaster, Sceloporus, 126. 
Sceloporus ferrariperezi, 126. 
Sceloporus torquatus, 122, 126, 217, 
221, 222, 228, 226, 229. 
melanorhinus, Sceloporus, 112, 1138. 
Seeloporus melanorhinus, 110, 112, 
225. 
melanostethus, Cnemidophorus, 189, 190. 
mentalis, Phyllodactylus, 46. 
merriami, Sceloporus, 105, 132. 
Sceloporus merriami, "132. 
Mesaspis, 198. 
moreletii, 198. 
metallicus, Anolis, 56, 59. 
mexicana, Cistudo, 34. 
Cistudo (Onychotria), 34, 
Lacerta, 39. 
Onychotria, 34. 
Rhinoclemmys, 30. 
Terrapene, 35. 
Terrapene mexicana, 34, 35, 226, 
228. 
mexicanum, Kinosternum, 23. 
mexicanus, Chamaeleo, 68. 
Cnemidophorus, 181. 
Corythophanes, 68. 
Crocodilus, 211. 
microcephalus, ’Dasia, 214. 
Diploglossus, 214, 
Euprepis, 214. 
Gerrhonotus, 214. 


121, 


BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


microlepidopterus, Sceloporus, 120. 
microlepidopus, Cnemidophorus, 180. 
microlepidotus, Sceloporus, 120. 
Sceloporus grammicus, 119, 120, 
220, 221, 222, 223, 225, 228, 229. 
Sceloporus microlepidotus, 120. 
microlepis, Sceloporus, 120. 
microscutata, Uta, 145. 
microscutatus, Urosaurus, 140, 145, 217. 
micrura, Limnochelone, 19 
Micrurus affinis mayensis, 225. 
milleri, Anolis, 57, 64, 224. 
minor, "Sceloporus j jarrovii, 123, 128, 221, 
222, 224, 225, 226, 229. 
Sceloporus torquatus, 128. 
mississippiensis, Alligator, 211. 
mitratus, Brachydactylus, 41. 
mitrella, ’Cristasaura, 7X 
Mocoa, 159. 
Mocoa’ cherriei, 157. 
modesta, Anota, 101. 
Barisia, 199, 201, 228. 
modestum, ’Phrynosoma, 95, 101, 219, 
220, 224, 226, 227, 229. 
modestus, Doliosaurus, 101. 
Gerrhonotus, 201. 
Pterogasterus, 201. 
Molinia, 210. 
monotropis, Diploglossus, 214. 
monserratensis, Sceloporus, 115. 
Sceloporus magister, 112, UU5; 27. 
moreletii, Crocodylus, 210, ‘211, 218, 
220, 227, 228, 229, 
Mesaspis, 198. 
motaguae, Cnemidophorus, 181. 
mucronatus, Sceloporus mucronatus, 
123, 124, 222, 225, 229. 
Sceloporus. torquatus, 124, 125. 
multicarinata, Elgaria multicarinata, 
205. 
multicarinatus, Cordylus (Gerrhonotus), 
205. 
multifasciatus, Gerrhonotus, 206. 
multiscutatus, Cnemidophorus tessella- 
tus, 191. 
Cnemidophorus tigris, 177, 
2G: 


multispinis, Ctenosaura, 74. 
multivirgatum, Plestiodon, 165. 
multivirgatus, Eumeces, 161, 165, 219. 
muralis, Phyllodactylus, 46, 47, 224. 
murinus, Seps, 175. 
mutilatus, Hemidactylus, 51, 52. 
Peropus, 51, 52, 223, 226. 
Mydas, 17. 
mydas, Chelonia, 17, 221, 223, 224, 225, 
226, 228, 229. 
Testudo, 17. 
Mydasea, 17. 


191, 


nana, Elgaria multicarinata, 205, 207, 
1 


nannodes, Anolis, 68. 


nanus, Gerrhonotus multi-carinatus, 
ia 


Gerrhonotus scincicauda, 207. 


INDEX 


nasuta, Caretta, 15. 
navarri, Hemidactylus, 51, 52. 
nebuloides, Anolis, 58, 66, 219, 220, 221, 
224, 225, 227. 
nebulosa, Chrysemys, 32, 217. 
Dactyloa, 55, 65. 
Pseudemys scripta, 31, 32, 217. 
nebulosus, Anolis, 58, 65, 220, 221, 222, 
223, 224, 225, 226. 
nelsoni, Phrynosoma, 103. 
Sceloporus, 138, 189, 219, 222, 223, 
226, 227. 
Terrapene, 34, 35, 223. 
Urosaurus bicarinatus, 
225. 
Uta, 147. 
Uta bicarinata, 147. 
nemoralis, Coleonyx elegans, 
220, 221, 223, 224. 
Neoseps, 155. 
newmanorum, Xenosaurus, 


141, 147, 


42, 43, 


207, 208, 


226. 
nigricauda, Uta, 145. 
nigricaudus, Urosaurus, 140, 145, 217. 


niloticus, Crocodylus, 210. 

nobilis, Elgaria, 206. 
Elgaria kingii, 205, 206. 
Gerrhonotus, 206. 

nolascensis, Uta, 148, 151, 227. 

Norops tropidonotus, 60. 
yucatanicus, 60. 

notata, Uma, 89. 
Uma notata, 89, 217. 

nuchalis, Basiliscus (Cristasaura), 72. 


oaxacae, Abronia, 196, 197, 224. 
Gerrhonotus, 197. 
oberon, Sceloporus jarrovii, 123, 129, 219. 
obesus, Euphryne, 79, 81. 
Sauromalus obesus, 79, 81. 
obscura, Lepidophyma flavimaculata, 


obscurus, Gerrhonotus, 200. 
Sceloporus, 109. 
obsoletum, Plestiodon, 160, 165. 


obsoletus, Cnemidophorus gularis gu- 
laris, 182. 
Eumeces, 161, 165, 219, 224, 228. 


obtusirostris, Ecphymotes, 213. 


occidentalis, Cnemidophorus communis, 
182. 


occipitalis, Aneuporus, 213. 
Tropidurus, 213. 
occulor, Lepidophyma smithii, 152, 153, 
225, 22 


ochoterenai, Abronia, 196, 197, 218. 
Eumeces, 162, 169, 221. 
197. 


Gerrhonotus vasconcelosii, 
Phrynosoma, 103. 
Sceloporus, 133, 135, 221, 223. 
octolineatus, Cnemidophorus, 184. 
Cnemidophorus gularis, 183. 
Oedicoryphus, 71. 


245 


oligoporus, Cnemidophorus deppii, 176, 
179, 218, 228. 
Sceloporus, 116. 
Sceloporus horridus, 110, 116, 220, 
221, 222, 223. 
Oligosoma gemmingeri, 159. 
olivacea, Chelonia, 14, 15. 
Lepidochelys, 14, ‘15, 218, 220, 221, 
224, 226. 
olivaceus, 'Gerrhonotus, 201. 
Sceloporus, 111, 117, 219, 224, 226, 
228 


rela Sceloporus variabilis, 129, 

130. 

omiltemanus, Sceloporus, 124. 
Sceloporus mucronatus, 123, 124, 


221, 225. 
Sceloporus torquatus, 124. 
Onychotria, 34. 
mexicana, 34. 
Ophisaurus, 194. 
ophiurus, Gerrhonotus, 204. 
Gerrhonotus liocephalus, 203, 204, 
225, 228. 
Pterogasterus, 204. 
orbiculaire, Agama, 97. 
orbiculare, Phrynosoma, 96, 97, 98. 
Phrynosoma orbiculare, 96, 97, 219, 
220, 222, 223, 224, 225, 226, 228, 
229. 
orbicularis, Lacerta, 94, 95, 97. 
Lacertus, 97. 
Tapaya, 94, 98. 
orcutti, Sceloporus orcutti, 111, 114, 217. 
ornata, Cistudo, 36. 
Emys, 31. 
Pseudemys scripta, 31, 33, 218, 222, 
224, 225, 226, 227, 229. 
Terrapene, 34, 36, 219, 227. 
Uta, 142. 
Uta ornata, 142. 
ornatus, Urosaurus ornatus, 142. 
Seeloporus, 127: 
Sceloporus ornatus, 122, 127, 219. 


pacificus, Crocodilus, 211. 

palearis, Cyclura, 76. 

pallida, Clemmys marmorata, 37, 217. 

palmeri, Uta, 148, 151, 227. 

panamensis, Anolis, 61. 

papillosus, Anelytropsis, 170, 226, 228. 

Paraloma, 71. 

parkeri, Ctenosaura, 75. 

parva, Ameiva undulata, Meio w2NS: 
24 


Uta, 149. 
parviauriculatus, 
219, 227. 
parviscutata, Uta, 145. 
arte Eumeces, 162, 166, 220, 223, 
parvus, Sceloporus, 131. 
Sceloporus parvus, 129, 131, 219, 
224, 226. 
paucicarinatus, Elgaria, 205, 206, 217. 
Gerrhonotus, 206. 


Eumeces, 162, 166, 


246 


pavimentatus, Eumeces, 160. 
Scincus, 160 


pectinata, Ctenosaura, 73, 75, 220, 221, 


922, 223, 224, 225, 226. 
Cyclura, 75. 


peninsularis, Coleonyx variegatus, 42, 


pentaprion, Anolis, 55, 57, 61, 218. 
Anolis (Coccoéssus), 61. 


ercarinatus, Corythophanes, 68, 69 ,218. 


eripia, 51. 
peronii, 51. 
peronii, Peripia, 51. 
Peropus, 51. 
Peropus, 40, 41, 51. 
mutilatus, 51, 52, 223, 226. 
peronii, BL. 
Perosuchus, 212; 
fuscus, 212. 
perplexus, Cnemidophorus, 184, 185. 
petersii, Anolis, 58, 65, 226, 228. 
Petrosaurus, 55, 90. 
repens, 90, 91, 217. 
thalassina, 90, 217. 
Phrynosoma, 54, 94, 102. 


asio, 95, 102, 218, 220, 221, 223, 224. 


blainvillii, 103. 

blainvillii frontale, 103. 

bufonium, 99. 

boucardi, 96, 98, 222. 

boucardii, 98. 

braconnieri, 95, 100, 224, 225. 

calidiarum, 101. 

cerroense, 97, 103, 217. 

cornutum, 94, 95, 99, 219, 220, 
224, 226, 227. 

cornutum planiceps, 99. 

coronatum, 94, 96. 

coronatum blainvillii, 97, 103, 217. 

coronatum coronatum, 97, 102, 217. 

coronatum frontale, 97, 103, 217. 

coronatum jamesi, 97, 102, 217. 

ditmarsi, 95, 104, 227. 

douglassii, 96, 100. 

count brachycercum, 96, 100, 
220 

douglassii 96, 100, 
219, 22h 

frontalis, 103. 

goodei, 102. 

harlanii, 99. 

hernandesi, 100. 

jamesi, 102. 

m’caillii, 96, 99, 217, 227. 

modestum, 95, 101, 219, 220, 224, 
226, 227, 229. 

nelsoni, 103. 

ochoterenai, 103. 

orbiculare, 96, 97, 98. 

orbiculare cortezii, 96, 98, 225, 228. 

orbiculare dugesii, 97, 98, 220. 

orbiculare hernandesi, 100. 

orbiculare orbiculare, 96, 97, 219, 
220, 222, 223, 224, 225, 226, 
228, 229. 

planiceps, 99. 


hernandesi, 


BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Phrynosoma platyrhinos, 95, 101. 
platyrhinos goodei, 96, 102, 227. 
platyrhinos platyrhinos, 96, 101, 

217 


regale, 104. 

schmidti, 103. 

solare, 96, 104, 217, 226, 227. 

solaris, 104. 

spinimentum, 102. 

taurus, 95, 104, 221, 225. 

wiegmanni, 97. 

Phyllodactylus, 41, 46. 
bordai, 46, 47, 221. 
deleampi, ‘46, ‘47, 221. 
homolepidurus, 46, 48, 219, 227. 
lanei, 47, 48, 220, 221, 222, 223, 226. 
magnatuberculatus, 47 E 48, 2211, 
magnus, 47, 48, 218, 221, 224. 
mentalis, 46. 
muralis, 46, 47, 224. 
pulcher, 46. 
tuberculosus, 47, 49, 217. 
tuberculosus homolepidurus, 48, 
unctus, 46, 47, 217. 
(Diplodactylus) unctus, 47. 
xanti, 49. 

Phymatolepis, 140, 144. 
bicarinatus, 140. 
bi-carinatus, 146. 

(Uta) irregularis, 146. 
picta, Testudo, 33. 

Verticaria, 186. 

Xantusia, 155. 
pictus, Claudius, 27. 

Chemidophorus hyperythrus, 

186, 217. 

Sceloporus, 121, 225, 229. 
pilsbryi, Sceloporus, 120. 
piperata, Holbrookia propinqua, 82, 85, 

228. 

planiceps, Phrynosoma, 99. 
Phrynosoma cornutum, 99. 

planifrons, Barisia imbricata, 199, 201, 

224. 

Gerrhonotus (Barissia), 201. 
Platypholis, 160. 
platyrhina, Anota, 101. 
platyrhinos, Doliosaurus, 101. 

Phrynosoma, 95, 101. 

Phrynosoma platyrhinos, 96, 101, 

217, 


175, 


Platysternidae, 18. 

Platythyra, 21. 
flavescens, 21, 24. 

Plestiodon bellii, 163. 
lagunensis, 168. 
multivirgatum, 165. 
obsoletum, 160, 165. 
quinquelineatum, 163. 
skiltonianum, 167. 
skiltonianus lagunensis, 168. 
tetragrammus, "165. 


Plethodontidae, 2. 
Pleurodira, 12. 


pleurostictus, Sceloporus, 119. 


INDEX 


plioporus, Sceloporus serrifer, 122, 124, 
218, 226, 227, 229. 

Plistodon sumichrasti, 164. 

Pnoépus, 49. 

podarga, Ameiva undulata, 171, 172, 
226, 228. 

poinsettii, Sceloporus, 121, 122, 125, 

219, 220, 224. 

Sceloporus torquatus, 125. 
polyphemus, Testudo, 28. 
pe Crotalus, 6. 

oriodogaster, 151. 

grayii, 151. 
praesignis, Aristelliger, 51. 
prezygus, Sceloporus, 107, 109, 218. 
Pristicercus, 55. 
propinqua, Holbrookia, 85. 

Holbrookia propinqua, 82, 83, 85. 
propus, Chamaesaura, 39. 

Pseudemys, 29, 31. 

elegans, 32. 

floridana texana, 31, 33, 224. 

scripta cataspila, 31, 32, 228. 

scripta elegans, 31, 32, 219, 228. 

scripta gaigeae, 31, 33. 

scripta hiltoni, 31, 32, 226. 

scripta nebulosa, 31, 32, 217. 

scripta ornata, 31, 33, 218, 222, 

224, 225, 226, 227, 229. 

texana, 33. 

troostii elegans, 32. 

umbra, 31, 33, 224. 
pseudo-caretta, Chelonia, 16. 
proedamy css Chelonia, 16. 

terogastenes, 203. 
Pterogasterus modestus, 201. 

ophiurus, 204. 
ptychopleurus, Tropidurus, 218. 
pulcher, Phyllodactylus, 46. 
pulcherrima, Emys, 30. 

Geoemyda pulcherrima, 29, 30, 220, 

221, 2238, 226. 
pulchra, Anniella, 209. 

Anniella pulchra, 209, 217. 

Holbrookia, 84. 

Holbrookia maculata, 82, 84. 
punctata, Testudo, 37. 
punctatissimus, Sphaerodactylus, 52. 
punctatum, Cinosternon, 26 
punctilineatus, Cnemidophorus, 189. 
pyrocephalus, Sceloporus, 105, 138, 139, 

220, 221, 222; 223. 
pyrrhocephalus, Sceloporus, 139. 


quadrilineata, Lacerta, 192. 
quadrilineatus, Eumeces, 167. 
quinquecarinata, Cyclura, 76. 
quinquecarinatus, Enyaliosaurus, 76, 224. 
quinquelineatum, Plestiodon, 168. 
quinquelineatus, Scincus, 163. 


rackhami, Xenosaurus, 208, 218. 

radula, Gaigeia, 153, 224. 

rafaeli, Barisia moreleti, 198, 199, 218. 
Gerrhonotus moreleti, 199. 


247 


Rapara, 20. 
rapicauda, Gekko, 49. 
Thecadactylus, 49. 
rapicaudus, Thecadactylus, 49, 229. 
regale, Phrynosoma, 104. 
remivaga, Caretta, 15. 
repens, Petrosaurus, 90, 91, 217. 
Uta, 90. 
Reptilia, 12. 
reticulatus, Crotaphytus, 92, 93, 219, 
224, 228. 
Rhinoclemys, 29. 
Rhinoclemmys mexicana, 30. 
rhinolopha, Iguana iguana, 72, 218, 220, 
221, 222, 223, 224, 225, 226, 227, 228. 
rhinolophus, I{guana] H[ypsilophus], 72. 
rhodostictus, Callisaurus, 87. 
Rhynchocephalia, 37. 
rodriguezii, Anolis, 64. 
Anolis limifrons, 57, 64, 218, 224, 
225, 229. 
rossignonii, Chelydra, 20. 
Emysaurus, 20. 
rostellum, Cinosternum, 25. 
rovirosae, HEumeces, 164. 
rozellae, Celestus, 195, 218. 
rubida, Chelopus, 30. 
Geoemyda, 29, 30, 220, 221, 223, 224. 
rubidus, Cnemidophorus, 191. 
Cnemidophorus tessellatus, 191. 
Cnemidophorus tigris, 176, 191, 217. 
rubigenosus, Anolis, 64. 
rubricaudatus, Eumeces gilberti, 161, 
167, 217. 
rubriventris, Sceloporus, 119. 
rudicollis, Barisia, 199, 202, 222. 
Gerrhonotus, 202. 
rufidorsum, Sceloporus, 115. 
Sceloporus magister, 112, 115, 217. 
rufopunctata, Uma, 89. 
ruthveni, Anolis, 60. 


sackii, Cnemidophorus, 181. 
Cnemidophorus sackii, 
QP, 223, 225, 
Cnemidophorus sexlineatus, 180, 181. 
sagraei, Anolis, 55. 
Salientia, 2. 
sallaei, Anolis, 67. 
salvini, Sceloporus, 108. 
Sceloporus malachiticus, 107, 108, 
PPAR PAL. 
salvinii, Staurotypus, 27, 224. 
Staurotypus (Stauremys), 27. 
Sator, 55, 139. 
angustus, 139, 217. 
grandaevus, 139, 140, 217. 
Sauria, 2, 12, 37, 39. 
Saurochelys, 20. 
Sauromalus, 54, 79. 
ater, 79, 80, 217. 
australis, 79, 80, 217. 
hispidus, 79, 80, 217. 
klauberi, 79, 80, 217. 
obesus obesus, 79, 81. 
obesus townsendi, 79, 81, 227. 


178, 180, 


248 


Sauromalus obesus tumidus, 79, 81. 


slevini, 79, 80, 217. 
townsendi, 81. 
varius, 79, 81, 227. 


scalaris, Cnemidophorus gularis, 182. 


Cnemidophorus sackii, 178, 182, 
219, 220. 

Sceloporus, 105, 137. 

Sceloporus scalaris, 136, 137, 220, 
221, 222, 223, 229. 


scapularis, Gymnodactylus, 42. 
Sceloporus, 55, 105. 


acanthinus, 108. 

aeneus, 137. 

aeneus aeneus, 136, 137, 220, 221, 
223, 225. 

aeneus bicanthalis, 186, 137, 222, 
224, 225, 228. 

asper, 107, 109, 221, 222, 223. 

binocularis, 126. 

bi-seriatus, 118. 

boulengeri, 113. 

bulleri, 122, 125, 222. 

carinatus, 134, 135, 218. 

cautus, 117, 219, 226, 229. 

chrysostictus, 105, 183, 218, 226, 
229. 

clarkii, 113. 

clarkii boulengeri, 111, 113, 222, 
223, 226, 227. 

clarkii clarkii, 111, 113, 219, 227. 

cochranae, 134. 

consobrinus, 118. 

couchii, 105, 129, 132, 219, 224. 

cozumelae, 129, 180, 226, 229. 

cupreus, 134. 

cyanogenys, 123, 125, 224, 228. 

delicatissimus, 131. 

digueti, 114. 

dispar, 120. 

disparilis, 120. 

dugesii, 127. 

dugesii dugesii, 122, 127, 220, 222, 
223% 

dugesii intermedius, 122, 127, 221, 
223 


edwardtaylori, 110, 112, 224. 

ferrariperezi, 126. 

ferrariperezi binocularis, 126. 

ferrariperezi melanogaster, 126. 

formosus, 107, 108. 

formosus formosus, 107, 108, 224, 
225, 228. 

formosus scitulus, 107, 109, 221. 

fulvus, 135. 

gadoviae, 138, 221, 223, 224, 225. 

goldmani, 136, 137, 219, 226. 

graciosus, 106. 

eracioene vandenburgianus, 119, 

grammicus, 105, 119. 

grammicus alpha, 120. 

grammicus disparilis, 119, 120, 219, 
220, 221, 222, 224, 226, 228, 229. 





BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


Sceloporus grammicus grammicus, 119, 


221, 224. 

grammicus microlepidotus, 119, 
120, 220, 221, 222, 223, 225, 228, 229. 

guentheri, 108. 

heterolepis, 119, 121, 222. 

heterurus, 120. 

horigyy albiventris, 110, 116, 222, 

horridus_horridus, 110, 116, 221, 
223, 225. 

horridus oligoporus, 110, 116, 220, 
221, 222, 223. 

humeralis, 134. 

intermedius, 127. 

jalapae, 136, 225, 229. 

jarrovii, 128. 

Jarrovii immucronatus, 123, 128, 
222, 225, 226, 229. 

jarrovii jarrovii, 122, 123, 128, 219, 
220, 227. 

jarrovii minor, 123, 128, 221, 222, 
224, 225, 226, 229. 

jarrovii oberon, 123, 129, 219. 

jarrovii sugillatus, 123, 128, 222. 

lateralis, 132. 

licki, 114. 

lineatulus, 115. 

lineolateralis, 122, 126, 220. 

lunaei, 107. 

lundelli gaigeae, 111, 112, 229. 

lundelli lundelli, 111, 112, 218. 

maculosus, 105, 133, 220. 

magister lineatulus, 110, 115, 217. 

magister magister, 111, 112, 114, 
217, 219, 220, 227. 

magister monserratensis, 112, 115, 
oii. 

magister rufidorsum, 112, 115, 217. 

magister zosteromus, 112, 115, 217. 

malachiticus acanthinus, 107, 108, 
218. 

malachiticus malachiticus, 107. 

malachiticus salvini, 107, 108, 225, 
229. 

malachiticus smaragdinus, 107. 

malachiticus taeniocnemis, 107, 218. 

marmoratus, 131. 

megalepidurus, 106, 121, 225, 228, 
229. 

melanogaster, 126. 

melanorhinus, 112, 113. 

melanorhinus calligaster, 110, 113, 
220, 221, 222, 223. 

melanorhinus melanorhinus, 110, 
112, 225. 

melanorhinus stuarti, 110, 113, 218. 

merriami, 105, 132. 

merriami annulatus, 132, 133, 219. 

merriami merriami, 132. 

microlepidopterus, 120. 

microlepidotus, 120. 

microlepidotus disparilis, 120. 

microlepidotus microlepidotus, 120. 

microlepis, 120. 

monserratensis, 115. 


INDEX 


Sceloporus mucronatus aureolus, 123, 


124, 225, 229. 

mucronatus mucronatus, 123, 124, 
222, 225, 229. 

mucronatus omiltemanus, 123, 124, 
221, 225. 

nelsoni, 138, 1389, 219, 222, 223, 
226, 227. 

obscurus, 109. 

occidentalis biseriatus, 117,118, 217. 

ochoterenai, 133, 135, 221, 223. 

oligoporus, 116. 

olivaceus, 111, 117, 219, 224, 226, 
228. 

omiltemanus, 124. 

orcutti licki, 111, 114, 217. 

orcutti orcutti, 111, 114, 217. 

ornatus, 127. 

ornatus caeruleus, 122, 127, 219. 

ornatus ornatus, 122, 127, 219, 

parvus, 131. 

parvus parvus, 129, 131, 219, 224, 
226. 


parvus scutulatus, 129, 131, 222, 
226. 

pictus, 121, 225, 229. 

pilsbryi, 120. 

pleurostictus, 119. 

poinsettii, 121, 122, 125, 219, 220, 
224. 

prezygus, 107, 109, 218. 

pyrocephalus, 105, 138, 139, 220, 
221, 222, 223. 

pyrrhocephalus, 139. 

rubriventris, 119. 

rufidorsum, 115. 

salvini, 108. 

scalaris, 105, 137. 

scalaris scalaris, 136, 137, 220, 221, 
222, 223, 229, 

scalaris slevini, 136, 138, 219, 220, 
224, 227. 

sealaris unicanthalis, 136, 188, 222. 

serrifer plioporus, 122, 124, 218, 
226, 227, 229. 

serrifer serrifer, 122, 123, 218, 229. 

siniferus, 105, 184. 

siniferus cupreus, 134, 225. 

siniferus siniferus, 134, 218, 221, 
223, 225. 

spinosus, 106, 107, 116. 

spinosus caeruleopunctatus, 110, 
116, 225. 

spinosus floridanus, 117. 

spinosus spinosus, 110, 111, 116, 
126; 2175220; 221; 222°:223, 225, 
228, 229. 

squamosus, 134, 135, 218. 


249 


Sceloporus torquatus melanogaster, 122, 


126, 217, 221, 222, 223, 226, 229. 

torquatus minor, 128. 

torquatus mucronatus, 124, 125. 

torquatus omiltemanus, 124. 

torquatus poinsettii, 125. 

torquatus torquatus, 122, 125, 220, 
221, 222, 223, 225, 229. 

undulatus, 106, 107. 

undulatus consobrinus, 117, 118, 
219, 220, 224, 229. 

undulatus virgatus, 117, 118, 219, 227. 

utiformis, 105, 135, 220, 221, 222, 
223, 226. 

vandenburgianus, 119. 

variabilis, 105, 130. 

variabilis marmoratus, 130, 131, 
219, 224, 228. 

variabilis olloporus, 129, 130. 

variabilis smithi, 130, 131, 225. 

variabilis variabilis, 130, 218, 221, 
222, 225, 226, 228, 229. 

viviparus, 108. 

westphalii, 127. 

zosteromus, 115. 


schiedii, Anolis, 57, 58, 63, 64. 


Dactyloa, 63. 


schlegelii, Dermochelys, 13. 


Sphargis coriacea, 13. 


schmidti, Anolis, 56, 58, 60, 220. 


Cnemidophorus hyperythrus, 175, 
187, 217. 

Eumeces, 164, 

Phrynosoma, 103. 

Urosaurus ornatus, 142, 143, 219. 

Uta ornata, 143. 

Verticaria hyperythra, 187. 


schottii, Urosaurus ornatus, 141, 144, 


223, 226, 227. 
Uta, 144. 
Uta ornata, 144. 


schwartzei, Eumeces, 160, 162, 218, 227, 
229. 
Scincella, 155, 156. 


assata assata, 157, 160, 218. 

assata taylori, 157, 160, 218, 220, 
221, 222, 224. 

caudaequinae, 157, 158, 224, 226. 

cherriei cherriei, 157, 218, 227. 

cherriei ixbaac, 157, 158, 218, 229. 

cherriei stuarti, 157, 158, 224, 228. 

Seed forbesorum, 157, 159, 
222. 


gemmingeri gemmingeri, 157, 159, 
222, 224, 228. 

incerta, 156. 

laterale, 157, 159, 219. 

silvicola, 157, 158, 224, 228. 


stejnegeri, 107, 109, 221. 

taeniocnemis, 107. 

teapensis, 129, 180, 218, 225, 
229. 

torquatus, 105, 106, 121. 

torquatus binocularis, 122, 
224. 

torquatus cyanogenys, 125. 


secincicauda, Tropidolepis, 205. 
Scincidae, 40, 155. 
227, | Scincus agilis, 156. 
lateralis, 159. 
pavimentatus, 160. 
126, quinquelineatus, 163. 
sloanii, 156. 
telfairii, 156. 


250 


Scincus ventralis, 203, 214. 
scitulus Sceloporus formosus, 107, 109, 
221 


scorpioides, Kinosternon, 21. 
scutulatus, Sceloporus parvus, 129, 131, 
222, 226. 
sealous, Cnemidophorus gularis, 183. 
semicristata, Cyclura, 74. 
semifasciatus, Cnemidophorus gularis, 
184. 
Cnemidophorus sackii, 178, 184, 
219. 
Seps murinus, 175. 
septemvittatus, Cnemidophorus, 184. 
sericea, Verticaria, 186. 
sericeus, Anolis, 58, 67, 218, 224, 226, 
227, 228. 
Cnemidophorus gularis, 183. 
Serpentes, 2, 12, 37. 
serpentina, Chelydra, 20, 21, 218, 228. 
Testudo, 20, 21. 
serratus, Laemanctus, 70, 218, 221, 222, 
224, 226, 228, 229. 
serrifer, Sceloporus serrifer, 122, 123, 
218, 229. 


severus, Claudius, 27. 
sexlineata, Lacerta, 174, 185. 
sexlineatus, Cnemidophorus, 177, 185. 
shawianum, Cinosternon, 23. 
shawii, Cyclura, 74. 
Siderolamprus, 194. 
enneagrammus, 194. 
silvicola, Scincella, 157, 158, 224, 228. 
silvicolum, Leiolopisma, 158. 
similis, Ctenosaura, 73, 218, 224, 226, 
227, 228, 229. 
Cyclura (Ctenosaura), 73. 
Iguana (Ctenosaura), 73. 
siniferus, Sceloporus, 105, 134. 
Sceloporus siniferus, 134, 218, 221, 
223, 225. 
sinistra, Ameiva undulata, 171, 174, 
220, 222, 223, 225. 
skiltonianum, Plestiodon, 167. 
skiltonianus, Eumeces, 161, 167, 217. 
slevini, Coleonyx variegatus, 42, 45, 217. 
Sauromalus, 79, 80, 217. 
Sceloporus scalaris, 136, 138, 219, 
220, 224, 227. 
Streptosaurus, 91, 217. 
Uta, 91. 
sloanii, Scincus, 156. 
eats Sceloporus malachiticus, 
ae Sceloporus variabilis, 130, 131, 


smithii, Lepidophyma, 152. 
Lepidophyma smithii, 152, 218. 
solare, Phrynosoma, 96, 104, 217, 

226 2am 


solaris, Phrynosoma, 104. 


BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


sonoriense, Cinosternum, 26. 
Kinosternon, 22, 26, 219, 220, 226. 
Kinosternum, 26. 

Thyrosternum, 26. 
sonoriensis, od am variegatus, 42, 

» 221. 

Dipsosaurus dorsalis, 77, 79, 226, 227. 
Spasmocnemis, 51. 
speciosa, Blepharactisis, 192. 
spengleri, Testudo, 29. 
Sphaerodactylus, 41, 52. 

anthracinus, 52, 2138. 

cinereus, 52. 

fantasticus, 52. 

glaucus, 53. 

glaucus glaucus, 52, 53, 218, 225, 

227, 229. 

glaucus torquatus, 52, 58, 225, 226. 

inornatus, 53. 

lineolatus, 52. 

punctatissimus, 52. 

sputator, 52. 

torquatus, 53. 

Sphargis coriacea schlegelii, 13. 

spinifera, Amyda, 18. 

spinimentum, Phrynosoma, 102. 

spinosus, Sceloporus, 106, 107, 116. 
Sceloporus spinosus, 110, 111, 116, 

126, 217, 220, 221, 222, 223, 225, 
228, 229. 

splendidus, Callisaurus, 87. 

Callisaurus draconoides, 86, 87, 217. 
sputator, Sphaerodactylus, 52. 
Squamata, 12, 37. 
squamata, Eretmochelys, 17. 

Uta, 148, 149, 217. 
squamosus, Sceloporus, 134, 135, 218. 
stansburiana, Uta, 148. 

Uta stansburiana, 148. 

Stauremys, 27. 


Staurotypus, 21, 27. 
biporecatus, 27. 
marmoratus, 27. 
salvinii, 27, 224. 
(Stauremys) salvinii, 27. 
triporcatus, 27, 227, 228. 
steindachneri, Diploglossus, 195. 
stejnegeri, Cnemidophorus, 191. 
Cnemidophorus tessellatus, 191. 
Kinosternon flavescens, 22, 24, 219, 
220, 226. 
Sceloporus, 107, 109, 221. 
Uta stansburiana, 148, 150, 217, 
219, 220, 227. 


stellata, Uta, 148, 150, 217. 
Stenodactylus fuscus, 45. 
variegatus, 44. 
stictogrammus, Cnemidophorus sackii,. 
178, 183, 219, 227. 
Streptosaurus, 55, 91. 
mearnsi, 91, 217. 
slevini, 91, 217. 
striatus, Celestus, 194. 
Cnemidophorus guttatus, 180. 


INDEX 


251 


stuarti, Ameiva undulata, 171, 173, 218, | Terrapene, 29, 34. 
2 


24, 227. 
Anolis, 63. 
Leiolopisma cherriei, 158. 
Lygosoma cherriei, 158. 
Sceloporus melanorhinus, 110, 113, 
21 


Scincela cherriei, 157, 158, 224, 228. 
sugillatus, Sceloporus jarrovii, 123, 128, 
222. 
suleata, Chalcides, 39. 
sulcifrons, Anolis, 61. 
sumichrasti, Epaphelus, 192. 
Eumeces, 161, 164, 218, 228. 
Gymnophthalmus, 192, 224. 
Plistodon, 164. 
suspectum, Heloderma, 193, 227. 
Swanka, 21. 
leucostoma, 25. 
maculata, 23, 25. 
sylvatica, Gaigeia, 153, 154, 222. 
Lepidophyma, 154. 
symmetrica, Uta, 144. 
Uta ornata, 144. 
symmetricus, Urosaurus ornatus, 
144, 217, 227. 


Tachybates, 49. 
taeniata, Abronia .taeniata, 196, 197, 
222, 225, 228. 
taeniatus, Abronia, 197. 
Gerrhonotus, 197. 
taeniocnemis, Sceloporus, 107. 
Sceloporus malachiticus, 107, 218. 


Tapaia, 94. 
Tapaja, 94, 
Tapaya, 94, 95. 
boucardii, 98. 
cornuta, 99. 
cortezii, 98. 
dugesii, 98. 
hernandesi, 100. 
hernandezii, 100. 
orbicularis, 94, 98. 
orbicularis longicaudatus, 97. 
Tapayaxin, 97. 
Tapayia, 94. 
taurus, Phrynosoma, 95, 104, 221, 225. 
taylori, Anolis, 57, 61, 221. 
Leiolopisma assatum, 160. 
Scincella assata, 157, 160, 218, 220, 
221, 222, 224. 
Uta, 148, 150, 227. 
teapensis, Sceloporus, 
225, 227, 2 
tehuanae, Lepidophyma smithii, 152, 
218, 224. 
Teiidae, 39, 40, 170. 
telfairii, Leiolopisma, 156. 
Scincus, 156. 
temporalis, Barisia moreleti, 199, 218. 
Gerrhonotus moreleti, 199. 


tenuis, Liolaemus, 213. 
teres, Cyclura, 74. 


142, 


129, 130, 218, 


carolina, 34. 
coahuila, 34, 36, 219. 
goldmani, 35. 
klauberi, 34, 35, 227. 
mexicana, 35. 
mexicana mexicana, 34, 35, 226, 
228. 
mexicana yucatana, 34, 35, 225. 
nelsoni, 34, 35, 223. 
ornata, 34, 36, 219, 227. 
triporcata, 27. 
yucatana, 35. 
tesselata, Ameiva, 188. 
tesselatus, Cnemidophorus, 3, 176, 188. 
tessellatus, Cnemidophorus, 3) 
Cnemidophorus tessellatus, 189. 
Gerrhonotus, 203. 
Testudines, 2, 12. 
Testudinidae, 19, 27. 
Testudo acutirostris, 20. 
agassizli, 28. 
berlandieri, 28. 
caouana, 15, 16. 
caretta, 16. 
cephalo, 16. 
clausa, 34. 
concentrica, 36. 
concinna, 31. 
coriacea, 13. 
imbricata, 16. 
mydas, 17. 
picta, 33. 
polyphemus, 28. 
punctata, 37. 
serpentina, 20, 21. 
spengleri, 29. 
viridis, 17. 
tetragrammus, Eumeces, 161, 165, 226, 
228. 
Plestiodon, 165. 
texana, Anniella, 209. 
Cophosaurus, 81. 
Holbrookia, 82, 85, 219, 220, 224, 
226, 227, 228. 
Pseudemys, 33. 
Pseudemys floridana, 31, 33, 224. 
texanus, Cophosaurus, 85. 
thalassina, Petrosaurus, 90, 217. 
Uta, 90. 
Thalassochelys (Colpochelys) kempii, 
15. 


Thecadactylus, 41, 49. 
rapicauda, 49. 
rapicaudus, 49, 229. 

thermophila, Holbrookia, 84. 
Holbrookia elegans, 84, 


Holbrookia maculata, 82, 84, 226, 
227. 
thomasi, Ameiva undulata, 171, 173, 


218. 
Thyrosternum, 21. 
sonoriense, 26. 
tigris, Cnemidophorus sexlineatus, 182. 
Cnemidophorus tigris, 177, 189, 
217, 227. 


252 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 


torquata, Agama, 126. 


torquatus, Sceloporus, 105, 106, 121. 


unicolor, Cnemidophorus, 180. 
unicus, Urosaurus, 141, 147, 219. 


Sceloporus torquatus, 122, 125, 220, | uniformis, Anolis, 59, 60. 


221, 222, 223, 225, 229. 
Sphaerodactylus, 53. 


Sphaerodactylus glaucus, 52, 53, 


225, 226. 
Tropidurus, 121. 
townsendi, Sauromalus, 81. 


Sauromalus obesus, 79, 81, 227. 


Trachyderma, 192. 
horridum, 192, 193. 
Trachypilus, 55. 
tridactylus, Bipes, 38, 221. 
Hemichirotes, 38. 
triliratum, Cinosternum, 23. 
Trionicidae, 18. 
Trionychidae, 18. 
Trionychoidea, 12, 18. 
triporcata, Terrapene, 27. 


triporcatus, Staurotypus, 27, 227, 228. 


Tropidogaster, 94. 
‘Tropidolepis, 105, 205. 
formosus, 108. 
intermedius, 127. 
scincicauda, 205. 


tropidonotus, Anolis, 56, 59, 60, 218, 


224, 225, 227, 228, 229. 
Norops, 60. 

‘Tropidopilus, 55. 

Tropidurus bocourti, 213. 
occipitalis, 213. 
ptychopleurus, 213. 
torquatus, 121. 

tuberculata, Iguana, 72. 

Uta, 147. 
Uta bi-carinata, 147. 


tuberculatus, Urosaurus bicarinatus, 


141, 147, 220, 222, 226, 227. 
tuberculosus, Gekko, 50. 
Phyllodactylus, 47, 49, 217. 


tumidus, Sauromalus obesus, 79, 81. 


turcica, Lacerta, 50. 
turcicus, Hemidactylus, 40, 50. 


Hemidactylus turcicus, 50, 218, 


227, 228, 229. 


typica, Cnemidophorus mexicanus, 181. 


Uma, 54, 89. 
exsul, 89, 219. 
notata, 89. 
notata cowlesi, 89, 90, 227. 
notata notata, 89, 217. 
rufopunctata, 89. 
umbra, Emys, 33. 
Pseudemys, 31, 33, 224. 
unctus, Diplodactylus, 47. 
Phyllodactylus, 46, 47, 217. 


Phyllodactylus (Diplodactylus), 47. 


undulata, Agama, 105. 
Ameiva undulata, 71, 174, 224. 
undulatus, Cnemidophorus, 174. 
Sceloporus, 106, 107. 
unica, Uta, 147. 


unicanthalis, Sceloporus scalaris, 136, 


138, 222. 


Anolis humilis, 56, 60, 218, 227. 


Urosaurus, 55, 140. 


auriculatus, 140, 146, 220. 

bicarinatus anonymorphus, 141, 
146, 218, 221, 225. 

bicarinatus bicarinatus, 141, 146, 
221, 223, 225. 

bicarinatus nelsoni, 141, 147, 225. 

bicarinatus tuberculatus, 141, 147, 
220, 222, 226, 227. 

clarionensis, 141, 142, 220. 

gadovi, 140, 145, 222, 223. 

graciosus, 140, 141, 144, 217. 

irregularis, 140, 146. 

microscutatus, 140, 145, 217. 

nigricaudus, 140, 145, 217. 

ornatus caeruleus, 141, 148, 219. 

ornatus chiricahuae, 142, 148. 

ornatus graciosus, 145. 

ornatus linearis, 142, 148, 219, 227. 

ornatus ornatus, 142. 

ornatus schmidti, 142, 143, 219. 

Orua te schottii, 141, 144, 223, 226, 


aren symmetricus, 142, 144, 217, 
2 


unicus, 141, 147, 219. 


ustus, Anolis, 58, 66, 218, 229. 
Uta, 1, 55, 146, 147. 


anonymorpha, 146. 
auriculata, 146. 

bicarinata, 146. 

bicarinata anonymorpha, 146. 
bi-carinata bi-carinata, 146. 
bicarinata nelsoni, 147. 
bi-carinata tuberculata, 147. 
caerulea, 143. 

clarionensis, 142. 

concinna, 148, 150, 217. 
elegans, 148, 149. 

gadovi, 145. 

graciosa, 144. 

gularis, 144. 

irregularis, 146. 

lateralis, 144. 
(Phymatolepis) lateralis, 144. 
mannophorus, 148, 149, 217. 
martinensis, 148, 149, 217. 
mearnsi, 91. 

microscutata, 145. 

nelsoni, 147. 

nigricauda, 145. 

nolascensis, 148, 151, 227. 
ornata, 142. 

ornata chiricahuae, 143. 
ornata lateralis, 144. 

ornata linearis, 143. 

ornata ornata, 142. 

ornata schmidti, 143. 
ornata schottii, 144. 

ornata symmetrica, 144. 
palmeri, 148, 151, 227. 
parva, 149. 


INDEX 


Uta parviscutata, 145. 
repens, 90. 
schottii, 144. 
slevini, 91. 
squamata, 148, 149, 217. 
stansburiana, 148. 
stansburiana elegans, 148, 150, 217. 
stansburiana hesperis, 148, 149, 217. 
stansburiana stansburiana, 148. 
stansburiana stejnegeri, 148, 150, 
217, 219; 220, 227. 
stellata, 148, 150, 217. 
symmetrica, 144. 
taylori, 148, 150, 227. 
thalassina, 90. 
tuberculata, 147. 
unica, 147. 
utiformis, Sceloporus, 
221, 222, 223, 226. 
utowanae, Anolis, 57, 62, 226. 


105, 135, 220, 


vandenburghi, Cnemidophorus, 191. 
vandenburgianus, Sceloporus, 119, 217. 
Sceloporus graciosus, 119, 217. 

variabilis, Sceloporus, 105, 130. 
Sceloporus variabilis, 130, 218, 221, 
222, 225, 226, 228, 229. 
variegatus, Coleonyx, 44. 
oleonyx variegatus, 42, 44, 217, 
224. 
Eublepharis, 44. 
Stenodactylus, 44. 
variolosus, Cnemidophorus, 190. 
varius, Sauromalus, 79, 81, 227. 
velox, Cnemidophorus gularis, 184. 
ventralis, Callisaurus, 88. 
Callisaurus draconoides, 86, 88, 227. 
Callisaurus ventralis, 88. 
Homalosaurus, 86, 88. 
Scincus, 203, 214. 
venusta, Emys, 31. 
veraepacis, Anolis ustus, 67. 


Verticaria, 174. 
beldingi, 186. 
caerulea, 186. 
ceralbensis, 188. 
espiritensis, 187. 
franciscensis, 187. 
hyperythra beldingi, 186. 
hyperythra hyperythra, 187. 
hyperythra schmidti, 187. 
picta, 186. 
sericea, 186. 


« 

253 

verus, Cnemidophorus gularis gularis, 
182 


vigilis, Xantusia, 154, 217. 

virgata, Chelonia, 17. 

virgatus, Sceloporus undulatus, 117, 118, 
219, 227. 

viridiflava, Barisia, 198, 200, 224. 

viridifiavus, Gerrhonotus, 200. 

viridis, Anolis, 55. 

Testudo, 17. 

vittatus, Basiliscus, 71, 218, 220, 221, 

222, 223, 224, 225, 227, 228, 229. 


webbii, Elgaria multicarinata, 205, 207, 
217 


Gerrhonotus, 207. 

Gerrhonotus seincicauda, 207. 
westphalii, Seeloporus, 127. 
wiegmanni, Anolis, 63. 

Phrynosoma, 97. 
wislizenii, Crotaphytus, 94. 

Gambelia, 93, 94. 

Gambelia wislizenii, 94, 217, 219, 

227. 


xanti, Phyllodactylus, 49. 
Xantusia, 151, 154. 
gilberti, 154, 155, 217. 
henshawi, 154, 155, 217. 
picta, 155. 
vigilis, 154, 217. 
Xantusiidae, 39, 151. 
Xenosauridae, 40, 207. 
Xenosaurus, 207. 
fasciatus, 207, 208. 
grandis, 207, 208, 225, 229 
newmanorum, 207, 208, 226. 
rackhami, 208, 218. 


Xerobates agassizii, 28. 
berlandieri, 28. 


yueatana, Cistudo, 35. 
Terrapene, 35. 
Terrapene mexicana, 34, 35, 225. 


yucatanicus, Norops, 60. 


Zablepsis, 154. 


zosteromus, Sceloporus, 115. 
Sceloporus magister, 112, 115, 217. 





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